Front Range, Eastern Slope, Rocky Mountains Checklist Flora of Native and Naturalized Vascular Plants of Golden and Vicinity, Jefferson County, Colorado (Continued)  

Tom Schweich  

Home Page
Topics in this Article:
Introduction
Geography
History of Botanic Exploration and Publication
Useful Publications
Methods
Results
Discussion - Native Plants
Discussion - Non-Native Plants
Conclusion
Acknowledgements
Literature Cited
Appendices
 Golden, Colorado sits in a valley formed by erosion along the Golden fault, the geotectonic boundary between the North American Cordillera and the Great Plains. Somewhat like Mono Lake, for which I have also prepared a checklist flora, it sits at a boundary, or perhaps ecotone. Things are always more interesting at the boundaries. I started this project when I realized no such list had been prepared for my newly adopted city. I hope you find this checklist flora helpful. Please write to me if you have questions or comments.

 

   

Biodiversity

 

Literature Cited:
- Colwell, Robert K., 2008.
- Savard, Jean-Pierre L., Philippe Clergeau, and Gwenaelle Mennechez, 2000.  

What is biodiversity?

 

 

   

Appendices

1 — Introduction

2 — Geography

3 — History of Botanic Exploration

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4 — Useful Publications

  • 411 — Floras
    • 4112017w — "411" & publication year & author.
  • 412 — Botanical but not a flora.

5 — Methods

6 — Results

  • 61 — Collections Found
  • 62 — Collections Made

63 — Discussion

  • 631 — Rare Plants
  • 632 — Notable Native Plants
    • 6321 -- Ferns, if any.
    • 6322 -- Gymnosperms, if any.
    • 6323 -- Dicots
      • 6323280Erinaugra -- "6323" & family & genus & species & subspecies or variety
    • 6324 -- Monocots
  • 633 — Notable Non-Native Plants
  • 633n — Noxious Weeds
    • 63402 — Non-Native Grasses in the Golden Landscape.
      • 63404020Agrcri -- "63404" & family & genus & species & sequence number, if needed.
    • 6342 — Non-Native Mustards in the Golden Landscape
      • 6342106Alyaly -- "6342" & family & genus & species & sequence number, if needed.
    • 6343 — Non-Native Sunflowers in the Golden Landscape
    • 6349 — Other Families
  • 637— Distribution of Natives
    • 6371 — Global
    • 6372 — Circumpolar
      • 63720510088Stelon — Stellaria longipes Goldie
    • 6373 — Eastern North America
    • 6374 — northern prairie
    • 6375 — Texas
    • 6376 — Rocky Mountains
    • 6377 — Cordillera
    • 6378 — southern Cordillera
      • 63780510250Giloph — Gilia ophthalmoides

7 — Conclusion

8 — Acknowledgements

  • 90 — Ecological Systems of Colorado
  • 91 — GIS Resources to Accompany the Checklist Flora
  • 92 — How the Flora is Built
  • 93 — Source Data
    • 931 — Types from the Golden Area
    • 932 — Namesakes of the Golden Area
    • 933 — Notes on All Taxa, Regardless of Nativity.
      • 9330300 — Notes on Ferns and Allies.
      • 9330400 — Notes on Gymnosperms.
      • 9330510 — Notes on Dicots.
      • 9330520 — Notes on Monocots.
    • 934 — Studies of Prairie or Foothill Ecosystems
    • 935 — Restoration, and Effects of Attempts Thereof
    • 936 — Recreation in Foothill and Prairie Ecosystems
    • 937 — Notes on Non-Native Species
      • 937 0520 — Notes on Non-Native Monocots -- Do not use, place with the 933s.
  • 94 — Keys
  • 95 — Schemas
  • 96 — Vegetation Descriptions
  • 97 — Reminders
  • 98 — Mystery Locations

 

   

Ecological Systems of Colorado

 
  The Colorado Natural Heritage Program page on Ecological Systems of Colorado is found at: http://www.cnhp.colostate.edu/download/projects/eco_systems/eco_systems.asp .

Literature Cited:
- Faber-Langendoen, Don, Ralph H. Crawford, and David L. Tart, 2009.
- Federal Geographic Data Committee, 2008.
- Jennings, Michael D., Don Faber-Langendoen, Orie L. Loucks, Robert K. Peet,m and David Roberts, 2009.  

 
  Comparison of published vegetation types.
 
CNHP, 2005O'Shea-Stone, 2002Kilburn & White, 1992Zeise, 1976
    Lichen-rock type. Lichen stand types. Areas of bare rock from steep lava cliffs to the conical peaks on the mesa surface.
  Grasslands
  • Short-grass grassland. Bouteloua gracilis, Bromus tectorum, with Alyssum parviflorum, and Opuntia sp., Echinocereus viridiflorus, Escobaria missouriensis, Escobaria vivipara var. vivipara. Also Hesperostipa comata (Syn: Stipa c.), and Yucca glauca. Some short shrubs of Chrysothamnus nauseosus ssp. graveolens, Prunus virginiana (Syn: Padus v.), Rhus aromatica ssp. trilobata. Celtis reticulata at edge of mesa. Traditionally dominated by Bouteloua gracilis and Buchloe dactyloides, but now dominated by Bromus tectorum.
Grassland type. Bromus tectorum and Alyssum simplex (Syn: A. minus. Occasional Achnatherum scribneri (Syn: Stipa s.) and Andropogon gerardii. Mixed-grass stand types. Dominated by Bromus tectorum and Agropyron sp. (Elymus sp. ?), with Buchloe dactyloides, and Alyssum alyssoides, Eriogonum umbellatum Torr., Heterotheca villosa, Opuntia compressa, Yucca glauca, and Ericameria nauseosa (Syn: Chrysothamnus nauseosus). West, south, and east exposures.
  Mixed-grass grassland. Stipa comata, Pascopyrum smithii, Bouteloua gracilis, Bromus tectorum, with Andropogon gerardii, Bouteloua curtipendula, Aristida purpurea, and Nassella viridula, with a large number of forbs. Mesa slopes and toe areas of STM.
RM Aspen Forest and Woodland - - -
RM Cliff, Canyon and Massive Bedrock - - -
RM Dry-Mesic and Mesic Montane Mixed Conifer Forest and Woodland - - -
SRM Pinyon-Juniper Woodland - - -
SRM Ponderosa Pine Woodland - - -
Rocky Mountain Lower Montane - Foothill Shrubland.
  • Cercocarpus montanus Shrubland Alliance
    • Series determination requires more data collection.
Upland shrubland. Cercoparpus montanus, with sparse cover of Bromus tectorum intermixed with Hesperostipa comata (Syn: Stipa c.), Yucca glauca, and many cacti.

Ravine shrubland. Skunkbush, chokecherry and Prunus americana, in dense thickets. Few plains cottonwoods and Salix amygdaloides

Shrubland type

  • Mixed shrub community. Symphoricarpos occidentalis, Cercocarpus montanus, Rhus [aromatica] ssp. trilobata, Ribes cereum, Prunus americana, and Prunus virginiana. Understory of Poa pratensis, Bromus tectorum, Elymus trachycaulus (Syn: Agropyron trachycaulum), Eriogonum umbellatum, Alyssum alyssoides, etc.
Mixed shrub stand types. Rhus [aromatica] ssp. trilobata, Ribes cereum Dougl., Symphoricarpos occidentalis, Cercocarpus montanus, Prunus virginiana L., Prunus americana Marsh. Acer glabrum in dense patches. Mostly north exposures.
    Shrubland type.

  • Mountain mahogany community. Cercocarpus montanus with an understory of Alyssum alysoides, Bromus tectorum, Agropyron cristatum (Syn: A. desertorum), Eriogonum umbellatum, etc.
Pure shrub stand types. Cercocarpus montanus, with Bromus tectorum, Alyssum alyssoides, and Eriogonum umbellatum.
    Grassland-shrub type. Common foothills species: Symphoricarpos occidentalis, Prunus americana, Rhus trilobata, and Ribes cereum. Grasses are Poa pratensis, Bromus tectorum, and Elymus trachycaulus (Syn: Agropyron trachycaulum). Also Cercoparpus montanus, Symphoricarpos rotundifolius (Syn: S. oreophilus), Prunus virginiana melanocarpa, Rosa arkansana, Physocarpus monogynus, and Ribes aureum. Shrub cover within grassland matrix is significant, but less than 50%. Shrub-grass stand types. Shrubs of Crataegus succulenta (Syn: C. erythropoda), Rosa sp., Rhus trilobata, Prunus virginiana, Prunus americana, Celtus reticulata, and Ribes cereum, with Agropyron sp. (Syn: Elymus sp. ?), Bromus tectorum, Achnatherum hymenoides (Syn: Oryzopsis h., and Alyssum alyssoides. Patches of shrubs in mixed grass-forb areas.
NA Arid West Emergent Marsh - - -
  Wetlands. Cottonwoods and willows, with Carex spp., and Juncus spp., and a variety of grasses and forbs. Patches of Typha spp.. Hydrology alteration.   Riparian (streamside) stand types. Salix exigua, Populus sargentii, Eleocharis macrostachya, Scirpus lacustris L., and Mentha spicata L.
    Woodland type.

  • Mountain maple community. Dense community of small Acer glabrum just below cliffs or in ravines with a dense understory of mixed shrub. North and east slopes.
  • Cottonwood woodland community. Scattered cottonwoods (Populus deltoides ssp. monilifera (Syn: P. sargentii), and P. angustifolia) along permanent and intermittent streams. In Big Ravine, Acer negundo, Salix exigua, and S. amygdaloides also occur. Other shrubs also form a dense understory.
  • Juniper Savannah community. Juniperus scopulorum with an understory of typical grassland.
Woodland stand types. Juniperus scopulorum, with Bromus tectorum.
WGP (Western Great Plains) Cliff, Outcrop, and Shale Barrens - - -
WGP (Western Great Plains) Closed Depression Wetland - - -
WGP (Western Great Plains) Foothill and Piedmont Grassland - - -
WGP (Western Great Plains) Riparian Woodland, Shrubland and Herbaceous - - -
WGP (Western Great Plains) Shortgrass Prairie - - -
  Disturbed/reclaimed    
  Developed    

Notes:

  1. [R3C3] Stipa comata grassland of 30-60 acres near western rim of mountain, may be partly due to effects of the 1988 fire.

   

Recognized Ecological Systems

 

   

Ecological Systems Recognized by the Colorado Natural Heritage System

 

Literature Cited:
- Colorado Natural Heritage Program, 2005.  

Ecological systems are dynamic assemblages or complexes of plant and/or animal communities that 1) occur together on the landscape; 2) are tied together by similar ecological processes, underlying abiotic environmental factors or gradients; and 3) form a readily identifiable unit on the ground. These systems provide a coarser level unit than plant associations and alliances as defined under the International Vegetation Classification standard, and are more easily identified on the ground.

The descriptions and summarized viability guidelines presented here are intended to serve as a tool for conservation and management planning by providing a context for conservation and management (i.e., what systems do we have in Colorado), and by providing easy access to ranking and evaluation criteria for key ecological attributes of each system (i.e., what is the condition of our systems).

System descriptions and viability guidelines are based on materials compiled by NatureServe or developed by the Colorado Natural Heritage Program. Funding for the development of these documents was provided in part by the Bureau of Land Management, The Nature Conservancy, and the USDA Forest Service (CNHP, 2005).

Literature Cited:
- Colorado Natural Heritage Program, 2005.  

 

   

Southwest Regional Gap Analysis Project (SWReGAP)

 

Literature Cited:
- Rondeau, R., K. Decker, J. Handwerk, J. Siemers, L. Grunau, and C. Pague, 2011.  

The Southwest Regional Gap Analysis Project (SWReGAP) was a mapping and assessment of biodiversity for the five-state region encompassing Arizona, Colorado, Nevada, New Mexico, and Utah. The area comprises approximately 150 million hectares (560,000 square miles) representing 1/5 the coterminous United States. The primary objective of the project was to use a coordinated approach to create detailed, seamless maps of the land cover, habitat for native terrestrial vertebrate species, land stewardship, and management status for the Southwest region. This information was analyzed to identify animal species habitats and natural land cover types that are underrepresented on land managed for their long term conservation. SWReGAP was a multi-institutional effort with scientists based in all five southwest states.

   

USNVC -- United States National Vegetation Classification

 
  http://usnvc.org/
  The U.S. National Vegetation Classification is supported by a formal partnership between the federal agencies, the Ecological Society of America (ESA), and NatureServe, working through the Federal Geographic Data Committee (FGDC) Vegetation Subcommittee. Primary signators include the U.S. Forest Service (which chairs the subcommittee), ESA, NatureServe, and the U.S. Geological Survey Core Science Systems (USGS/CSS). Together we are committed to supporting the implementation and maintenance of the National Vegetation Classification (NVC) Standard (FGDC 2008).
  The overall objective of the Vegetation and Information Standards is to support the use of a consistent national vegetation classification system (NVCS) to produce uniform statistics in vegetation resources from vegetation cover data at the national level. It is important that, as agencies map or inventory vegetated Earth cover, they collect enough data accurately and precisely to translate it for national reporting, aggregation, and comparisons. Adoption of the Vegetation Classification and Information Standards in subsequent development and application of vegetation mapping schemes will facilitate the compilation of regional and national summaries. In turn, the consistent collection of such information will eventually support the detailed, quantitative, geo-referenced basis for vegetation cover modeling, mapping, and analysis at the field level.

   

EcoVeg

 

Literature Cited:
- Faber-Langendoen, Dom, Todd Keeler-Wolf, Del Meidinger, Dave Tart, Bruce Hoagland, Carmen Josse, Gonzalo Navarro, Serguei Ponomarenko, Jean-Peirre Saucier, Alan Weakley, and Patrick Comer, 2014.  

http://www.natureserve.org/biodiversity-science/publications/ecoveg-new-approach-vegetation-description-and-classification

 

   

GIS Resources to Accompany the Checklist Flora

 

 

   

How the Flora is Built

 

 

   

Source Data

 
 

Mentzelia

 

Literature Cited:
- Hufford, Larry, Michelle M. McMahon, Anna M. Sherwood, Gail Reeves, and Mark W, Chase, 2003.  

Names recognized by Harrington (1954) Names recognized by Snow (2009) Names recognized by Weber and Wittmann (2012) Names recognized by Ackerfield (2015) Names recognized by FNANM Mentzelia (s.l.) represented by collections in Jefferson County, Colorado
Section Bartonia
Mentzelia multiflora (Nutt.) Gray
(Syn: M. speciosa Osterhout., Nuttallia multiflora (Nutt.) Greene, N. speciosa (Osterh.) Greene, N. sinuata Rydb.)
Mentzelia multiflora (Nutt.) A. Gray var. multiflora Nuttallia multiflora (Nuttall) Greene
(Incl: N. sinuata, N. speciosa)
Mentzelia multiflora (Nutt.) Gray Mentzelia multiflora (Nutt.) Gray M. multiflora (Nutt.) Gray
Mentzelia sinuata (Rydb.) R. J. Hill Mentzelia speciosa Osterh. Mentzelia speciosa Osterh. Mentzelia sinuata (Rydb.) R. J. Hill
Mentzelia speciosa Osterh. Mentzelia speciosa Osterhout
Mentzelia nuda (Pursh) T. & G.
(Syn: Nuttallia nuda (Pursh) Greene)
Mentzelia nuda (Pursh) Torr. & A. Gray Nuttallia nuda (Pursh) Greene Mentzelia nuda (Pursh) Torr. & A. Gray Mentzelia nuda (Pursh) Torr. & A. Gray M. nuda (Pursh) Torr. & Gray
Section Trachyphytum
Mentzelia albicaulis Dougl ex Hook. Mentzelia albicaulis (Douglas ex Hook.) Douglas ex Torr. & A. Gray Acrolasia albicaulis (Douglas) Rydberg Mentzelia albicaulis (Douglas ex Hook.) Douglas ex Torr. & A. Gray
(Syn: M. montana (Davidson) Davidson)
M. albicaulis (Dougl. ex Hook.) Dougl. ex Torr. & Gray M. albicaulis (Dougl. ex Hook.) Dougl. ex Torr. & Gray
Not recognized Mentzelia montana (Davids.) Davids. Not recognized M. montana (Davidson) Davidson M. montana (Davidson) Davidsona
Mentzelia dispersa Wats. Mentzelia dispersa S. Watsonb Acrolasia dispersa (S. Watson) Davidson Mentzelia dispersa S. Watson M. dispersa S. Wats. M. dispersa S. Wats.
Harrington, H. D. 1954. Manual of the plants of Colorado. Denver, CO.: Sage Books., 1954. Snow, Neil. 2009. Checklist of Vascular Plants of the Southern Rocky Mountain Region (Version 3). 316 p. Weber, William A., and Ronald C. Wittmann. 2012. Colorado Flora: Eastern Slope. 4th Edition. Boulder, Colorado: University Press of Colorado, 2012. Ackerfield, Jennifer. 2015. The Flora of Colorado. Fort Worth, TX. Botanical Research Institute of Texas. Flora of North America Editorial Committee, eds. 1993+. Flora of North America North of Mexico. 16+. New York and Oxford. Published on the Internet; http://www.efloras.org/ (accessed 2013 through 2019) Source: Southwest Environmental Information Network, SEINet. 2014. http//:swbiodiversity.org/seinet/index.php. Accessed on September 04, 2014.

Notes:

  1. bMentzelia dispersa S. Watson. Snow (2009) follows Dorn (2001) in not recognizing varieties in our region.
  2. aMentzelia montana (Davidson) Davidson is represented by two collections made by George E. Osterhout with Ira W. Clokey, Osterhout's #3095 (RM95508) and #5741 (RM162001). The collections were made 22 June 1918 and the locality is Morrison, Jefferson County, Colorado.
 

Symphoricarpos

 

Literature Cited:
- Bell, Charles D., 2010.  

“Towards a Species Level Phylogeny of Symphoricarpos (Caprifoliaceae), Based on Nuclear and Chloroplast DNA”

Literature Cited:
- Bell, Charles D., 2010.  

Authors for Symphoricarpos in FNANM are: Applequist, Wendy L./wendy.applequist at mobot.org and Bell, Charles D./valerianaceae1969 at gmail.com. Caprifoliaceae will be contained in Volume 18, which, as of this date, 3 August 2014, is under production.

 

   

Botanical Explorers

 

   

Thomas Nuttall

 
 

1810-1811 Missouri River

 
 

1815-1816 The Carolinas

 
 

1816-1817 Ohio River to South Carolina

 
 

1818-1820 Arkansas

 

 

   

Names of Historical Reports

 
 

The following is an intentionally empty table …

Abbreviation Full Title Availability Online
  • …abbreviation…
… full name of publication … URL: … if available …

Literature Cited:
- Plukenet, Leonard, 1696.  

One of several pre-Linnaean botanical texts.

Abbreviation Full Title Availability Online
Pluk. Alm. Plukenet, Leonard. 1696. Almagestum botanicum. Londini : Sumptibus auctoris 1696 URL: https://bibdigital.rjb.csic.es/idurl/1/10874

Cited by Lamarck (1783) as a source for Arabis reptans (=Draba reptans).

 

Literature Cited:
- Dillenius, Johan Jacob, 1732.

Other articles:
• Golden Checklist Flora:  Notes on Symphyotrichum ericoides;  

Dillenius, 1732 Hortus Elthamensis

Abbreviation Full Title Availability Online

Dill. elt.
Hortus Elthamensis, seu Plantarum rariorum, quas in horto suo Elthami in Cantio coluit ... Jacobus Sherard ... Guilielmi ... frater delineationes et descriptiones URL: http://bibdigital.rjb.csic.es/ing/Libro.php?Libro=1386
Dillenius was born at Darmstadt and was educated at the University of Giessen, earlier the family name had been changed from Dillen to Dillenius. In 1721, at the instance of the botanist William Sherard, he moved to England. In 1734 Dillenius was appointed Sherardian professor of botany at Oxford, in accordance with the will of Sherard, who at his death in 1728 left the university £3000 for the endowment of the chair, as well as his library and herbarium, all on the condition that Dillenius should be appointed the first professor.

In 1732 he published Hortus Elthamensis, a catalogue of the rare plants growing at Eltham, London, in the collection of Sherard's younger brother, James (1666—1738), who, after making a fortune as an apothecary, devoted himself to gardening and music. For this work Dillenius himself executed 324 plates; it was described by Linnaeus, who spent a month with him at Oxford in 1736, and afterwards dedicated his Critica Botanica to him, as opus botanicum quo absolutius mundus non vidit, "a botanical work of which the world has not seen one more authoritative". Further, Linnaeus would later name a genus of tropical tree Dillenia in his honor. In the section on Symphyotrichum ericoides refers to the work of Paul Herman, Prodromus of Dutch Flowers, perhaps published posthumously, as follows:

Ab Aftere Novae Angliaw Linariae folio, Chamaemeli floribus, Hermannus Par. Bat. p. 95. …
This appears to refer to:
D. Pauli Hermanni Florae Batavae flores, Ejusdem Paradisi Batavi Prodromus; & tendem post idsius obitum Paradisus ipsa :
That I found in: Joannis Raii (John Ray) … 1704 … Historiae plantarum: tomus tertius … : accessit Historia Stirpium ins. Luzonis & reliquarum Philippinarum a R. P. Gee. Jos. Camello … S. J. conscripta : item D. Jos. Pitton Tournefort … Grollarium institutionum rei herbariae

 

Literature Cited:
- Gronovius, Jan Frederik, and John Clayton, 1738-1743.

Other articles:
• Golden Checklist Flora:  Notes on Celtis occidentalis, Linnaeus, 1753;  Notes on Symphyotrichum ericoides;  

Gronovius, 1738, Flora Virginica

Abbreviation Full Title Availability Online

Gron. Virg.
Flora Virginica, exhibens plantas URL: https://www.biodiversitylibrary.org/item/122722

Jan Frederik Gronovius (also seen as Johann Frederik and Johannes Fredericus) (10 February 1690 in Leiden – 10 July 1762 in Leiden) was a Dutch botanist notable as a patron of Linnaeus.

John Clayton, a plant collector in Virginia sent him many specimens, as well as manuscript descriptions, in the 1730s. Without Clayton's knowledge, Gronovius used the material in his Flora Virginica (1739–43, 2nd ed. 1762).

He was the son of Jakob Gronovius and grandson of Johann Friedrich Gronovius, both classical scholars. In 1719, he married Margaretha Christina Trigland, who died in 1726, and Johanna Susanna Alensoon in 1729. His son Laurens Theodoor Gronovius (1730–1777) was also a botanist.

 

Literature Cited:
- Van Royen, Adrianus, 1740.

Other articles:
• Golden Checklist Flora:  Notes on Symphyotrichum ericoides;  

Royen, 1740, Flora Leydensus Prodromus

Abbreviation Full Title Availability Online

Roy. lugdb.
Flora Leydensus Prodromus

Florae leydensis prodromus :exhibens plantas quae in Horto academico Lugduno-Batavo aluntur.

URL: https://www.biodiversitylibrary.org/item/13843

Adriaan van Royen (11 November 1704 in Leiden – 28 February 1779 in Leiden) was a Dutch botanist. He was a professor at Leiden University and is associated with Carl Linnaeus.

He is best known for his work on flora of Southeast Asia. Adriaan van Royen formed a close relationship with Linnaeus, who had visited the Leiden Botanic Garden during a stay in Leiden between 1737 and 1738.

He was elected a Fellow of the Royal Society in 1728.

He died in Leiden in 1779.

 

Literature Cited:
- Gmelin, Johan Georg, 1747-1769.  

Gmelin, 1747-1769, Publication Details

Abbreviation Full Title Availability Online

Gmel. Sibir.
Gmelin, Johan Georg. 1747-1769. Flora Sibirica sive Historia Plantarum Siberiae. St. Petersburg: Imperial Academy of Sciences. URL: https://gdz.sub.uni-goettingen.de/id/PPN330224352

 

Other articles:
• Golden Checklist Flora:  Delphinium carolinianum, Walter, 1788;  

Walter, 1788, Flora Caroliniana

Thomas Walter (1740-1789) was a British-born American botanist. He is best known for his book Flora Caroliniana (1788), an early yet fairly complete catalog of the flowering plants of South Carolina.

Abbreviation Full Title Availability Online

Fl. Carol.
Flora caroliniana : secundum systema vegetabilium perillustris Linnaei digesta; characteres essentiales naturalesve et differentias veras exhibens; cum emendationibus numerosis: descriptionum antea evulgatarum: adumbrationes stirpium plus mille continens: necnon, generibus novis non paucis, speciebus plurimis novisq. ornata URL: https://www.biodiversitylibrary.org/item/38202

 

Literature Cited:
- Schreber, Johann Christian Daniel van, 1791.

Other articles:
• Golden Checklist Flora:  Agoseris, Schreber, 1791;  

Schreber, 1789-1791, Publication Details

The following is an intentionally empty table …

Abbreviation Full Title Availability Online

Gen. Pl., ed. 8[a]
Genera Plantarum Eorumque Characteres Naturales Secundum Numerum, Figuram, Situm, & Proportionem Omnium Fructificationis Partium. (Ed. 8[a]). URL: … if available …

 

Literature Cited:
- Gaertner, Joseph, 1788-1807.  

Gaertner, 1791, Publication Details

Joseph Gaertner (1732 – 1791) was a German botanist, best known for his work on seeds, De Fructibus et Seminibus Plantarum (1788-1792).

He was born in Calw, and studied in Göttingen under Albrecht von Haller. He was primarily a naturalist, but also worked at physics and zoology. He travelled extensively to visit other naturalists. He was professor of anatomy in Tübingen in 1760, and was appointed professor of botany at St Petersburg in 1768, but returned to Calw in 1770. Much of his productive work involved hybridization of plants and expression of traits in progeny.

Abbreviation Full Title Availability Online

Fruct. Sem.
De Fructibus et Seminibus Plantarum: accedunt seminum centuriae quinque priores cum tabulis Aeneis LXXIX. URL: https://www.biodiversitylibrary.org/page/37208351

Gaertner (1791) appears here because of his publication of Troximon. Today we treat Troximon as a synonym of Krigia Schreb. and the published names in Troximon have been dispersed primarily to Agoseris with one to Krigia and two to Nothocalais. Krigia biflora (Walter) S. F. Blake “Two-Flower Dwarf Dandelion” is found in Colorado, including one collection near Deckers in southwest Jefferson County. Nothocalais cuspidata (Pursh) Greene “Prairie False Dandelion” is common around Golden s.l.

 

Literature Cited:
- Fraser, John, 1813.
- Greene, Edward L., 1889.

Other articles:
• Golden Checklist Flora:  Astragalus crassicarpus, Nuttall, 1813;  Glycyrrhiza lepidota, Nuttall, 1813;  Sphaeralcea coccinea, Fraser, 1813;  Notes on Escobaria vivipara, Fraser, 1813;  Oenothera cespitosa, Fraser, 1813;  Ratibida columnifera, Fraser, 1813;  

Fraser's Catalogue, 1813

Abbreviation Full Title Availability Online
Cat. Pl. Upper Louisiana
Fraser's Cat.
Fraser catal.
Catalogue of New and Interesting Plants Collected in Upper Louisiana and Principally on the River Missouri, North America, for Sale at Messrs. Fraser's Nursery for Curious American Plants, Sloane, Square, King's Road, Chelsea, [London 1813] A reprint is available at URL: https://www.biodiversitylibrary.org/item/52475#page/119/

Remarks the International Plant Names Index (accessed 16 August 2020): Originally distributed by Messrs. Fraser’s Nursery for Curious American Plants and was referred to as Frasers’ catalogues. Reprinted in: Pittonia. 2: 114-119. 1890. Although Nuttall is not cited as the author, he has been generally accepted as the author (see St. Louis Code Art. 46 Ex. 26). Regarding the precise date of publ.: a reference to this Cat. Is made in Bot. Mag. Ad t. 1574. 1 Aug 1813. For several names in the supplement and addenda et corrigenda portions, Pursh (Fl. Amer. Sept. 2: 727-751. Dec 1813) referred to Fraser's Catalogue, e.g, pp. 728, 729, 750. For the validly and invalidly published names in this Catalogue, see: J. L. Reveal, Rhodora 70: 25-54. 1968.

  • Amorpha canescens Nutt., Cat. Pl. Upper Louisiana [1] (1813), nom. Inval.
  • Amorpha nana Nutt., Cat. Pl. Upper Louisiana no. 5 (1813).
  • Astragalus crassicarpus Nutt., Cat. Pl. Upper Louisiana no. 6 [unpaged] (1813).
  • Cactus viviparus Nutt. (= Escobaria vivipara (Nutt.) Britton & Rose), Cat. Pl. Upper Louisiana no. 22 (1813). .
  • Eriogonum flavum Nutt., Cat. Pl. Upper Louisiana [unpaged] (1813).
  • Glycyrrhiza lepidota Nutt., Cat. Pl. Upper Louisiana no. 45 (1813).
  • Malva coccinea Nutt. (=Sphaeralcea coccinea (Nutt.) Rydb.), Cat. Pl. Upper Louisiana no. 51 (1813).
  • Oenothera albicaulis Nutt., Cat. Pl. Upper Louisiana no. 54 (1813), nom. Inval.
  • Oenothera cespitosa Nutt., Cat. Pl. Upper Louisiana no. 53 (1813).
  • Rudbeckia columnifera Nutt., Cat. Pl. Upper Louisiana no. 75 (1813).
  • Yucca glauca Nutt., Cat. Pl. Upper Louisiana no. 89 (1813).

Literature Cited:
- Reveal, James L., 1968.

Other articles:
• Golden Checklist Flora:  Ratibiba columnifera, Reveal, 1968;  

Multiple authors have considered the names published in Fraser's 1813 Catalogue, the author of the names, and the legitimacy of the names. One recent such paper was Reveal (1968) “On the Names in Fraser's 1813 Calalogue.” Original copies of Fraser's Catalogue are quite rare, leading Greene (1890) to republish. Reveal acknowledges that Greene essentially accurately reprinted the catalogue. One change that Greene acknowledges was the addition “[ By T. Nuttall. ]” to the title page, which he says Nuttall wrote in ink on the original.

Possibly among other taxa, the legitimacy of the names in Fraser's Catalogue is important for our Priairie Coneflower Ratibida columnifera (Nutt.) Woot. & Standl.

Literature Cited:
- Cronquist, A, D. D. Keck, and B. Maguire, 1956.
- Graustein, Jeannette E., 1967.
- Reveal, James L., 1968.
- Shinners, Lloyd H., 1955.
- Shinners, Lloyd H., 1956.  

Reveal (1968) thought the author must have been Nuttall, referring to several other papers on the question.

… The interested reader on this subject is invited to refer to the series of arguments presented by Shinners (1949, 1955, 1956) for Nuttall not being the author and the Catalogue as an invalid source of publication; Graustein's (1956) contentions that Nuttall was the author but that he considered the species not validly published because of an agreement with his sponsor, Benjamin Smith Barton of Philadelphia, not to publish any new species without Barton's consent; and Cronquist, Keck, and Maguire (1956) who believe that Nuttall was the author and that the Catalogue is a valid place of publication.
 

It has been common knowledge that Nuttall was at least associated with the names found in the Catalogue. Pennell (1936) and Graustein (1967) point out that he was in England and associated with the Fraser Brothers' Nursery at the time of publication. From Nuttall's two subsequent publications (1817, 1818) which followed shortly after the Catalogue, we know that he considered at least some of the names as his own. Several of the early authors attributed the names in the Catalogue to Nuttall, and they certainly could have asked Nuttall if they had any doubt as who the author was. Pennell (1936) gives the impression that Nuttall was a rather shy and inhibited person. It would seem out of character for a man of such temperament to assume responsibility for the names in Fraser's Catalogue if he was not in any way responsible for them. It is inconceivable to me that someone else could have assigned names to Nuttall's own collection without Nuttall stating this fact later. If Nuttall was not associated with the entities, why should he later accept some of them as his unless he actually had given the names to the Fraser Brothers, helped someone who was employed by them to prepare the Catalogue, or perhaps have written the text himself. It should be noted here, however, that I do not say that Nuttall was the author of the paper, although Greene (1890) suggests this after seeing the copy of the Catalogue in Philadephia. What I do contend is that Nuttall is the author of the names in the paper, and thus, as provided by the International Code (1966), the species should be cited as "Nutt. In Fras."
 

Shinners (1956) goes to great length to show that several of the names in Fraser's Catalogue were not claimed by Nuttall in his later publications, and while this is true, I suspect that it was for reasons other than those given by Shinners. Some species were found to have been adequately described between 1813 and 1818 with names acceptable to Nuttall, but for those that were not, he used his own names that he had published in the Fraser's Catalogue. Nuttall occasionally cited only "Frasers Catalogue" and did not give himself credit for the name. Shinners uses this argument to show that Nuttall was not the author. The species with which I am most familiar that was published in the Catalogue is Eriogonum flavum. The name is credited to “Fras. Catal. 1813” and not starred as a new species in Nuttall's 1818 book, The Genera of North American Plants, and for this reason, Shinners suggested that Nuttall did not consider this ispecies as his. Nothing is further from the truth. In a detailed paper on this species in particular, and the genus Eriogonum in general, which Nuttall published in 1817, he makes a point of stating that he published the name E. flavum in Eraser's Catalogue. The reason why Nuttall simply stated “Fras. Catal. 1813.” instead of “T. N. in Fras. Catal. 1813.,” as he sometimes did, was probably to save space. To say that Nuttall was not the author of Fraser's Catalogue I believe is on more tenuous bases than to say that he was indeed the sole author. As no one else has come along to claim authorship of the species published in the list, this, I believe, proves the point that only Nuttall could have given the names to his own collection.
 

The International Code of Botanical Nomenclature (1966) states in Article 34 that "A name is not validly published … when it is not accepted by the author who published it …" As Nuttall did not specifically reject the names in the Catalogue, they must be considered as published. The arguments put forth by Graustein (1956) are immaterial. She states that under the terms of a contract, Nuttall's journals and observations became the exclusive property of Barton, and Nuttall was not supposed to do anything else but what was specifically stated in the con- tract. Thus, Graustein believes that Nuttall could not legally publish any new species in Fraser's Catalogue, for if he did, he would be breaking the terms of the contract. How- ever, as McKelvey (1955) has pointed out, Nuttall's mere presence with the Overland Astorians as they ascended the Missouri River was breaking his contract with Barton, as was the shipping of his plants to England instead of Philadelphia. Certainly one more step in the breaking of the contract by publishing his findings would not be totally surprising.
Reveal (1968) then goes on to review each name published in Fraser's 1813 Catalogue, rendering an opinion about their validity.

 

Literature Cited:
- Pursh, Frederick, 1814.

Other articles:
• Golden Checklist Flora:  Notes on Juniperus communis depressa, Pursh, 1814;  Glycyrrhiza lepidota, Pursh, 1814;  Agoseris glauca, Pursh, 1816;  Ratibida columnifera, Pursh, 1814;  Pseudoroegneria spicata, Pursh, 1814;  

Pursh, 1814-1816, Publication Details

Pursh (1814-16) is an often cited reference for taxa found in Golden s.l.

Abbreviation Full Title Availability Online
Fl. Sept. Americ.
Pursh Fl. Sept. Americ.
Pursh, Frederick. 1814. Flora Americae Septentrionalis; or, A Systematic Arrangement and Description of the Plants of North America. 2 vols. London: White, Cochrane, and Co., 1814. URL: https://biodiversitylibrary.org/item/1987

There are forty-four taxa named by Pursh that are known to occur in Golden s.l..

  • Juniperus communis var. depressa Pursh. Common juniper
  • Calligonum canescens Pursh (=Atriplex canescens (Pursh) Nutt.)
  • Musineon divaricatum (Pursh) Raf.. Leafy Wild Parsley
  • Agoseris glauca (Pursh) Raf.. Pale Goat-Chicory, as Troximon glaucum Pursh
  • Arnica fulgens Pursh. Shining Leopardbane
  • Balsamorhiza sagittata (Pursh) Nutt.. Arrow-Leaf Balsamroot
  • Dieteria canescens (Pursh) Nutt.. Hoar False Tansy-Aster
  • Erigeron compositus Pursh. Dwarf Mountain Fleabane
  • Gaillardia aristata Pursh. Great Blanket-Flower
  • Grindelia squarrosa (Pursh) Dunal. Curly-Cup Gumweed
  • Gutierrezia sarothrae (Pursh) Britt. & Rusby. Kindlingweed
  • Heterotheca villosa (Pursh) Shinners. Hairy False Golden-Aster
  • Lygodesmia juncea (Pursh) D. Don ex Hook.. Rush Skeleton-Plant
  • Nothocalais cuspidata (Pursh) Greene. Wavy-Leaf Prairie-Dandelion
  • Xanthisma spinulosum (Pursh) D.R. Morgan & R.L. Hartman. Lacy Sleepy Daisy
  • Mertensia lanceolata (Pursh) DC.. Prairie Bluebells
  • Phacelia heterophylla Pursh. Variable-Leaf Scorpion-Weed
  • Cleome serrulata Pursh. Rocky Mountain Beeplant
  • Euphorbia marginata Pursh. Snow-on-the-Mountain
  • Amorpha canescens Pursh. Leadplant ???
  • Glycyrrhiza lepidota Pursh. American Licorice
  • Lupinus argenteus Pursh. Silver-Stem Lupine
  • Oxytropis lambertii Pursh. Stemless Locoweed
  • Pediomelum argophyllum (Pursh) J. Grimes. Silver-Leaf Indian-Breadroot
  • Psoralidium lanceolatum (Pursh) Rydb.. Lemon scurfpea
  • Psoralidium tenuiflorum (Pursh) Rydb.. Slimflower scurfpea
  • Ribes aureum Pursh. Golden Currant
  • Linum lewisii Pursh. Prairie Flax
  • Mentzelia decapetala (Pursh ex Sims) Urb. & Gilg ex Gilg. Gumbo-Lily
  • Mentzelia nuda (Pursh) Torr. & A. Gray. Goodmother
  • Claytonia lanceolata Pursh. Lance-Leaf Springbeauty
  • Lewisia rediviva var. rediviva Pursh. Bitter root
  • Mirabilis linearis (Pursh) Heimerl. Narrow-Leaf Four-O'clock
  • Oenothera albicaulis Pursh. White-Stem Evening-Primrose
  • Castilleja sessiliflora Pursh. Great Plains Indian-Paintbrush
  • Mimulus lewisii Pursh. Great Purple Monkey-Flower
  • Ipomopsis aggregata (Pursh) V. Grant. Scarlet Skyrocket
  • Rumex venosus Pursh. Veiny Dock
  • Androsace occidentalis Pursh. Western Rock-Jasmine
  • Clematis hirsutissima Pursh. Sugarbowls
  • Ranunculus cymbalaria Pursh. Alkali Buttercup
  • Geum triflorum Pursh. Old-Man's-Whiskers
  • Scrophularia lanceolata Pursh. Lance-Leaf Figwort
  • Viola nuttallii Pursh. Yellow Prairie Violet
  • Pseudoroegneria spicata (Pursh) A. Löve. Bluebunch-Wheat Grass

Literature Cited:
- Rafinesque, C. S., 1818b.  

Rafinesque (1818) had the following to say in his review of Pursh (1814) Flora Americae Septentrionalis.

Original Text
We have abundant proofs that [Pursh] has introduced many new species, not discovered by, nor belonging tohimself, and without the leave of the owners. Mr. Bradbury, for instance, has loudly complained to us, against him, for having so many of his new plants collected in the Missouri, without his permission, which he meant to publish them himself ; but the blame, if any, appears to lay with those friends who put the plants into the hands of Mr> Pursh, since the author of a general Flora is perfectly at liberty to avail himself of all the materials which come to his knowledge, and ought rather to be blamed for omitting them than otherwise.
It is advisable therefor for the discoverers of new plants to publish them speedily themselves, or keep them out of sight, and buried, as misers do their gold.
Many new plants are stated by Mr. P. to grow on the Mississippi and Missouri, without reference to their discoverers, and as he never was there himself, it is to be presumed they were collected by Mess. Lewis or Nuttall, and probably the latter ; the circumstance from which they were derived is illiberal and disingenuous.
After all, let no one suppose that we despise the labours of Mr. P : far from it. Few can value them more than we do ; we shal at all times be happy to give credit to any botanist for his real personal exertions : for such, and for the compilation of his Flora, Mr. Pursh does really deserve our thanks, notwithstanding so many errors and omissions. We therefore with him complete success in his future labours, and particularly in the completion of a Flora Canadensis, which we understand he has lately undertaken on the spot.

C. S. R.

Literature Cited:
- Robin, C. C. (Charles-Ce´sar), 1817.  

The following is an intentionally empty table …

Abbreviation Full Title Availability Online
Fl. Ludov. Florula Ludoviciana ; or, A Flora of the State of Louisiana URL: https://www.biodiversitylibrary.org/item/275247

 

Literature Cited:
- Fraser, John, 1813.
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Nuttall, 1841, publication details;  Delphinium carolinianum virescens, Nuttall, 1919;  Notes on Erysimum asperum, Nuttall, 1818;  Amelanchier alnifolia, Nuttall, 1818;  Glycyrrhiza lepidota, Nuttall, 1818;  Sphaeralcea coccinea, Nuttall, 1818;  Mentzelia multiflora, Nuttall, 1818;  Collomia linearis, Nuttall, 1818;  Dracocephalum, Nuttall, 1818;  Solanum triflorum, Nuttall, 1818;  Orthocarpus luteus, Nuttall, 181;  Ambrosia tomentosa, Nuttall, 1818;  Notes on Artemisia ludoviciana;  Cirsium undulatum, Nuttall, 1818;  Cyclachaena xanthifolia, Nuttall, 1818;  Notes on Erigeron pumilis;  Notes on Erigeron pumilus, Nuttall, 1818;  Notes on Erigeron pumilis, Nuttall, 1818;  Ratibida columnifera, Nuttall, 1818;  Senecio integerrimis, Nuttall, 1818;  Solidago speciosa, Nuttall, 1818;  Bouteloua gracilis, Nuttall, 1818;  Elymus elymoides, Nuttall, 1818;  Notes on Hordeum pusillum;  Schedonnardus paniculatus, Nuttall, 1818;  

Nuttall, 1818, Publication Details

Abbreviation Full Title Availability Online

Nutt. 
Gen. Am.
Nuttall, Thomas. 1818. The Genera of North American Plants and a catalogue of the species to the year 1817. 2 Vols.. Philadelphia: 1818. http://www.biodiversitylibrary.org/bibliography/290

Types published by Nuttall in this volume that are found in Golden s.l.

  • Atriplex canescens (Pursh) Nutt. (Calligonum canescens Pursh)
  • Ambrosia tomentosa Nutt.
  • Artemisia ludoviciana Nutt. Silver Wormwood.
  • Cirsium undulatum (Nutt.) Spreng. Wavy Leaved Thistle, as Cnicus undulatus Nutt.
  • Erigeron pumilis Nutt. Shaggy Fleabane.
  • Iva xanthiifolia Nutt. Carelessweed, =Cyclachaena xanthiifolia (Nutt.) Fesen.
  • Senecio integerrimus Nutt. Columbia Ragwort.
  • Solidago speciosa Nutt. Showy Goldenrod.
  • Collomia linearis Nutt. Tiny Trumpet.
  • Solanum triflorum Nutt. Cutleaf Nightshade.
  • Orthocarpus luteus Nutt. “Yellow Owls Clover.”
  • Delphinium virescens Nutt. (=Delphinium carolinianum subsp. virescens (Nutt.) R. E. Brooks)
  • Buchloe dactyloides (Nutt.) Engelm. (=Bouteloua dactyloides (Nutt.) Columbus) Buffalo Grass
  • Hordeum pusillum Nutt. Little Barley, on the plains of the Missouri.
  • Munroa squarrosa (Nutt.) Torr. False Buffalograss.
  • Schedonnardus paniculatus (Nutt.) Trel. Tumblegrass, as Lepturus paniculatus.

    Nuttall types published elsewhere.

  • Sphaeralcea coccinea (Nutt.) Rydb., published in Fraser (1813) as Malva coccinea.
  • Oenothera cespitosa Nutt. Tufted Evening Primrose, published in Fraser's (1813) Catalogue.
  • Monarda pectinata Nutt. “Plains Beebalm”, published in 1848, Plants Collected by Gambel.
  • Physalis longifolia Nutt. “Longleaf Groundcherry,” published in 1834, Flora of Arkansas
  • Agoseris parviflora (Nutt.) D. Dietr. “Steppe Goat-Chicory,” published in 1841 as Troximon parviflorum
  • Antennaria parvifolia Nutt. “Small-Leaf Pussytoes.” published in 1841.
  • Coreopsis tinctoria Nutt. Golden Tickseed, published in Arkansa report, 1821.
  • Crepis occidentalis Nutt. Largeflower Hawksbeard, published in report of plants collected by Nathaniel Wyeth, 1834a
  • Helianthus pumilus Nutt. Little Sunflower. Published in 1841.
  • Heliomeris multiflora Nutt. Showy Golden Eye. Published in 1841.
  • Ratibida columnifera (Nutt.) Woot. & Standl. (Syn: Rudbeckia columnifera Nutt.) “Upright Prairie Coneflower.” Published in Fraser (1813) Catalogue, with the name attributed to Nuttall.
  • Senecio plattensis Nutt. =Packera plattensis (Nutt.) W. A. Weber & Á. Löve, published in Nuttall (1841).
  • Solidago missouriensis Nutt. Missouri Goldenrod, published in report of plants collected by Nathaniel Wyeth, 1834a.
  • Solidago nana Nutt. Baby Goldenrod, published in Nuttall (1841).
  • Leucocrinum montanum Nutt. Ex A. Gray. Star Lily, published by Gray (1848).
  • Aristida purpurea Nutt. Purple Threeawn, published in Nuttall (1834) account of his trip to the Arkansa Territory.
  • Muhlenbergia montana (Nutt.) Hitchc. Mountain Muhly, published as Calycodon montanus by Nuttall in his 1848 account of the collections by Gambel.

 

Literature Cited:
- Rafinesque, C. S., 1818b.

Other articles:
• Golden Checklist Flora:  Ratibida columnifera, Rafinesque, 1818;  

American Monthly Magazine and Critical Review, Rafinesque, 1817-1818

One of Rafinesque's several publication series.

Abbreviation Full Title Availability Online
Amer. Monthly Mag. & Crit. Rev. American Monthly Magazine and Critical Review. Vols. 1-4, 1817-18 [1819] Available through Google Books.

 

Literature Cited:
- Elliott, Stephen, 1821-1824.

Other articles:
• Golden Checklist Flora:  Brickellia, Elliott, 1823;  

Elliott, 1821-1824, Publication Details

Elliott (1821-1824) published his Sketch of the Botany of South Carolina and Georgia.

Abbreviation Full Title Availability Online

A sketch of the botany of South Carolina and Georgia
Sketch Bot. S. Carolina
Botany of South-Carolina and Georgia
Elliott's Botany 
A sketch of the botany of South Carolina and Georgia. Elliott, Stephen, 1771-1830. Hoff, John, , printer. Charleston, S.C. :J.R. Schenck,1821-24. URL: https://www.biodiversitylibrary.org/bibliography/9508

 

Literature Cited:
- Nuttall, Thomas, 1821.

Other articles:
• Golden Checklist Flora:  Coreopsis tinctoria, Nuttall, 1821;  

Nuttall, 1821, Publication Details

Nuttall (1821) describes his nearly disastrous exploration of the Arkansas River in 1818-1820.

Abbreviation Full Title Availability Online
J. Acad. Nat. Sci. Philadelphia 2: (1821). A Description of some new species of Plants, recently introduced into the gardens of Philadelphia, from the Arkansa territory. Journal of the Academy of Natural Sciences of Philadelphia. 2: URL: https://www.biodiversitylibrary.org/page/26373612#page/124/

 

Literature Cited:
- Roemer, Johann Jacob, and Josef August Schultes, 1817-1830.
- Sprengel, Curt Polycarp Joachim, 1826.  

Sprengel, 1826, Publication Details.

Abbreviation Full Title Availability Online
Syst. Veg., ed. 16 Systema vegetabilium [Caroli Linnaei ... ]. Editio decima sexta. Gottingae URL: https://www.biodiversitylibrary.org/bibliography/822

Although purporting to be 16th edition Linnaeus' Systema vegetabilium, this work must be attributed to Sprengel. In fact this should count as the 17th edition, as the real 16th edition that of Roemer & Schultes.

 

Other articles:
• Golden Checklist Flora:   1827;  Eriogonum umbellatum;  Acer glabrum;  

Torrey, 1828, Rocky Mountain Plants of Dr. Edwin P. James.

Torrey's final account of plants collected by Edwin James in 1820.

Abbreviation Full Title Availability Online

Ann. Lyceum Nat. Hist. New York ii. (1828)
Some account of a Collection of Plants made during a journey to and from the Rocky Mountains in the summer of 1820, by Edwin P. James, M. D. Assistant Surgeon U. S. Army. URL: https://www.biodiversitylibrary.org/part/240415

   

Rafinesque, 1833, Publication Details

Abbreviation Full Title Availability Online

Atlantic J
Atlantic Journal, and Friend of Knowledge. Philadelphia, PA. http://www.biodiversitylibrary.org/item/104571

 

Other articles:
• Golden Checklist Flora:  Clematis columbiana, Nuttall, 1834;  Phlox longifolia, Nuttall, 1834;  Notes on Crepis occidentalis;  Notes on Solidago missouriensis, Nuttall, 1834;  Maianthemum racemosum amplexicaule, Nuttall, 1834a;  

Nuttall, 1834a, Wyeth Expedition

Nuttall (1834a) described plants brought back by Nathaniel Wyeth from his expedition to the Oregon Territory.

Abbreviation Full Title Availability Online
J. Acad. Nat. Sci. Philadelphia 7: pp (1834). Journ. Acad. Nat. Sci. Philad., Vol. VII. A Catalogue of a Collection of Plants made chiefly in the Valleys of the Rocky Mountain or Northern Andes, towards the sources of the Columbia River, by Mr. Nathaniel B. Wyeth, and described by T. Nuttall Journal of the Academy of Natural Sciences of Philadelphia. Philadelphia, PA

Read February 18, 1834.

URL: https://www.biodiversitylibrary.org/item/79407#page/39
New names of plants found in Colorado that were published in this paper.
  • Phlox longifolia Nutt. “Longleaf Phlox.”
  • Crepis occidentalis Nutt. Largeflower Hawksbeard.
  • Solidago missouriensis Nutt. Missouri Goldenrod.

   

Nuttall, 1834b, Flora of the Territory of Arkansas

The following is an intentionally empty table …

Abbreviation Full Title Availability Online
Trans. Amer. Philos. Soc. ser. 2, 5: 145 (1835). Article VI. Collections towards a Flora of the Territory of Arkansas. By Thomas Nuttall. Read before the American Philosophical Society April 4, 1834. URL: https://www.biodiversitylibrary.org/item/110958

  • Physalis longifolia Nutt. “Longleaf Groundcherry.”
  • Aristida purpurea Nutt. Purple Threeawn.

 

Literature Cited:
- Fresenius, Georg, 1836.

Other articles:
• Golden Checklist Flora:  Cyclachaena xanthifolia, Fresnius, 1836;
Full Size ImageFootnotes from Index Seminum in which Cyclachaena is proposed  

Fresenius, 1836, Publication Details

Johann Baptist Georg (George) Wolfgang Fresenius (1808-1866) was a German physician and botanist, known for his work in the field of phycology. He was a native of Frankfurt am Main.

He studied medicine at the Universities of Heidelberg, Würzburg and Giessen, earning his doctorate at the latter institution in 1829. Afterwards he settled in Frankfurt am Main, where he worked as a general practitioner of medicine while maintaining an active interest in botany.

As a student in Heidelberg and afterwards, he studied botany with his friend George Engelmann (1809-1884), who later became a renowned German-American botanist. From 1831 Fresenius was curator of the Senckenberg herbarium and a teacher at the Senckenberg Research Institute (Forschungsinstitut Senckenberg). With his student Anton de Bary (1831–1888), he conducted microscopic investigations of algae and fungi. He died in Frankfurt on 1 December 1866 at the age of 58.

Abbreviation Full Title Availability Online
Index Sem. (Frankfort/M) Index Seminum. Frankfort am Main. Universita¨t Frankfurt am Main. Botanischer Garten. URL:
https://www.biodiversitylibrary.org/bibliography/84379
https://www.biodiversitylibrary.org/item/160600

In the online publication, the various parts are mixed up as to the date order.

Full Size Image
Page 1 from Index Seminum Frankfort/Am Main

 

Literature Cited:
- Lawson, Peter & Son, 1836.

Other articles:
• Golden Checklist Flora:  Notes on Pinus ponderosa, Lawson, 1836;  

Lawson, 1836, Agriculturist's Manual

Published in Edinburgh; see Linzer Biol. Beitr. 28: 1048. 1996

Abbreviation Full Title Availability Online

Agric. Man.
The Agriculturalist's manual : being a familiar description of the agricultural plants cultivated in Europe, including practical observations respecting those suited to the climate of Great Britain, and forming a report of Lawson's Agricultural Museum in Edinburgh URL: https://www.biodiversitylibrary.org/item/165048

Names in this publication:

  • Pinus ponderosa Douglas ex Lawson & C. Lawson. “Ponderosa Pine”

 

Other articles:
• Golden Checklist Flora:  Clematis columbiana, Torr. & A. Gray, 1838;  Delphinium carolinianum virescens, Torrey & A. Gray, 1838;  Rorippa sinuata, Torrey & Gray, 1838;  Holodiscus dumosus, Torr. & A. Gray;  Vicia ludoviciana, Torr. & A. Gray, 1838;  

Torrey & A. Gray, 1838-1843, Flora of North America

Torrey & A. Gray (1838-1943) in two volumes …

Abbreviation Full Title Availability Online

Fl. N. Amer.
A flora of North America :containing abridged descriptions of all the known indigenous and naturalized plants growing north of Mexico, arranged according to the natural system URL: https://www.biodiversitylibrary.org/bibliography/9466
Names published in this volume:
  • Vicia ludoviciana Nutt. ex Torr & A. Gray

 

Literature Cited:
- Nuttall, Thomas, 1840-1841.

Other articles:
• Golden Checklist Flora:  Nuttall, 1818, publication details;  Troximon glaucum, Nuttall, 1841;  Agoseris parviflora, Nuttall, 1841;  Franseria discolor, Nuttall, 1841;  Antennaria parvifolia, Nuttall, 1841;  Crepis occidentalis, Nuttall, 1814;  Notes on Helianthus pumilus;  Notes on Heterotheca foliosa, Nuttall, 1841;  Notes on Packera plattensis, Nuttall, 1841;  Solidago nana, Nuttall, 1841;  

Nuttall, 1841, Tour across the Continent to the Pacific, a Residence in Oregon, and a Visit to the Sandwich Islands

In 1834, Nuttall resigned his post and set off west again on an expedition led by Nathaniel Jarvis Wyeth, this time accompanied by the naturalist John Kirk Townsend. They travelled through Kansas, Wyoming, and Utah, and then down the Snake River to the Columbia. Nuttall then sailed across the Pacific Ocean to the Hawaiian Islands in December. He returned in the spring of 1835 and spent the next year botanizing in the Pacific Northwest, an area already covered by David Douglas. On the Pacific coast, Nuttall heard of the ship Alert leaving San Diego in May 1836, bound for Boston.

Abbreviation Full Title Availability Online

Trans. Amer. Philos. Soc. ser. 2, 7 (1841).
Descriptions of new Species and Genera of Plants in the natural Order of the Compositae, collected in a Tour across the Continent to the Pacific, a Residence in Oregon, and a Visit to the Sandwich Islands and Upper California, during the Years 1834 and 1835. Transactions of the American Philosophical Society Held at Philadelphia for Promoting useful Knowledge. Series 2, Volume 7. Philadelphia. URL: https://www.biodiversitylibrary.org/item/95918

Nuttall types published in this volume:

  • Agoseris parviflora (Nutt.) D. Dietr., published in 1841 as Troximon parviflorum
  • Antennaria parvifolia Nutt. Small-Leaf Pussytoes.
  • Brickellia grandiflora (Hook.) Nutt. Tasselflower Brickellbush.
  • Helianthus pumilus Nutt. Little Sunflower.
  • Heliomeris multiflora Nutt. Showy Golden Eye.
  • Psilochenia occidentalis Nutt. (Syn: Crepis occidentalis Nutt.)
  • Senecio plattensis Nutt. =Packera plattensis (Nutt.) W. A. Weber & Á. Löve
  • Solidago nana Nutt. Baby Goldenrod.

No mention is made of any Salix.

 

Other articles:
• Golden Checklist Flora:  Notes on Salix exigua;  

Nuttall, 1842-1849, North American Sylva

Nuttall extended Michaux's three-volume North American Sylva, with three additional volumes, containing all the forest trees discovered in the Rocky Mountains, the territory of Oregon, down to the shores of the Pacific and into the confines of California, as well as in various parts of the United States.

Abbreviation Full Title Availability Online
    N. Amer. Sylv. [Nuttall]
The North American Sylva; or, A description of the forest trees of the United States, Canada, and Nova Scotia, considered particularly with respect to their use in the arts, and their introduction into commerce; to which is added a description of the most useful of the European trees … Tr. from the French of F. Andrew Michaux …, The. Philadelphia URL: https://www.biodiversitylibrary.org/item/92584

Dates


Vols. 1-3 (issued as vols. 4-6 of a 6-volume series), 1842-1849
1(1): 1-56. med 1842
1(2): 57-136. Jul-Dec 1842
2: 1-123. 1846
3: 1-148. 1849.

Remarks

With Illustrated by 122 finely colored plates. By Thomas Nuttall … The whole forming six volumes, and comprising 278 plates --Note on t.-p.

Names first published in these volumes:

  • Salix exigua Nutt.

 

Literature Cited:
- Fremont, John C., 1845.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum alatum, Frémont 1845;  

Fremont, 1845, Exploring Expedition to the Rocky Mountains

The following is an intentionally empty table …

Abbreviation Full Title Availability Online

Frémont Rep.
Report of the Exploring Expedition to the Rocky Mountains in the year 1842 and to Oregon and North California in the years 1843-'44. URL: https://www.biodiversitylibrary.org/item/101327

The Catalogue of Plants for Frémont's 1842 expedition starts on page 75. The Polygonaceae is listed on page 96, those listed being Eriogonum ovalifolium Nutt., E. caespitosum Nutt., E. umbellatum Torr., E. fremontii n. sp. (=E. brevicaule Nutt.), and E. annuum Nutt.

The descriptions of plants for Frémont's 1843-'44 expedition begins on page 311. The Eriogonum described include E. inflatum Torr. and Frém., E. reniforme Torr. and Frém., and E. cordalum Torr. and Frém. (=E. cordatum Torr & Frém., and is impossible to place unequivocally, per FNANM.)

No mention is made of E. alatum though Frémont and Torrey were aware of it.

 

Other articles:
• Golden Checklist Flora:  Agoseris glauca, Dietrich, 1847;  Agoseris parviflora, Dietrich, 1847;  

Dietrich, 1847, Publication Details

David Nathaniel Friedrich Dietrich (1800 – 1888) was German botanist and gardener. In 1828 Dietrich worked as a botanical gardener in Jena. In 1836 he received his doctorate at the University of Jena, and later served as a curator at the botanical garden in Jena. Dietrich wrote pamphlets on poisonous plants, mosses, and forest flora and fauna of Germany as well as several botanical encyclopedias. His five-volume, 1839-1852 Synopsis Plantarum, cited here, included about 80,000 species and 524 genera. The five-volume Flora of Germany published from 1833 to 1864 contains 1150 colored panels. The two-volume Forst Flora and the 476 booklets of the comprehensive Flora Universalis are his most famous work (Wikipedia, 2020).

Abbreviation Full Title Availability Online

Syn. Pl.
Synopsis Plantarum seu enumeratio systematica plantarum…. 4. Issued in five volumes: Vol. 1, Jul 1839, Vol. 2, 1-20 Dec 1840, Vol. 3, late Dec 1842, Vol. 4, early Jan 1847, Vol. 5, Apr 1852 URL: https://www.biodiversitylibrary.org/page/688583

 

Literature Cited:
- Roemer, M. J., 1847.

Other articles:
• Golden Checklist Flora:  Saskatoon Serviceberry;  

Roemer, 1847, Fam. Nat. Syn. Monogr.

Max Joseph Roemer (1791–1849) was a German botanist who worked in Weimar. He served as a Landrichter (country judge) in the Bavarian town of Aub, afterwards working as a private scientist in Würzburg. He is the taxonomic authority of the genera Heteromeles, Pyracantha and Erythrocarpus as well as of numerous plant species.

Abbreviation Full Title Availability Online

Fam. Nat. Syn. Monogr.
Familiarum naturalium regni vegetabilis synopses monographicae; seu, Enumeratio omnium plantarum hucusque detectarum secundum ordines naturales, genera et species digestarum, additis diagnosibus, synonymis, novarumque vel minus cognitarum descriptionibus.

Monographic synopsis of the families of plants; or, An enumeration of all the natural orders of plants according to those that hitherto have been detected, the genera and species in four parts, with the addition of diagnoses, synonyms, and descriptions of new or less known species.


URL: https://www.biodiversitylibrary.org/bibliography/49482
Fascicle 1-2: https://www.biodiversitylibrary.org/item/104538
Fascicle 3-4: https://www.biodiversitylibrary.org/item/104545

 

Literature Cited:
- Emory, William H., 1848.  

Emory, 1848, Publication Details

Abbreviation Full Title Availability Online

Emory's Rep.
Emory, Notes milit. Reconn. 1848
Emory, William H. 1848. Notes of a military reconnaissance, from Fort Leavenworth, in Missouri, to San Diego, in California, including parts of the Arkansas, Del Norte, and Gila rivers. Washington: Wendell and Van Benthuysen, printers, 1848. https://www.biodiversitylibrary.org/item/99257

 

Literature Cited:
- Nuttall, Thomas, 1848b.

Other articles:
• Golden Checklist Flora:  Heliomeris multiflora;  Gray, 1849, publication details;  Notes on Quercus gambelii;  Notes on Eucycla, Nuttall, 1848;  Notes on Eriogonum effusum;  Mentzelia multiflora, Nuttall, 1847;  Ipomopsis spicata, Nuttall, 1848;  Monarda pectinata, Nuttall, 1848;  

Nuttall, 1848, Plants collected by William Gambel

This paper describes collected by William Gambel in the vicinity of Santa Fe, New Mexico, between June and September, 1841, and in February-July, 1842, in California. Nuttall also used the paper to describe additional plants he collected on his 1834 trip to Oregon Territory. The publication appears in two forms.

Abbreviation Full Title Availability Online
  • Nutt. Pl. Gamb. in Proceedings, Acad, Philad. vol. 4.
Descriptions of Plants collected by William Gambel, M. D., in the Rocky Mountains and Upper California. URL: https://www.biodiversitylibrary.org/item/84785
Abbreviation Full Title Availability Online
  • Nutt. Pl. Gamb. In Jour, Acad, Philad. N. ser. 1
Art. XIII. — Descriptions of Plants collected by William Gambel, M. D., in the Rocky Mountains and Upper California. URL: https://www.biodiversitylibrary.org/item/95920

A related publication would be Gray's (1849) Planta Fendlerianae Novi-Mexicanae.

Names published in this volume that Gambel collected:

  • Quercus gambelli Nutt. “Gambel's Oak.”

Names published int his volume that Nuttall collected:

  • Eriogonum effusum Nutt. “Spreading Buckwheat.”
  • Eucycla Nutt., now treated as a subgroup of Eriogonum Michaux.

No mention is made of any willows.

 

Other articles:
• Golden Checklist Flora:  Abronia fragrans, Hooker, 1853;  

Hooker's journal of botany and Kew Garden miscellany, 1849-1857

The following is an intentionally empty table …

Abbreviation Full Title Availability Online
Hooker's J. Bot. Kew Gard. Misc. Hooker's journal of botany and Kew Garden miscellany. URL: https://www.biodiversitylibrary.org/bibliography/236

Names published here:

  • Vol. 5, Abronia fragrans Nutt. ex Hook.

 

Literature Cited:
- Gray, Asa, 1849.

Other articles:
• Golden Checklist Flora:  Nuttall, 1848, publication details;  Holodiscus dumosus, Gray, 1849;  Sphaeralcea coccinea, Gray, 1849;  Mentzelia multiflora, Gray, 1849;  Gray, 1849;  Brickellia californica, Gray, 1849;  Notes on Erigeron tracyi, Gray, 1849;  Packera fendleri, Gray, 1849;
• Field Notes:  Gray, A., 1849;  

Gray, 1849, Plantae Fendlerianae Novi-Mexicanae

This publication describes some of the plants collected by Augustus Fendler in the vicinity of Santa Fe, New Mexico, October 1846 to August 1847. It ends with the phrase “To be continued,” though it was not.

Abbreviation Full Title Availability Online

Gray, Pl. Fendl.
Mem. Am. Acad. 4
Plantae Fendlerianae Novi-Mexicanae; An Account of a Collection of Plants made chiefly in the Vicinity of Santa Fe, New Mexico, by Augustus Fendler; with Descriptions of the New Species, Critical Remarks, and Characters of other undescribed or little known Plants from surrounding Regions. Memoirs of the American Academy. IV(I):1-116. URL:
https://www.biodiversitylibrary.org/item/55373
https://www.biodiversitylibrary.org/item/49963

 

Literature Cited:
- Gray, Asa, 1852-1853.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum alatum, Gray, 1852;  Notes on Oenothera brachycarpa;  

Gray, 1852-1853, Plantæ Wrightianæ

Gray (1852-1853) account of plants collected by Charles Wright.

Abbreviation Full Title Availability Online

Plantæ Wrightianæ

Wright

Plantae Wrightianae: Texano — Neo – Mexicanae: an account of a collection of plants made by Charles Wright in an Expedition from Texas to New Mexico, in the Summer and Autumn of 1849, with Critical Notices and Characters of Other New or Interesting Plants from Adjacent Regions, &c. by Asa Gray, M. D., Fisher Professor of Natural History in Harvard University. URL: https://www.biodiversitylibrary.org/item/55374
 

PLANTAE WRIGHTIANAE TEXANO-NEO-MEXICANAE:
AN ACCOUNT
of
A COLLECTION OF PLANTS MADE BY CHARLES WRIGHT, A. M.,
IN AN EXPEDITION FROM TEXAS TO EL PASO, NEW MEXICO, IN THE
SUMMER AND AUTUMN OF 1840.
Mr. Charles Wright, who has for several years past devoted much attention to the botany of Texas, returned to Galveston from the North in the spring of 1849, and proceeded to San Antonio, purposing to avail himself of the opportunity afforded by the movement of a body of United States troops from this place across the country to El Paso, in Southern New Mexico, to investigate the natural history, and especially the botany, of this hitherto untrodden region. A recommendation from the War Department, that all proper facilities should be furnished to Mr. Wright, in furtherance of his arduous and entirely scientific undertaking, procured for him only the free transportation of his paper for preserving specimens, and of the collections he was enabled to make. This favor he owes to the kindness of Captain French, the quartermaster of the expedition, to whom and to Major Henry and Major Van Horn, Mr. Wright desires to express his thanks. The train left the frontier settlement of Castroville about the first of June, and reached El Paso early in September. The remainder of that month was devoted to making collections in the vicinity of that interesting station. Finding that much time would necessarily be lost in passing the long winter in New Mexico, Mr. Wright retraced his steps, and accompanied his rich collections back to Texas by the return train, leaving El Paso in October, and reaching San Antonio late in November. In returning he was enabled to add to his collection some species which had escaped notice during the outward journey, or which were not then in season, as well as largely to increase his collection of seeds, and of living Cactaceous plants. Specimens of the latter have been placed in the hands of Dr. Engelmann, of St. Louis, for examination. The seeds have been divided between the Botanic Garden of Harvard University, under my charge, and the Royal Botanic Gardens at Kew, under the direction of Sir Wm. Hooker.
A full set of the plants here enumerated or described is retained in my own herbarium ; another becomes the property of the Smithsonian Institution, which has efficiently patronized this exploration. It will there form, with similar sets of the collections of Fendler and Lindheimer, made in New Mexico and Texas, the nucleus of an important and authentic North American herbarium, destined to be enriched, I trust, by continued accessions, especially from our newly-acquired ter- ritories, until it shall comprise representatives of our whole flora, and specimens of all the vegetable products of our wide country.
Another set of these plants will be found in the herbarium of John A. Lowell, Esq., of Boston, who has liberally patronized Mr. Wright's scientific explorations. The others, eight or nine in number, are about to be issued to the subscribers who have applied for them.
The numbers prefixed to the names are those under which the specimens are distributed. Those marked with a †, in place of a number, were collected in single specimens, or at least not in sufficient quantity for distribution.
The whole will give a good idea of the vegetation, and consequently of the climate, general character, and capabilities, of the region traversed. I append, from time to time, notices or characters of plants gathered by other collectors in adjacent regions, especially by Dr. Wislizenus in the valley of the Rio Grande and in Chihuahua, and by the indefatigable Dr. Gregg * in the same district and in the northern provinces of Mexico, — chiefly from materials obligingly furnished by Dr. Engelmann.
Orders or genera elaborated by Dr. Engelmann, Dr. Torrey, Mr. Bentham, or others, have the name of the author preflxed.
A proper account of the topography and physical character of the region traversed by the United States troops in their march from Texas to New Mexico will doubtless be officially published, before the printing of this memoir is com- pleted. It is therefore unnecessary for me to attempt to compile any such account from Mr. Wright's disjoined and necessarily imperfect memoranda.
A. GRAY.
Harvard University, Cambridge, May, 1850.
* Tidings of the lamented death of this most assiduous collector, in California, have just been received.

 

Literature Cited:
- Sitgreaves, L., 1853.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum alatum, Sitgreaves, 1853;

Locations: Zuni River.  

Sitgreaves, 1853, Expedition Down the Zuni and Colorado Rivers

Sitgreaves, 1953 …

Abbreviation Full Title Availability Online

Sitgr., Rep. Exped. Zuni & Colorado Rivers

L. Sitgreaves, Rep. Exped. Zuni Colorado Rivers

Report of an expedition down the Zuni and Colorado Rivers, by Captain L. Sitgreaves, Corps Topographical Engineers. URL: https://www.biodiversitylibrary.org/item/124223

Report on the Natural History of the country passed over by the exploring expedition under the command of Brevet Captain L. Sitgreaves, U. S. Topographical Engineers, during the year 1851. By S. W. Woodhouse, M.D., Surgeon and Naturalist to the Expedition. … pae 31

Mr. Wright, an enterprising botanist, has passed over this route several times, and the plants have been described by Doctors Torrey and Gray, many of which have been already published by them in the Smithsonian Contributions, under the title, “Plantae Wrightianae.”

Zoology. Mammals and Birds, by S. W. Woodhouse, M. D. Reptiles, by Edward Hallowell, M. D. Fishes, by Prof. S. F. Baird and Charles Girard. … page 41

Botany. by Professor John Torrey. … page 153

For Eriogonum alatum, the citation is “(Torr., l. c.;)” Two entries up, i.e., that for Eriogonum orthocladon, the citation is “(Torr. mss., in D. C. Prodr. ined.:)” which I think means “I, Torrey, described this in a manuscript I gave to DeCandolle for his use in his Prodromo.”

 

Literature Cited:
- Marcy, Randolph B., 1854.  

Marcy, 1854, Publication Details

Abbreviation Full Title Availability Online

Explor. Red River Louisiana
Marcy, Randolph B., 1854. Exploration of the Red River of Louisiana. Exploration of the Red River of Louisiana. Washington: A. O. P. Nicholson, Public Printer, 1854. URL: https://www.biodiversitylibrary.org/item/95580

 

Literature Cited:
- Beckwith, Lt. E. G., 1855.

Other articles:
• Golden Checklist Flora:  Torrey & Gray, 1855. Pac. R. R. Rep.;  

Pacific Railroad Reports, Beckwith

The following is an intentionally empty table …

Abbreviation Full Title Availability Online
  • Pac. R. R. Report
Reports of explorations and surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean URL: https://www.biodiversitylibrary.org/item/268872#page/13

Volume II, first article gives a narrative of the Gunnison expedition, and describes the death of the Captain.

October 25.—Captain Gunnison, with Mr. R. H. Kern, Mr. F. Creutzfeldt, and Mr. Wm. Potter, (guide,) with John Bellows, and a corporal and six men from the escort, left camp at a late hour this morning to explore the vicinity of Sevier lake, supposed to be distant some fifteen or eighteen miles. From Captain Gunnison’s journal I extract the description of the country and operations of the party during the day, written after they had encamped for the night: “I came down the river southwest for nine miles, and then, bearing more west for two miles, concluded to encamp, as the water below might prove too salt. The route was through heavy artemisia for five miles, when we came upon more open plains to the nine-mile point, where we met with sloughs alive with geese, ducks, brant, pelicans, and gulls. A few hawks were careering in the high wind, and the black-eared and black-tailed rabbits were very numerous in the large artemisia.
“The mountains wore all day their white night mantle of snow, and we had squalls from the north, with snow falling on the high mountains on all sides of us. Towards sunset it brightened up a little, and our hunters brought in four ducks of as many different varieties.”
The remainder of the surveying party left under my charge, with the escort under Captain Morris, crossed the river at an excellent ford at the point of our encampment, immediately after the departure of Captain Gunnison, and, agreeably to his request, proceeded up the river in a northeast direction, encamping at a late hour on the river bottom where it is unusually wide. The river at this point makes a long bend in the plain to the south, passing through drifting sand-hills partially covered with artemisia. We had passed southward to avoid these hills in crossing from Cedar spring to the river, and to-day we passed to the west of the largest of them, yet our route was very heavy and the labor severe on our animals. The day, too, was cold and blustering, with occasional slight squalls of snow in the plains, while in the mountains it fell during the greater part of the day. Those of us who were mounted halted frequently—the wagons coming on very slowly—and built fires of sage, which being resinous burnt very freely, with a large flame for the moment, giving out abundant heat. With the setting sun the wind went down, and the night was clear and cold; and at a late hour the pure mountain snows reflected beautifully the clear light of the waning moon, while all around was quietness and repose. The gap by which the Sevier river passes the Un-kuk-oo-ap mountains is called a canon; but at this distance—six miles directly in front of us to the northeast—it appears like anything but a canon passage, although the river may wind from side to side, striking against the foot of the mountains, preventing an easy passage for wagons in its natural state. A large Indian trail, however, passes directly up the river into it. This range of mountains, as seen from our present and last camps, seems to terminate a few miles to the north, leaving a broad, open passage of several miles between it and the succeeding range to the west, in which the waters of the Sevier and Great Salt lakes are divided only by gentle slopes. Distance, 14.27 miles.
October 26.—The morning was clear and cold, and Mount Nebo, seen through the Sevier river gap, with its pure mantle of snow, half enveloped in floating misty clouds, mildly reflecting the rays of the rising sun, presented one of the most beautiful mountain scenes I have ever witnessed. Our animals were kept out to graze until a late hour. At 11 o’clock, however, a party was despatched to ascertain the practicability of the passage around the mountain and thence north to the Great Salt lake ; and, without moving camp again , until the return of Captain Gunnison, it was intended to examine the passage of the Sevier river the following day. But the first party had scarcely proceeded a hundred yards from camp, when it was met by a man, weak and exhausted, reeling breathless into camp, barely able to communicate, by a few broken sentences, as he sank into a seat, the painful intelligence that Captain Gunnison’s party had been surprised in their camp by a large party of Indians, and, he thought, all but himself massacred. Orders were instantly given by Captain Morris, and promptly obeyed by all the men remaining with him of his escort, to replenish their ammunition ; and having brought up and saddled their horses, in thirty minutes they were moving rapidly towards the scene of that fatal disaster, hoping to rescue all who might yet survive and perform the last mournful duties of humanity to those who were known to have fallen.
The man who first reached camp was the corporal of the escort, who had made his escape on his horse, and had ridden him until he could go no further, leaving him at our camp of the previous day, whence he ran on foot fourteen miles—twenty-five in all—arriving, without arms, in the condition I have described. Another of the escort reached camp on horseback, before Capiain Morris’ departure; and two others were met by him in the course of the afternoon, making their way towards camp. The horses of Dr. Schiel, who had accompanied Captain Morris, and of his sergeant and several of his men, gave out during their rapid march, and their riders were left straggling behind; but eventually all arrived safely in camp—-the sergeant and some of the men, from their own folly, in a poor plight.
Before Captain Morris’ departure the train animals were driven in, with the intention at first of removing the train to a more secure place, with the two young gentlemen, Homans and Snyder, and the teamster force, some of whom were unarmed, to guard it. But it was subsequently determined to break up the camp altogether, and move towards a convenient point, where Captain Morris and myself could meet on the following or succeeding day, and take such measures for future operations as circumstances might require, with better means within our reach than we could command at any other point.
Crossing the river at the camp, we took the shortest line to escape from the sand, which proved far heavier than that of the previous day, but it was 7.44 miles in a southeast course, beyond a border of small cedars a mile wide, among which the sand was so drifted that it was only by innumerable windings and contortions of teams and wagons, that we at last escaped from it and reached the plain of grass a mile or two wide, which here lies on the gradual slope of the mountain. Here we encamped. In crossing the sand-hills numerous fresh Indian tracks were seen, notwithstanding the prevalence of a high wind; but the night passed quietly, and at sunrise we travelled southward along the base of the mountain, hoping to escape a continuation of the sand of the previous day, in which we were only partially successful, however, as it continued heavy for ten miles. We then passed a spur of the mountain and changed our course from south to southeast, and struck the trail we had made in going from Cedar spring to the Sevier river, six miles from the spring, at which we found a large camp of Mormons, on their way to settle at Parawan, near Little Salt lake. Here we encamped and turned our animals out to graze on the hills; the 20.93 miles of to-day being the severest day’s labor performed by them, although the day was cool, during the whole course of our long summer journeyings. The last of our animals were not out of harness when Captain Morris arrived, confirming our worst fears for the fate of our late comrades.
Captain Gunnison had encamped early in the afternoon, while the wind and storm were yet fresh, and doubtless feeling the security which men come to indulge after passing long periods of time surrounded by savages without actually encountering them. The abundant grass and fuel of a little nook in the river bottom, sheltered by the high second bank of the river on one side, and thick willows, distant scarcely thirty yards, on two of the others, with the river in front, offered a tempting place of comfort and utility, which was perhaps accepted without even a thought of danger. It was known to the party that a band of Indians was near them, for we had seen their fires daily since entering the valley; but an unusual feeling of security against them was felt, as Captain Gunnison had learned that a recent quarrel, resulting in several deaths, which they had had with emigrants, had terminated, and that notwithstanding this difficulty they had remained at peace with the neighboring settlers, which had been confirmed and guarantied for the future in a “talk” held with some of the Indians of this band, by an agent of the governor of the Territory, during our stay near Fillmore. This information, Captain Gunnison told me before leaving, relieved him from any apprehension he might otherwise have felt regarding this band, and was the reason for his haying asked for so small an escort to accompany him, which he as well as his guide, an experienced citizen of the Territory, deemed sufficient.
The usual precaution of a camp guard had been taken, each of the party (including the commander) in turn having performed that duty during the night. At the break of day all arose and at once engaged in the usual duties of a camp preparatory to an early start, to reach that day the most distant point of exploration fur the present season. The sun had not yet risen, most of the party being at breakfast, when the surrounding quietness and silence of this vast plain was broken by the discharge of a volley of rifles and a shower of arrows through that devoted camp, mingled with the savage yells of a large band of Pah-Utah Indians almost in the midst of the camp; for, under cover of the thick bushes, they had approached undiscovered to within twenty-five yards of the camp-fires. The surprise was complete. At the first discharge, the call to “ seize your arms” had little effect. All was confusion. Captain Gunnison, stepping from his tent, called to his savage murderers that he was their friend; but this had no effect. They rushed into camp, and only those escaped who succeeded in mounting on horseback, and even then they were pursued for many miles. The horse of one fell near camp, tumbling his rider under a bush, where he lay for six or seven hours, while the Indians were passing him on every side, until finally he could no longer hear them near him or in the camp, when he left, and was met soon afterwards by Capt. Morris’ party, which reached the fatal spot just before night. Two Indians were seen a mile or two from camp by Lieutenant Baker and Mr. Potter, brother of the guide, but they were not able to come up with them before night enabled them to escape. The bodies of the slain were not all found at dark, and hope still lingered as a bright fire was built to assure any survivor of safety. But the long weary night, rendered hideous by the howling of wolves, wore away, as this little band of armed men, barely larger than that which had already been sacrificed, lay near the fatal spot, and day dawned only to discover the mutilated remains of their recent comrades, none of them being scalped—a barbarity which some of the tribes on this part of the continent seldom indulge. Borne of their arms were, however, cut off at the elbows, and their entrails cut open; and, the wolves having had access to them during the day and to those exposed during the night, their bodies were in such a condition that it was not deemed possible to bring them away—not even that of Captain Gunnison, who had fallen pierced with fifteen arrows.
The statement which has from time to time appeared (or been copied) in various newspapers of the country since the occurrence of these sad events, charging the Mormons or Mormon authorities with instigating the Indians to, if not actually aiding them in, the murder of Captain Gunnison and his associates, is, I believe, not only entirely false, but there is no accidental circumstance connected with it affording the slightest foundation for such a charge.

 

Literature Cited:
- Torrey, John, and Asa Gray, 1855.

Other articles:
• Golden Checklist Flora:  Beckwith, 1855. Pac. R. R. Rep.;  Notes on Suckleya suckleyana;  Torrey & Gray, 1855;  

Pacific Railroad Reports, Botany, Torrey & Gray

The following is an intentionally empty table …

Abbreviation Full Title Availability Online
  • Pac. R. R. Report
Reports of explorations and surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean URL: https://www.biodiversitylibrary.org/bibliography/11139

Volume II, about in the middle, in the Report on the Botany of the Expedition. Part II of that report covers the Botany of the expedition led by Gunnison.

PART II. Plants collected by Mr. F. Creutzfeldt, under the direction of Captain J. W. Gunnison, U. S. Army, in charge of explorations for a railroad from Fort Leavenworth, by the way of the Kansas and Arkansas rivers, to Bent's Fort; thence by the Huerfano river and Sangre de Crista Pass to the valley of San Luis; thence west from that valley to Grand and Green rivers; thence into the Great Basin, Utah, to the vicinity of the Sevier or Nicollet lake. The collection was commenced at Westport, in Missouri, in June, 1853, and fnished late in October.

 

Literature Cited:
- Engelmann, Georg, and John M. Bigelow, 1856.

Other articles:
• Golden Checklist Flora:   Notes on Opuntia cymochila, Engelmann & Bigelow, 1856;   Notes on Opuntia tortispina, Engelmann & Bigelow, 1856;  

Pacific Railroad Reports, Whipple, Botany, Cactaceae, Engelmann & Bigelow

George Engelmann and John M. Bigelow described to Cactaceae from the Whipple expedition.

Abbreviation Full Title Availability Online
    Descr. Cact. [G. Engelmann & J. Bigelow] iv. 42 (1856)
No. 3, Description of the Cactaceae.
Report of the Botany of the Expedition, Volume 5.
Route Near the Thirty-fifth Parallel, Exploted by Lieutenant A. W. Whipple, Topographical Engineers, in 1853 and 1854.
Reports of explorations and surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean
URL: https://www.biodiversitylibrary.org/item/237542

Two names of interest that were first described in this report are Opuntia tortispina and O. cymochila.

 

Literature Cited:
- DeCandolle, Augustus Pyramus, 1857.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum alatum, DeCandolle, 1857;  

DeCandolle, 1824-1874, Prodromo

Multi-volume …

Abbreviation Full Title Availability Online

D. C. Prodr. ined.
Prodromus systematis naturalis regni vegetabilis, sive, Enumeratio contracta ordinum generum specierumque plantarum huc usque cognitarium, juxta methodi naturalis, normas digesta
URL: https://www.biodiversitylibrary.org/bibliography/286
URL, Vol. 14: https://www.biodiversitylibrary.org/item/109211

 

Literature Cited:
- Hayden, F. V., 1870.

Other articles:
• Golden Checklist Flora:  Notes on Cryptantha virgata, Porter, 1870;  

Hayden, F. V., 1870, Wyoming

Abbreviation Full Title Availability Online

Hayden Report
Hayden's Rep.
Preliminary report of the United States Geological Survey of Wyoming : and portions of contiguous territories, (being a second annual report of progress)

Part V.—Catalogue of Plants, by Thomas C. Porter

URL: . https://www.biodiversitylibrary.org/item/124638

This catalogue embraces the plants collected in Wyoming Territory, by Dr. F. V. Hayden, during the geological survey of 1870 — at Camp Carlin, from July 25 to 30, on the route from Fort D. A. Russell, via Fort Fetterman, Sweetwater, South Pass, Wind Eiver Mountains, and Green River, to Fort Bridger, from August 1 to September 13 ; in the Uinta Mountains, south of Henry's Fork of Green River, in the latter half of September, and on Henry's Fork, in the month of October. To these are added his collection in the North Park, Colorado Territory, August, 1868, and another, made by Mr. B. H. Smith, in the region around the city of Denver, during the summer of 1869.

 

Other articles:
• Golden Checklist Flora:  Notes on Amaranthus blitoides, Watson, 1879;  Notes on Calochortus gunnisonii;  

Watson, 1871, Botany, United States Geological Explorations on the Fortieth Parallel

Sereno Watson was the botanist and wrote the Botany volume, with assistance from D. C. Eaton and others.

Abbreviation Full Title Availability Online
  • Botany [Fortieth Parallel]
United States Geological Expolration [sic] of the Fortieth Parallel. Botany. Washington, DC URL: https://www.biodiversitylibrary.org/item/104490

 

Other articles:
• Golden Checklist Flora:  Porter & Coulter, 1874;  

Porter & Coulter, 1874, Synopsis of the flora of Colorado

John M. Coulter was expedition botanist on Hayden's 1873 Expedition to Colorado.

Abbreviation Full Title Availability Online
  • …abbreviation…
Synopsis of the flora of Colorado https://www.biodiversitylibrary.org/bibliography/46971
 

The work is based chiefly on collections made, in 1861 and succeeding years, by Dr. C. C. Parry, whose indefatigable labors have added so much to our knowledge of the flora of the region; in 1862 by Messrs. Hall and Harbour; in 1867, by Dr. W. A. Bell, of Manitou Springs; in 1868, by Dr. F. V. Hayden; in 1869, by B. H. Smith, Esq., of Denver; in 1871, by Dr. George Smith and W. M. Canby, Esq.; in 1871 and 1873, by Messrs. Meehan and Hooper; in 1872, by J. H. Redfield, Esq.; in 1872 and 1873, by T. S. Brandegee, Esq., of Canon City, Rev. E. L. Greene, of Pueblo, and T. C. Porter; and in 1873, by J. M. Coulter.

 

Literature Cited:
- Wheeler, George M., 1878.  

Wheeler, 1878, Report upon United States Geographical surveys west of the one hundredth meridian

The Wheeler Survey was a survey of a portion of the United States lying west of the 100th meridian. It comprised multiple expeditions, and was supervised by First Lieutenant (later Captain) George Montague Wheeler.

Abbreviation Full Title Availability Online

Wheeler's Rep.
Wheeler report
Wheeler survey
Report upon United States Geographical surveys west of the one hundredth meridian

Title Variants:

Explorations and Surveys West of the One Hundredth Meridian
Explorations and surveys.
Geographical and Geological Explorations and Surveys West of the One Hundredth Meridian
Geographical Surveys West of 100th Meridian
Geographical Surveys West of the One Hundredth Meridian
Report upon explorations and surveys west of the one hundredth meridian.
Report upon geographical and geological explorations and surveys west of the one hundredth meridian.
Report upon United States Geographical and Geological Explorations and Surveys West of the One Hundredth Meridian
United States Geographical and Geological Explorations and Surveys West of the One Hundredth Meridian
United States Geographical Surveys West of 100th Meridian
United States Geographical Surveys West of the One Hundredth Meridian

URL: ttps://www.biodiversitylibrary.org/bibliography/49058
URL of Vol. 6: https://www.biodiversitylibrary.org/item/115739

 

Other articles:
• Golden Checklist Flora:  Notes on Pinus ponderosa, Watson, 1880;  

Brewer, et al., 1889 , Botany, California.

Botany report form the Geological Survey of California.

Abbreviation Full Title Availability Online

Bot. California [W. H. Brewer]
Geological Survey of California. J. D. Whitney, State Geologist. Botany. Cambridge, MA URL: https://www.biodiversitylibrary.org/bibliography/22424

 

Other articles:
• Golden Checklist Flora:  Coulter, 1885;  

Coulter, 1885, Botany of the Rocky Mountain Region

Coulter (1885) ...

Abbreviation Full Title Availability Online
  • …abbreviation…
Manual of the botany (Phaenogamia and Pteridophyta) of the Rocky mountain region, from New Mexico to the British boundary https://www.biodiversitylibrary.org/bibliography/29024

 

Literature Cited:
- Sargent, Charles Sprague., 1897.

Other articles:
• Golden Checklist Flora:  Notes on Juniperus scopulorum, Sargent 1897;  

Sargent, 1888-1897, Garden & Forest

Garden & Forest was edited by Charles S. Sargent, who was the Director of the Arnold Arboretum and Professor of Agriculture in Harvard College, etc.

Abbreviation Full Title Availability Online

Gard. & Forest
Garden and Forest; a Journal of Horticulture, Landscape Art and Forestry
Vols. 1-10, 1888-1897
URL: https://www.biodiversitylibrary.org/bibliography/50609#/summary

Names published in this publication:

  • Juniperus scopulorum Sarg. “Rocky Mountain Juniper”

 

Literature Cited:
- Heller, A. A., 1898.

Other articles:
• Golden Checklist Flora:  Holodiscus dumosus, Heller, 1898;  

Heller, 1898, Catalogue of North American Plants North of Mexico

Amos Arthur Heller (March 21, 1867 – May 19, 1944) was an American botanist. Heller was born in Danville, Pennsylvania. In 1892, Heller received a Bachelor of Arts degree from Franklin & Marshall College. In 1897, he received a Master's degree in Botany from Franklin & Marshall College. From 1896 to 1898, Heller was a professor of Botany at the University of Minnesota. From 1898 to 1899, Heller worked on the Vanderbilt Expedition to Puerto Rico under the auspices of the New York Botanical Garden. Starting in 1905, Heller was a professor of Botany at the California Academy of Sciences in San Francisco, California. After moving to California, Heller and his wife, Emily Gertrude Heller, founded the botanical journal Muhlenbergia and Heller continued to edit that journal until 1915.

Abbreviation Full Title Availability Online

Cat. N. Amer. Pl.
Catalogue of North American Plants North of Mexico, Exclusive of the Lower Cryptograms URL: https://www.biodiversitylibrary.org/bibliography/38325

 

Literature Cited:
- Rydberg, Per Axel, 1906.

Other articles:
• Golden Checklist Flora:  Rydberg, 1906;
• Field Notes:  Erigeron eximius, Greene, 1898;  

Rydberg, 1906, Flora of Colorado

Rydberg (1906) …

Abbreviation Full Title Availability Online
  • …abbreviation…
Flora of Colorado https://www.biodiversitylibrary.org/bibliography/2486
 

Some systematic collection was done by Professor James Cassidy, Professor of Botany (1881-1889), in the intervals of his many duties, up to the time of his death in 1889. His successor, Professor C. S. Crandall, gave much time to the work, especially after the establishment of the experimental grass station in cooperation with the U. S. Department of Agriculture. This afforded occasion and opportunity for trips in search of promising species of native grasses. Subsequent collections, especially of forest products and grasses for the World's Fair in 1893, gave rise to further collecting trips to various parts of the State. Enthusiastic aid was given by Mr. J. H. Cowan, an energetic and promising student, who became Professor of Horticulture and Botany upon the resignation of Professor Crandall, but whose promising career was terminated by death before he had entered upon the active duties of the position.
 

The author has also consulted the various publications on the flora of Colorado. The most important of these are :

T. C. Porter and J. M. Coulter, Synopsis of the Flora of Colorado; J.M. Coulter, Manual of the Botany of the Rocky Mountain Region; T. S. Brandegee, Flora of South-western Colorado; Alice Eastwood, Flora of Denver and Vicinity; John Torrey's report on E. James' collection in Long's Expedition; Asa Gray's reports on the collections of C. C. Parry, E. Hall and Harbour ; Professor E. L. Greene's various publications in Pittonia, Plantae Bakerianae and Leaflets and the publications of Professors T. S. Brandegee, Aven Nelson and M. E. Jones, Mr. G. E. Osterhout and Miss Alice Eastwood in the Botanical Gazette, Bulletin of the Torrey Botanical Club, Zoe, Erythea and the Proceedings of the California Academy of Sciences.

 

Literature Cited:
- Rydberg, Per Axel, 1917.

Other articles:
• Golden Checklist Flora:  Rydberg, 1917;  

Rydberg, 1917. Flora of the Rocky Mountains and adjacent plains

Rydberg (1917) ...

Abbreviation Full Title Availability Online
  • …abbreviation…
Flora of the Rocky Mountains and adjacent plains : Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and neighboring parts of Nebraska, South Dakota, North Dakota, and British Columbia https://www.biodiversitylibrary.org/bibliography/7757
Accepts both Erigeron superbus Greene and E. eximius Greene, places them in different sections, and distinguishes between Sections IX. Macranthi and X. Glabelli on the basis of reduced upper stem leaves and leaves 3-nerved or not. Title Flora of the Rocky Mountains and adjacent plains, Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and neighboring parts of Nebraska, South Dakota, and British Columbia Title Variants Abbreviated: Fl. Rocky Mts https://www.biodiversitylibrary.org/bibliography/147

Literature Cited:
- Rydberg, Per Axel, 1922.

Other articles:
• Golden Checklist Flora:  Rydberg, 1922;  

Rydberg (1922) ...

Abbreviation Full Title Availability Online
  • …abbreviation…
Flora of the Rocky Mountains and adjacent plains, Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and the neighboring parts of Nebraska, South Dakota, and British Columbia https://www.biodiversitylibrary.org/bibliography/161373

Rydberg's (1922) 2nd edition made no changes to our Colorado Chrysothamnus or Ericameria.

 

Literature Cited:
- Turland, N. J., et al., 2018.

Other articles:
• Golden Checklist Flora:  Notes on Suckleya suckleyana;  

Turland, et al., 2018, ICBN

International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017.

Abbreviation Full Title Availability Online
  • …abbreviation…
… full name of publication … URL: … if available …

Cite this Code as: Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. & Smith, G. F. (eds.) 2018: International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Glashütten: Koeltz Botanical Books. DOI https://doi.org/10.12705/Code.2018

Previous versions of the Code.

Year of publicationInformal name
1867Laws of botanical nomenclature
1883Laws of botanical nomenclature, ed. 2
1906Vienna Rules
1912Brussels Rules
1935Cambridge Rules
1950Amsterdam Code
1952Stockholm Code
1956Paris Code
1961Montreal Code
1966Edinburgh Code
1972Seattle Code
1978Leningrad Code
1983Sydney Code
1988Berlin Code
1994Tokyo Code
2000St Louis Code
2006Vienna Code
2012Melbourne Code
2018Shenzhen Code (current)

 

   

Types from the Golden Area

 

   

Namesakes of the Golden Area

 

 

   

General Information, and Mysteries Regarding Other Non-Types or Non-Namesakes

 

 

Other articles:
• Golden Checklist Flora:  Erythranthe guttata;  

Erythranthe guttata (Fisch. ex DC.) G.L. Nesom “Seep Monkeyflower”

(Syn: Mimulus guttatus DC.)

   

Equisetum L. “Horse Tail”

 

 

Other articles:
• Golden Checklist Flora:  Equisetum hyemale ssp. affine;  

Notes on Equisetum hyemale L. ssp. affine “Tall Scouring Rush”

 

Literature Cited:
- Calder, James A., and Roy L. Taylor, 1965.  

The publication of ssp. affine by Calder & Taylor (1965) was just a nomenclatural change without comment.

Equisetum hyemale ssp. affine (Engelm.) Calder & Taylor, comb. nov.
E. robustum var. affine Engelm., Am. Jour. Sci. 46: 88. 1844.
E. hyemale var. affine A. A. Eaton, Fern. Bull. 11: 111. 1903.

 

Other articles:
• Golden Checklist Flora:  Equisetum laevigatum;  

Equisetum laevigatum A. Braun “Smooth Horsetail”

 

Literature Cited:
- Braun, Alexander, and George Engelmann, 1844.  

George Engelmann translated Alexander Braun's (1844) monograph of North American Species of Equisetum, making some additions.

 

Literature Cited:
- Windham, Michael D., 1987.  

Argyrochosma, Windham, 1987

Original Text
Argyrochosma (J. Smith) Windham, stat. nov. — Notholaena sect. Argyrochosma J. Smith, J. Bot. (Hooker) 4:50. 1841. — Lectotype (chosen by Christensen, 1906, Ind. Fil., p. XL); Pteris nivea Poir. [Argyrochosma nivea (Poir.) Windham].
... [Diagnosis omitted.] ...
Distribution. — A strictly American genus of approximately 20 species occupying rupestral or (rarely) terrestrial habitats from near sea level to an elevation of 4200 m in the Andes. Ranging from Missouri, Wyoming, and California to Chile (including the Juan Fernandez Islands), Argentina and the highlands of southeastern Brazil. There is a large geographic gap between the North and South American elements of the genus (only A. incana is found in Central America and the West Indies), with the greatest diversity of species occurring in the highlands of central and northern Mexico.

 

Other articles:
• Golden Checklist Flora:  Argyrochosma fendleri;  

Argyrochosma fendleri (Kunze) Windham “Fendler's False Cloak Fern”

 

Literature Cited:
- Kunze, Gustav, 1851.  

First collected by A. Fendler 1847 in New Mexico, his No. 1017, described by Kunze from a specimen apparently deposited in the General Herbarium of Berlin (https://hdl.handle.net/2027/uc1.c026306023?urlappend=%3Bseq=181). The quality of the scan presents difficulty for one unfamiliar with German to transcribe

Literature Cited:
- Windham, Michael D., 1987.  

Original Text
4) Argyrochosma fendleri (Kunze) Windham, comb. nov. — Notholaena fendleri Kunze, Farnkr. 2:87, t. 136. 1851. — Pellaea fendleri (Kunze) Prantl

 

Literature Cited:
- Fee, A. L. A., 1852.

Other articles:
• Golden Checklist Flora:  Cheilanthes feei;  

Cheilanthes feei T. Moore “Slender Lipfern”

First described as Myriopteris gracilis by Fee (1852) citing habitat on rocks around Hillsboro, in North America.

 

Other articles:
• Golden Checklist Flora:  Selaginella densa;  

Notes on Selaginella densa Rydb. “Rocky Mountain Spikemoss”

 

 

Other articles:
• Golden Checklist Flora:  Selaginella underwoodii;  

Selaginella underwoodii Hieron “Underwood's Spikemoss”

Described from specimens collected by Fendler, in 1847, in the mountains near Santa Fe.

 

Other articles:
• Golden Checklist Flora:  Cystopteris fragilis;  

Cystopteris fragilis (L.) Bernh. “Brittle Bladderfern”

The fern was first described by Linnaeus (1753) as Polypodium fragile from habitats described as the cooler hills of Europe. Cystopteris was proposed by Bernhardi in 1805, who placed C. fragilis therein. This was done in German which, I confess, I have not tried to translate.

 

Other articles:
• Golden Checklist Flora:  Woodsia oregana ssp. cathcartiana;  

Woodsia oregana D.C. Eaton ssp. cathcartiana (B.L. Rob.) Windham “Rocky Mountain Woodsia”

 

Literature Cited:
- Robinson, B. L., 1908.  

Robinson (1908, v. 10, n. 110, p. 30) published ...

Original Text
Woodsia Cathcartiana, n. sp. W. scopulinae affnis et simillima, sed minute glanduloso-puberula nee hispidula; frondibus 2-3 dm. altis obscure viridibus firmiusculis lanceolatis 25-55 mm. latis bipinnatifidis; pinnis oblongis, inferioribus distantibus, lobis sinubus modice latis separatis oblongis denticulatis; soris submarginalibus ; indusio obscuro eo W. scopulinae simile. — W. scopulina D. C. Eaton apud Gray, Man. ed. 6, 691 (1890), non D. C. Eaton, Can. Nat. ii. 90 (1865). — Nearly related and very similar to W. scopulina, but minutely glandular-puberulent, not hispidulous; fronds 2-3 dm. high, dull green, rather firm in texture, lanceolate, 25-55 mm. wide, bipinnatifid; pinnae oblong, the lower distant; lobes oblong, denticulate, separated by rather wide sinuses; sori snbmarginal; indusium obscure, similar to that of W. scopulina. — Taylor's Falls of the St. Croix River, Minnesota, 1874, Miss Ellen Cathcart (type, in hb. Gray); also on rocks, Lower Falls of the Menomine River, Michigan, 31 August, 1892, C. F. Wheeler. This species was sent to Dr. Gray soon after its original collection. It was referred by him to Prof. Eaton, who evidently was somewhat puzzled by it. He reported it as belonging to his W. scopulina remarking, however, upon its peculiar glandular puberulence. Much additional material of the real W. scopulina, a species frequent from the Rocky Mountains westward, is now at hand and the constancy of the distinctions pointed out above is such as to warrant the publication of the plant of Minnesota and Michigan as a separate species.

 

Other articles:
• Golden Checklist Flora:  Azolla mexicana;  

Azolla mexicana C. Presl “Mexican Mosquito Fern”

 

Literature Cited:
- Presl, Karl B., 1845.  

Carl Presl (1845, ser. 5, part 3, p. 580) barely noted the presenence of a new species of Azolla. Some sources give the page number as 150. Others note that the publication on p. 580 is as isonym. I did note see anything on page 150 that looked like publication of a name in Azolla

Original Text English Translation
— Nova Azollae species est: Azolla mexicana; fronde pinnata, foliolis imbricatis laevibus subrotundis coloratis, radicibus capillaribus. Habitat in Mexico, ubi legit clar. Schiede. Affinis videtur A. portoricensi, differt foliolis margine non hyalinis. A new species of Azolla is: Azolla mexicana; fronds pinnate, subrotund overlapping leaflets smooth colored capillary roots. Lives in Mexico, where it is described by Schiede. It seems related to A. portoricensis; the edge of the leaves are not hyaline.

   

Notes on Juniperus L. “Juniper”

We have five species of Juniperus in Colorado, although our authors do not agree on which ones we have, or the nomenclature to be applied.

We definitely have something in section Juniperus, that is the circumpolar Juniperus communis but whether we apply the circumpolar ssp. alpina (Weber & Wittmann, 2012) or the North American variety depressa (Ackerfield, 2022) is apparently still up for debate.

In section Sabina one clade with smooth leaf margins includes our common Rocky Mountain Juniper, J. scopulorum, and the occasionally found garden escapee, J. virginiana. Another clade with sawtooth margins includes both J. monosperma and J. osteosperma, the former occurring in Platte Canyon in Jefferson County.

Literature Cited:
- Mao, Kangshan, Gang Hao, Lianquan Liu, Robert P. Adams, and Richard I. Milne., 2010.  

Mao, et al. (2010) placed both J. scopulorum and J. virginiana in Clade III of section Sabina, though not immediately adjacent to each other. These are the smooth-leaved American species plus the Eurasian J. sabina and a couple of others. J. monosperma and J. osteosperma are Clade II, though at some distance from each other. This clade is described as the serrate-leaved junipers of North America. Section Sabina probably originated in Asia. The presence of an American var. depressa within J. communis represents a recent diversification event.

Literature Cited:
- Adams, Robert P., and Andrea E. Schwarzbach, 2013.  

Adams & Schwarzbach (2013) suggest that migration of J. sabina and others across the Bering Land Bridge (17.6-5.5.mya) gave rise to the current, cold climate, western hemisphere species such as J. scopulorum, whereas the serrate, semi-arid junipers, such as J. osteosperma migrated from the eastern to the western hemisphere via the North American Land Bridge ca. 47-30.3 mya.

 

Other articles:
• Golden Checklist Flora:  Juniperus comminis depressa;  

Juniperus communis var. depressa “Common Juniper”

 

Other articles:
• Golden Checklist Flora:  Pursh, 1814, publication details;  

Pursh (1814, vol. II, p. 646) described our common juniper from plants he had seen in New York and Maine.

710. JUNIPERUS. Gen. pl. 1552.
communis 1. J. foliis ternis patentibus mucronatis bacca longioribus Willd. sp. pl. 4. p. 853.
erecta. α. J. ramis erectis.
depressa. β J. ramis depressis.
  α. About rocks, near the falls of rivers, in Canada and the western part of New York. β. in New York, and particularly in the province of Maine, in rocky or gravelly situations. ♄ May. v. v. The Common Juniper may probably have been originally brought from Europe ; but the variety β., or probably a distinct species, seems to be really an original native ; it does not grow above a foot or two high, and one single root will cover sometimes a space of from fifteen to twenty feet in diameter.

 

Other articles:
• Golden Checklist Flora:  Juniperus scopulorum;  

Juniperus scopulorum “Rocky Mountain Juniper”

The Rocky Mountain Juniper — Juniperus scopulorum Sargent — is found in all the hilly areas around Golden s.l. It was originally treated as J. virginiana L. or the Red Cedar. C. S. Sargent (1897) recognized J. scopulorum as a separate species. The Rocky Mountain Juniper is found as far east of South Dakota and Nebraska, where it is known to hybridize with its eastern relative J. virginiana in zones of contact in the Missouri River basin. To the west, J. scopulorum is known to occur in Oregon, Nevada, and Arizona, though not California. Hybrids of J. scopulorum and J. osteosperma are known from from Walnut Canyon [Arizona?] north into Utah and east to Mesa Verde.

Literature Cited:
- Sargent, Charles Sprague., 1897.

Other articles:
• Golden Checklist Flora:  Sargent, 1888-1897, Garden & Forest, publication details.;  

Original Text
Juniperus Virginiana has usually been considered to cross the continent to the shores of Puget Sound and Vancouver Island, and to be pretty widely distributed through the interior Rocky Mountain region from the northern border of the United States to northern New Mexico and Arizona.
After having seen, however, a good deal of this western tree during the past two seasons, I am inclined to believe that the so-called western Red Cedar as in grows in Wyoming, Montana and Colorado, at least, and perhaps everywhere, will have to be considered another species, and should this supposition prove correct on further investigation, I should propose the name of Juniperus scopulorum for it.
The habit of the Rocky Mountain tree, as may be seen in our illustration near the Mammoth Hot Springs in the Ywllowstone National Park, where this Juniper is very common, and the only arborescent, and where it grows on gravelly slopes at elevations of six or seven thousand feet with Pinus flexilis.
It has the slender branchlets and opposite leaves in pairs of the eastern tree, but the fruit is larger, and does not ripen until the second year, while that of our Red Cedar ripens during its first autumn.
The branches are stouter and covered with more scaly bark, and the bark of the trunk, which is often forked near the ground is unlike that of the eastern tree, which separates into thin narrow scales fringed on the margins, but, like that of some other western Junipers, divides into irregular, narrow, connected flat ridges, which break up on the surface more or less freely into persistent shreddy scales.
The wood has the same fragance as that of the eastern tree, although it is rather less powerful, and the color is a duller red.
The habit and the character of the bark may be due, perhaps, to differences of soil and climate, which might also affect the color of the wood, and the only really tangible character by which the western tree can be separated from the eastern is the biennial fruit.
The fact, moreover, is significant that unless the eastern and western trees come together in north-western Nebraska, the meeting place of the eastern and western floras, they are separated by a continuous belt of country through the middle of the continent several hundred miles wide; and moreover, with the exception of Juniperus communis, which encircles the northern hemisphere, and the White Spruce, which crosses this continent far northward and reaches the Pacific coast within the Arctic Circle, no coniferous tree grows in both eastern and western North America.
But before the question of the distribution of the Red Cedar can be satisfactorily determined more observations should be made on the time of ripening of the fruit, expecially in Colorado, Utah, New Mexico and Arizona, in the valley of the Columbia River and on Vancouver Island, for it is, of course, possible, although hardly probable, that the proposed Juniperus scopulorum may be confined to the northern Rocky Mountains and that Juniperus Virginiana really reaches the southern part of that range and even the Pacific coast.

According to Sargent (1897) the only character to distiguish J. scopulorum from J. virginiana is the biennial development of the fruit. FNANM, key by Robert P. Adams, distinguish the two by the extent of which the scalelike leaves overlap, by not more than 1/5 for J. scopulorum, and by the shapes of exfoliating bark, in addition to development of the fruit. J. scopulorum is distinguished from J. osteosperma by the leaf margin entire for the former, whereas leaf margin denticulate for the latter. Ackerfield's (2015) Flora of Colorado follows the same scheme, but does not accept J. virginiana occurring in Colorado, though the author has found it as a garden escapee.

 

Other articles:
• Golden Checklist Flora:  Juniperus virginiana;  

Notes on Juniperus virginiana L. “Eastern Redcedar”

 

 

Other articles:
• Golden Checklist Flora:  Pinus ponderosa;  

Pinus ponderosa Douglas ex Lawson & C. Lawson. “Ponderosa Pine”

 

Literature Cited:
- Lawson, Peter & Son, 1836.

Other articles:
• Golden Checklist Flora:  Lawson, 1836, publication details;  

Lawson & Lawson (1836) described the ponderosa pine from plants they were growing in pots at their Agricultural Museum near Edinburgh, Scotland. Material to grow the plants (seeds?) was obtained through David Douglas from his second and most successful trip to the American Pacific Northwest.

Original Text
XXV. PINUS PONDEROSA — Heavy Wooded Pine.
From not possessing the advantage of deriving information from any previous description of this species, and having only young trees in the Museum to refer to, an oppostunity has not been afforded of giving any details regarding its flowers, cones, &c. It is hoped, however, the following will be sufficient to enable those less acquainted with the general appearance of this seemingly valuable and highly interesting tree to distinguish it from others of the three-leaved pines. In its habit of growth P/. ponderosa seems to surpass all others of the genus for strength and luxuriance, the branches are few, regularly verticillated, horisontal, and seem incluned to assume a pendulous or drooping habit as the three becomes older ; central or top-shoot often more than an inch in diameter, and or proportional length ; buds large and free from resin ; leaves thickly set, nine inches to a foot or fourteen inches in length, thick rigid, and nearly straight, rounded on the exterior, and having a longitudinal prominent rib, together with minute channels on the interior side ; smooth, with very indistinctly serrated margins ; sheaths short, of a dull blackish colour, and lacerated or torn at their extremities ; timber said to be so ponderous as almost to sink in water. Introduced by Mr Douglas from the west coast of North American in 1828.
Such of the above remarks as cannot be supposed to apply to small plants in pots, are derived from a specimen, perhaps the finest in Scotland, growing in the Caledonian Horticultural Society's Gardens, Inverleith Row. Judging from its rapid growth and hardiness, this seems to be one of the most valuable species which has yet been introduced, particularly should the wood, when grown in this country, be found to possess the valuable qualities which have been ascribed to it.

Literature Cited:
- Brewer, W. H., Sereno Watson, and Asa Gray, 1880.

Other articles:
• Golden Checklist Flora:  Brewer, 1880, publication details;  

Our Rocky Mountain variety scopulorum was proposed by George Engelmann, writing in his section about Tribe III, the Abietineae in Volume II, Watson's completion of the Botany report of the California Geological Survey. Engelmann included variety scopulorum that is not known from California and other comments about conifers in the western United States in what was perhaps intended to provide regional treatment to the family.

Original Text Comments
Order CIII. CONIFERAE  
 
Tribe III. ABIETINEAE (By Dr. George Engelmann.) Engelmann's tribe Abietineae contained Abies, Pseudotsuga, Tsuga, Picea, and Pinus, and thus contains taxa in our current family Pinaceae.
 
11. PINUS, Tourn. ; Link. Pine.  
 
8. P. ponderosa, Dougl. One of the largest pines known (200 to 300 feet high and 12 to 15 feet in diameter), with very think red=brown bark, deeply furrowed and split in large plates : leaves on stout branchlets in the axils of strongly fringed somewhat presistent bracts, 5 to 9 or even 11 inches long ; the thin sheaths at first 9 or 10 (later 3) lines long : male flowers cylindric, flexuous, 1 1/2 to 2 inches long, crowded into a short head ; involucre of 10 or 12 bracts ; anthers with a large semicircular scarcely dentate crest : cones oval, 3 or 4 (rarely 5) inches long, 1 1/2 to 2 inches thick, of a rich brown color, sessile or subsessile, spreading or slightly recurved, often 3 to 5 together ; umbo high, with a stout straight or incurved prickle : seeds dark brown, 4 lines long ; wing 10 to 12 lines long, widest above the middle : cotyledons 6 to 9. — Loud. Arbor. iv. 2243 ; Newberry, 1. c. 36, t. 4 ; Parlat. 1. c. 395 ; Engelm. Wheeler's Rep. vi. 261. P. Benthamiana, Hartw. Journ. Hort. Soc. ii. 189. P. Beardsleyi and Craigana, Murr. Edinb. New Phil. Journ. i. 286.  
Var. Jeffreyi. A tree 100 to 200 feet high, with a more rounded top, more finely cleft and darker bark, and paler leaves 4 to 9 inches long : male flowers 1 1/4 inches long : cones larger, 5 to 12 inches long, lighter brown, on short peduncles, fewer in a cluster, with thinner apophyses, and slender prickles hooked backward : seeds 4 to 7 lines long ; wings 12 or 13 lines long : cotyledons 7 to 11. — P. Jeffreyl, Murr. l. c. xi. 224, t. 8, 9 ; Parlat. l. c. 393. I have included the Jeffrey pine here because that is the dominant pine on the east side of the Sierra Nevada, for example, at Sagehen Meadow.
Var. scopulorum. A smaller tree (80 to 100 feet high) : leaves 3 to 6 inches long, often in pairs : male flowers an inch long : cones smaller, 2 or 3 (rarely 4) inches long, grayish brown, with stout prickles : seeds 2 1/2 to 3 1/2 lines long, the wings 9 to 12 lines : cotyledons 6 to 9. — P. ponderosa of the Rocky Mountain floras.  

 

Other articles:
• Golden Checklist Flora:  Pseudotsuga menziesii glauca;  

Pseudotsuga menziesii (Mirb.) Franco var. glauca (Beissn.) Franco. “Douglas Fir”

 

Literature Cited:
- Lambert, Aylmer Bourke, Esq., F.R.S, F.S.A, 1802.  

The first publication of “Douglas fir” was as the “Nootka fir” from a specimen that Lambert saw in Banks' herbarium.

Banks was a member of the landed gentry who who inherited extensive family estates. Banks was educated at Eton College and at Oxford University where he developed a keen interest in natural history. He made his name as a naturalist on voyages first to Newfoundland and Labrador in Canada in 1766 and subsequently on Captain Cook’s Endeavour voyage during 1768-1771. He was a friend and advisor to King George III and pivotal in the early development of the Royal Botanic Gardens, Kew.

Lambert is best known for his work A description of the genus Pinus, issued in several parts 1803–1824, a sumptuously illustrated folio volume detailing all of the conifers then known. A second folio edition was produced between 1828 and 1837, and a third, smaller (octavo) edition in 1832. Individual books even of the same edition are often very different from one another, which causes problems when the illustrations have been used as types to fix the application of names.

Lambert was also the host of Frederick Pursh while the latter worked on his Flora of North America.

Original Text Comments
TAB. 33.  
27. PINUS TAXIFOLIA  
NOOTKA FIR  
Pinus taxifolia, foliis solitariis planis integerrimis, strobilis oblongis, antheris inflato-didymis.

Habitat ad Americae borealis oras occidentales.

Pinus taxifolia, leaves in a single plane, entire, cone oblong, anthers inflated in two lobes.

Lives on the western coast of North America.


 
DESCRIPTIO  
Habitus P. canadensis, at folia angustiosa et paululum longiora, integerrima. Amenta mascula ovata, subsessilia, multiflora ; antheris inflato-didymis, cristâ reflexâ, minimâ. P. canadensis is a synonym of Tsuga canadensis (L.) Carrière

 
The figure was taken from a specimen in the Banksian herbarium, brought home by Mr. Menzies, by whom it was discovered on the North-west coast of America, and who has favoured me with the following particulars respecting this species.

In general habit this tree resembles P. canadensis, and attains considerable height and size. The leaves are also very like those of the species just mentioned, but narrower, and their edges are entire, whereas the others are visibly serrated. The inflorescentia is much larger than in P. canadensis and there are more antherae. As for the Cones, I can give no account of them, those which were brought by Mr. Menzies having been unfortunately mislaid. That gentleman however, informs me that they differ in their form from the cones of P. canadensis, and that they are longer.

 
Explanation of the figure is omitted.  

Literature Cited:
- Mayr, Heinrich, 1890.  

The epithet “glauca” was first proposed by Mayr (1890).

Original Text Translation and Comments.
Pseudotsuga Douglasii var. glauca, Colorado Douglasia ist ausgezeichnet durch eine auffallende, hellweissliche Farbung der Nadeln; doch zeigen nur die einjahrigen Nadel, diese schone Bereiftheit; an den zwei- und mehrjahrigen Trieben verschwindet wiederum die weissliche Farbe, weshalb eigentlich nur jungen Exemplaren besondere Schonheit verliehen ist; auch andere solche glauca- Varietaten und -Arten, wie Pinus pungens, zeigen dieses Verhalten. Colorado Douglasia is characterized by a striking, bright white color of the needles; but only the one-year-old needles show this beautiful frostiness; the whitish color disappears on the biennial and perennial shoots, which is why only young specimens are particularly beautiful; other such glauca varieties and species, such as Pinus pungens, also show this behavior.
Die glauca - Douglasia ist in Colorado, New-Mexico und Arizona (Santa Rita) heimisch. Auch der Zapfen zeigt einige Verschiedenheiten ; er is namlich kleiner und armer an Schuppen als die westliche Form (Tafel VI); die kleinfruchtige Douglasia van Montana bildet hierin die Verbindung zwischen der glauca und der typischen Douglasia. In trockenem un im Winter kalterem Klima erwachsen hat sich diese Varittat im Osten der Union als frosthart, das heisst wohl in den allermeisten Fallen als trockenhart, wenn man so sagen kann, erwiesen. Sie is in ihrer Heimat wie uberall, wo sie kultivert wird (auch im deutschen Walde) langsanwuchsig (verglichen mit der typischen Form), eine unangenehme Eigenschaft, die ihre sonstigen Vorzuge wieder aufhebt; auch als Nutzholz soll sie weit hinter der Kustenform zuruckstehen. The glauca - Douglasia is native to Colorado, New Mexico and Arizona (Santa Rita). The cone also shows some differences; it is namely smaller and poorer in scales than the western form (Plate VI); the small-fruited Douglasia from Montana forms the link between the glauca and the typical Douglasia. Grown up in dry climates and colder climates in winter, this variety has proven to be frost-hardy in the east of the Union, that is to say, in most cases, dry-hardy, if you can say that. In her homeland, as everywhere where she is cultivated (also in the German forest), she is long-growing (compared to the typical shape), an unpleasant quality that cancels out her other advantages; Even as timber, it should lag far behind the coastal shape.
 

Pseudotsuga taxifolia (Poir.) Britton ex Sudw., in U. S. Dept. Agric., Div. For., Bull. No. 14, 46 (1897); et in U. S. Dept. Agric., Div. For., Bull. No. 14, 46 (1897) Bull. No. 17, 24 (1898), in adnot.; Rehder apud Sprague & M. L. Green in Kew Bull. 1938, 80.

I have been unable to find a copy of Division of Forestry, No. 14 in any online form.

Literature Cited:
- Editors of Wikipedia, 2021.
- Franco, João do Amaral, 1950.
- Heywood, Vernon, 2009.  

Joao do Amaral Franco (1921-2009) was a professor of botany at the Instituto Superior de Agronomia from 1950 until his retirement in 1991. At the beginning of the 1950s, he started his duties as a teacher, maintaining constant research on conifers, which resulted in the publication of several notes and the proposal of several taxonomic arrangements in that group. Over time his research interests broadened, establishing contact with European researchers, with emphasis on those based at the Royal Botanical Garden in Kew and at the Natural History Museum in London, institutions where he worked.

Original Text Translation and Comments
CEDRUS LIBANENSIS
ET PSEUDOTSUGA MENZIESII
Cedrus libanensis
and Pseudotsuga menziesii
a by
JOÃO DO AMARAL FRANCO
( Olisiponis Instituti Superioris Agronoraiae Assistente )
João do Amaral Franco
(Higher Institute of Agronomy at the Technical University of Lisbon)
Recebido em 14 de Janeiro de 1950. Received 14 January 1950.
Discussion of Cedrus libanensis Mirb. omitted.  
Pseudotsuga Menziesii (Mirb.) Franco, nov. comb. Pseudotsuga Menziesii (Mirb.) Franco, nov. comb.

Pinus taxifolia Lamb., Descript. Gen. Pinus ed. 1, 1: 51, t. 33 (1803) ; non Salisb. (1796).
Abies taxifolia (Lamb.) Poir. in Lam., Encycl. Méth. Bot. VI : 523 (1804) ; non Du TOUR (1803).
Abies Menziesii Mirb. in Mém. Mus. Hist. Nat. (Paris) XIII: 63, 70 (1825) «Menziezii».
Abies Douglasii Hort, ex Loud., Hort. Brit. ed. 1: 388 (1830) ; nom. nud.
Pinus Douglasii Sabine ex D. Don in Lamb., Descript. Gen. Pinus ed. 3, II: 1 p., 1 t. (1832).
Abies mucronata Raf., Atl. Journ. I: 120 (Autumnus 1832).
Abies Douglasii (Lamb.) Lindl, in Penny Cycl. 1: 32 (1833).
Picea Douglasii (Lindl.) Link in Linnaea XV : 524 ( 1841) «Douglassi ».
Tsuga Douglasii (Lindl.) Carr., Tr. Conif. ed. 1: 192 (1855).
Pseudotsuga Douglasii (Lindl.) Carr., Tr. Conif. ed. 2 : 256 (1867).
Pseudotsuga taxifolia (Lamb.) Britt., in N. Y. Acad. Sei. Trans. VIII : 74 (1889) comb, illegit.
Pseudotsuga mucronata (Raf.) Sudw. ap. Holz., U. S. Dept. Agr. Div. Botany, Contrib. U. S. Natl. Herbarium 3: 266 (1895).
Pseudotsuga taxifolia ( Poir. ) Britt. ex Sudw., U. S. Dept. Agr. Div. Forestry Bull. 14: 46 (1897).
Abietia Douglasii (Lindl.) Kent in Veitch, Man. Conif. ed. 2: 476 (1900).
Pseudotsuga taxifolia (Poir.) Rehd. ap. Sprague et Green in Kew Bull. Misc. Inform. 1938 (2) : 80 (23-III-1938) comb, superfl.
 
In hodiernum haec species novissime Pseudotsuga taxifolia (Poir.) Britt. ex Sudw. vocabitur.

Pinus taxifolia Lamb. (1803) hujus speciei primum nomen est sed ante art. 61 Nomenclaturae Botanicae Regularum (1935) legitimum non est ob vetustiorem homonymum Pinum taxifoliam Salisb. (1796) Abietis balsameae (L.) Mill, synonymum. Ob earn causam Pseudotsuga taxifolia (Lamb.) Britt. (1889) legitima combinatio non est.

POIRET (1804) Abietem taxifoliam (Lamb.) Poir. hanc speciem nominavit. Ut combinatio nova Abies taxifolia (Lamb.) Poir. nomen legitimum non est sed ut nomen novum Abies taxifolia Poir. ante art. 69 licet. SPRAGUE et GREEN (1938) nominem Abietem taxifoliam Poir. in Indice Kewense non perscriptum a cl. REHDER ex occasione inventum dicent. Notandum est quod HENRY in ELWES et HENRY, Trees Great Brit. & Irel. IV : 814 (1909) ut Pseudotsugae Douglasii (Lindl.) Carr. synonymum nominem « Abies taxifolia Poiret, in Lamarck, Dict. vi. 523 (1804)» refert. SPRAGUE et GREEN ad REHDER novam combinationem imputaverunt et sie hanc speciem nominaverunt : « Pseudotsuga taxifolia (Poir.) Rehd. » Autem LITTLE in Amer. Journ. Bot. XXXI (9): 594 (1944) demonstravit hanc combinationem novam non esse quoniam SUDWORTH (1897) ad BRITTON jam imputavit.

Novissime duo nova nomina in Indice Kewense non perscripta reperimus quae hujus speciei nomenclaturam omnino mutant.

Re ipsa Du TOUR in Nouv. DiCt. Hist. Nat. XX (1803) in articulo «SAPIN» generem Abietem recognoscet et ut prima species (op. cit. 114) describet:

 
«SAPIN COMMUN, SAPIN ARGENTÉ, SAPIN BLANC, SAPIN A FEUILLES D'IF, Pinus picea Linn.; Abies alba Mill.; Abies taxifolia Mus., très-grand arbre don’t la tige est droite et nue jusqu'à son sommet, et don’t les branches sont parallèles à l'horizon; sa tête forme une pyramide. Son bois, tendre et résineux, est revêtu d'une écorce blanchâtre, sèche et friable. Ses feuilles sont étroites, assez longues, échancrées à leur extrémité et blanchâtres en dessous ; ses fleurs mâles disposées en grappes axillaires, et ses cônes rougeâtres,… Ce bel arbre habite les hautes montagnes et les pays élevés où il forme de vastes forêts ; il est très-commun en Suisse, en Allemagne, dans les environs de Strasbourg…»
“COMMON FIR, SILVER FIR, WHITE FIR, YF-LEAF FIR, Pinus picea Linn .; Abies alba Mill .; Abies taxifolia Mus., A very tall tree with the stem straight and bare to the top, and the branches parallel to the horizon; his head forms a pyramid. Its wood, soft and resinous, is covered with a whitish bark, dry and crumbly. Its leaves are narrow, fairly long, indented at their end and whitish below; its male flowers arranged in axillary clusters, and its reddish cones,… This beautiful tree inhabits high mountains and high countries where it forms vast forests; it is very common in Switzerland, in Germany, in the surroundings of Strasbourg… ”
In exemplo supra nomen Abies taxifolia vestustior quam Abies taxifolia Poir. et alia specie assignatum liquet.

In Du TOUR articulo Mus. verisimiliter Musaei abbreviatio est, in hoc casu Lutetiae Musaei Historiae Naturalis.

 
Adversum synonymiam etiam nunc cognitam expectanda erat, casu non legitimae fuissent combinationes supra Pinum taxifoliam Lamb, vel Abietem taxifoliam Poir. fundatae, inter Pinum Douglasii Sabine ex D. Don (1832) et Abietem mucronatam Raf. (1832) vetustatis electio. Autem hoc supervacuum agnovimus quia MIRBEL (1825: 63) explanat :

« Les bords du Tachoutché-Tessé, lequel coule à l'ouest des Rocheuses, sont garnis des mêmes arbres que les bords de l'Oungigah. Cette riche végétation se porte dans la Nouvelle--Hanovre jusqu'aux plages de l'Océan Pacifique, par 52° 20' de latitude, vis-à-vis l'île Nootka, où le naturaliste Menziez, compagnon de Vancouver, découvrit un Abies, que Lambert a nommé taxifolia, et que j'appellerai Menziezii pour le distinguer de l'Abies taxifolia du Jardin du Roi, qui a pour lui l'antériorité. »
 
Idem auctor (1825: 70) in enumeratione generis Abietis specierum sequentem offert :  
« 11. Menziezii. Mirb. Nootka. Nouvelle-Géorgie taxifolia. Lamb, non Desf. (lat. 51° à. …)»
 
Notandum est quod MIRBEL (1825) europaeam speciem ut Abies taxifolia Desf. refert.  
Tali modo Abies Menziezii Mirb. primum legitimum nomen est et hinc novam combinationem Pseudotsugam Menziesii (Mirb.) Franco proponimus. Haec species in honorem ARCHIBALD MENZIES qui eam anno 1797 in Nootka sinu detexit nominata est et ideo epithetum a Menziezii ad Menziesii emendamus.  
Notandum est quod multis Coniferarum operibus nomen Abies Menziesii (Dougl. ex D. Don) Lindl. (1833) supra Pinum Menziesii Dougl. ex D. Don (1832) fundatum et Piceae sitchensis (Bong.) Carr. synonymum reperitur. Tamen Abies Menziesii (Dougl. ex D. Don) Lindl, ut posterior detrimentum afferre Abiete Menziesii Mirb. non potest.  
Pseudotsugae Menziesii (Mirb.) Franco specimen typicum sub nomine Pini taxifoliae Lamb, et ab ARCHIBALD MENZIES lecto in Herbario Musei Historiae Naturalis Londinensis servatur ubi aestate priore cum cl. J. RAMSBOTTOM permissu eum vidimus.  
Diversas varietates formasque continet inter quas :  
α — Var. viridis (Schwer.) Franco, nov. comb.

Pseudotsuga Douglasii var. viridis Schwer, in Mitt. Deutsch. Dendr. Ges. XVI : 257 (1907).

 
for. densa (Slavin) Franco, nov. comb.

Pseudotsuga taxifolia for. densa Slavin in Chittenden, Conif. Cultiv. 137 (1932).

 
for. dumosa (Carr.) Franco, nov. comb.

Pseudotsuga Douglasii [var.] dumosa Carr., Tr. Conif. ed. 2: 258 (1867).

 
β — Var. caesia (Schwer.) Franco, nov. comb.

Pseudotsuga Douglasii var. caesia Schwer, in Mitt. Deutsch. Dendr. Ges. XI : 86 (1902).

 
γ — Var. glauca (Mayr) Franco, nov. comb.

Pseudotsuga Douglasii var. glauca Mayr, Wald. Nordam. 307 (1890).

P. glauca (Mayr) Mayr in Mitt. Deutsch. Dendr. Ges. XI : 86 (1902).

P. taxifolia var. glauca (Mayr) Schneid, in Silva-Tarouca, Uns. Freiland-Nadelholz. 269 (1913).

 
for. pumila (Beissn.) Franco, nov. comb.

Pseudotsuga Douglasii var. pumila Beissn., Handb. Nadelholzk. ed. 2: 110 (1909).

P. taxifolia var. pumila (Beissn.) M. L. Green in Kew Bull. Misc. Inform. 1938 (2): 85 (23-III-1938)

 

Literature Cited:
- Little, Elbert L., 1952.  

Little (1952, p. 181) rejects Pseudotsuga menziesii (Mirb.) Franco, and accepts the Rocky Mountain variety as P. taxifolia var. glauca (Beissn.) Sudw.

Literature Cited:
- Harrington, H. D., 1954.  

Harrington (1954) accepts Pseudotsuga taxifolia (Poir.) Britt. ex Sudworth, U. S. D. A. For Div. Bull. 14:46. 1897

Literature Cited:
- Harrington, H. D., 1964, 2nd ed..  

Harrington (1964, 2nd ed.) continues to accepts Pseudotsuga taxifolia (Poir.) Britt. ex Sudworth, U. S. D. A. For Div. Bull. 14:46. 1897

Literature Cited:
- Lipscomb, Barney, 1993.  

Typically, the varieties of P. menziesii are distinguished geographically. But what of those that might be planted? If one encounters an itinerant Douglas fir around Denver, is it more likely the local variety glauca, or an import from the Northwest? Flora of North America (Lipscomb, 1993) distinguishes the varieties as follows:

Original Text
1 Bracts straight, appressed; seed cones 6-10 cm; leaves yellowish green; Pacific Coast region. 2a var. menziesii
+ Bracts spreading, often reflexed; seed cones 4-7 cm; leaves bluish green to dark green or gray-green; Rocky Mountain region. 2b var. glauca

Other articles:
• Field Notes:   Coll. No. 1502, 12 Jul 2016;
Full Size ImageColl. No. 1502, Pseudotsuga menziesii var. glauca  

My collection no. 1502, 13-Jul-2016, made on the northwest slope of North Table Mountain looks to be variety glauca.

 

There are two Douglas firs planted in my neighborhood. Are they the local variety? Or, the non-native variety?

 

Other articles:
• Golden Checklist Flora:  Acer platanoides;  

Acer platanoides L. “Norway Maple”

 

 

Other articles:
• Golden Checklist Flora:  Acer saccharinum;  

Acer saccharinum L. “Silver Maple”

 

 

Other articles:
• Golden Checklist Flora:  Populus;  

Populus L. “Poplar”

 

Literature Cited:
- Stettler, R. F., H. D. Bradshaw, Jr., P. E. Heilman, and T. M. Hinckley, editors, 1996.  

 

 

Other articles:
• Golden Checklist Flora:  Populus alba;  

Populus alba L. “White Cottonwood”

 

 

Other articles:
• Golden Checklist Flora:  Populus angustifolia;  

Populus angustifolia E. James “Narrowleaf Cottonwood”

 

Literature Cited:
- James, Edwin, ed., 1823.  

James (1822, v. 1, p. 497) ... entry for July 4, 1820, while encamped on the South Platte River near Brighton.

Several valuable plants were here collected, and among others a large suffruticose species of Lupine. The long leaved cotton-wood* of Lewis and Clark, which is according to their suggestion, a species of populus, is here of very common occurrence. It is found intermixed with the common cotton-wood, resembling it in size and general aspect. Its leaves are long and narrow, its trunk smoother, and its branches more slender and flexile, than those of the Populus angulata. Some of its fruit was fortunately still remaining, affording us an opportunity to be entirely satisfied of its relation to this genus.

* Populus angustifolia, J.

 

Other articles:
• Golden Checklist Flora:  Populus deltoides ssp. monilifera;  

Populus deltoides Marshall ssp. monilifera (Aiton) Eckenw. “Plains Cottonwood”

(Syn: Populus sargentii Dode)

Literature Cited:
- Marshall, Humphry, 1785.  

Marshall (1785, p. 106) ...

1. Populus deltoide. White Poplar, or Cotton Tree of Carolina.
(Bartram's Catalogue.)
This becomes a tall tree, with a large erect trunk, covered with a white, fmoothifh bark, refembling that of the Afpen tree. The leaves are large, generally nearly triangular, toothed or indented with fharp and deep ferratures, of a fhining full green on their upper furface, but fomewhat lighter or hoary underneath; ftanding upon long flender footftalks, and generally reftlefs or in motion. The timber is white, firm, and elaftic, principally ufed for fence rails. It grows naturally upon rich low lands, on the banks of large rivers in Carolina and Florida.

Literature Cited:
- Aiton, William, 1789.  

Aiton (1789, v. 3, p. 406) ...

monilife-
ra
.
8. P. foliis fubcordatis glabris bafi glandulofis : ferraturis cartilagineis hamatis pilofiufculis ; nervis patulis, petiolis compleffis, ramis teretibus.
  Canadian Poplar Tree.
  Nat. of Canada.
  Introd. about 1772, by John Hope, M. D.
  Fl. May. H. ♄

Literature Cited:
- Eckenwalder, James E., 1977.  

Eckenwalder (1977, v. 58, n. 3, p. 93-208) ...

Populus deltoides subsp. monilifera (W. Aiton) Eckenwalder, stat. nov.
Populus monilifera W. Aiton, Hortus Kewensis 3: 406. 1789. Type: (Canada) cult. Kew Gardens, W. Aiton (holotype, BM!).
Populus deltoides var. occidentalis Rydberg, Mem. New York Bot. Gard. 1:115. 1900.
Populus sargentii Dode, Bull. Soc. Hist. Nat. Autun 18: 198. 1905.
Populus deltoides var. monilifera (W. Aiton) A. Henry, Gard. Chron. Ser. 3. 56: 2. fig. 4. 1914.
Populus texana Sargent, Bot. Gaz. (Crawfordsville) 67: 211, 212. 1919.
Most modern discussion of P. deltoides has revolved around the distinctness of the cottonwoods of the Great Plains region from their eastern counterparts (Fowells, 1965). The contrasts of foliar and bud characteristics usually advanced to distinguish a western species (P. sargentii) or variety (Populus deltoides var. occidentalis) from the eastern cottonwood (Sargent, 1913) really apply solely to the southern cottonwood, P. deltoides subsp. deltoides. The cottonwoods of the Great Lakes and other representatives of the northern cottonwood are, in fact, quite similar to the trees of the plains, and I treat them all as members of a single subspecies. They are similar in their short pedicels, pubescent winter buds, and long-acuminate, deltoid-ovate leaves (Figure 2D), generally with a single pair of basilaminar glands (although these are absent in a few Texas plants). There is a general decline in tooth number southwestward from the northeastern part of its range. This subspecies occurs from the Great Lakes region westward to the Prairie Provinces and south to the Texas panhandle.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

“ ... [like a string of beads, alluding to the racemes of fruits] ...” (Weber & Wittmann, 2012, p. 347).

 

Other articles:
• Golden Checklist Flora:  Populus tremuloides;  

Populus tremuloides Michx. “Quaking Aspen”

 

Literature Cited:
- Michaux, Andre, 1803.  

Michaux (1803, v. 2, p. 243) ...

tremuloides. P. foliis parvulis, suborbiculatis, abrupte acuteque acuminatis, serrulatis, margina pubescentibus.
Hab. in Canada et Noveboraco.

 

Other articles:
• Golden Checklist Flora:  Salix alba;  

Salix alba L. “White Willow”

 
 

Linnaeus (1753, v. 2, p. 1021) ...

29. SALIX foliis lanceolatis acuminatis ferratis utrinque pubefcentibus : ferraturis intimis glandulotis. Hort. cliff. 473. Fl. fusc. 812. Mat. med. 449. Dalib. parif. 297. Roy. lugdb. 83 alba.  
Salix foliis elliptico-lanceolatis ferratis fubtus fericeis , vimine fragili. Hall. helv. 152.  
Salix vulgaris alba arborefcens. Bauh. pin. 473.  
Salix alba. It. fcan. 200.  
Habitat ad pagos & urbes Europae. ♄ She lives in the villages and cities of Europe.
Eft arbor maxima cum Salice fragili. There is a very large tree with the Salice (catkins?) fragile.

 

Other articles:
• Golden Checklist Flora:  Salix amygdaloides;  

Salix amygdaloides Andersson “Peachleaf Willow”

 

Literature Cited:
- Andersson, Nils Johan, 1858.  

Andersson, N. J. 1858. Enumeratio Salicum nobis ex America boreali huc usque cognotarum “The Willow Flora known to us from north America” Öfversigt af Kongl. Vetenskaps-akademiens forhandlingar “Overview of the Proceedings by the Royal Academy of Sciences” v. 15, p. 114-133 https://www.biodiversitylibrary.org/item/54173

Andersson (1858, v. 15, p. 114) ...

3. S. amygdaloides n. sp. ; triandra; amentis lateralibus pedunculatis, rigidiusculis ; pedunculo foliato; capsulis ovato-conicis, glabris, pedicello nectarium 6:ies superante, stylo subnullo, stigmatibus brevissimis, partitis; foliis late lanceolatis, utrinque glaberrimis, subtus pallidioribus, margine glanduloso-serratis, exstipulatis.
Hab. »Missouri: Fort Pierre (Priuz Neuwied)» Hb. vindob.
Haec species, prae ceteris affinibus, S. amygdaliam nostram latifoliam refert. Folia tamen iis S. pentandrae etiam similia; capsulae longius pedicellatae, squamae mox caducae.

 

Other articles:
• Golden Checklist Flora:  Salix exigua;  Nuttall, 1842-1849, publication details;  

Salix exigua Nutt. “Coyote Willow”

 

Literature Cited:
- Nuttall, Thomas, 1840-1841.
- Nuttall, Thomas, 1842-1849.
- Nuttall, Thomas, 1848b.  

The species was described by Nuttall (1842, vol. 1, p. 90) in his extension of Michaux's North American Sylva as being found in the Territory of Oregon without any additional information about collector, location, date, &c. At the time Oregon extended from central Wyoming to the Pacific coast.

Original Text Comments
SLENDER WILLOW.  
Salix exigua. Foliis linearibus utrinque acutis subintegerrimus sericeis, stipulus nullis, amentis scrotinis elongatis, capsulis lanceolatis sessilibus, demum nudiusculis.  
This species is also a native of the Territory of Oregon, and grew with the preceding, which it strongly resembles: it is, however, a smaller species; the serrulations are mostly wanting, though very minute ones are sometimes seen: the capsules are smaller and not pedicellated. The male plant I have not seen. The branches are reddish brown and smooth. The preceding was River Willow Salix fluviatilis that is now treated as a synonym of Salix melanopsis Nutt.

It is a little curious that Nuttall mentions no willows in either his report of his residency in Oregon (Nuttall, 1840) or in his descriptions of plants collected by William Gambel (Nuttall, 1848).

Literature Cited:
- Argus, George W., 2010.  

The treatment of Salix in FNANM, written by Geoge Argus, treats S. exigua with one variety hindsiana, which is not known to occur in Colorado. Argus (2010) treats S. interior at the rank of species saying:

Sometimes Salix interior is treated as a subspecies of S. exigua (R. D. Dorn 1998). Salix exigua and S. interior hybridize and apparently intergrade in the western Great Plains; because the area of overlap is relatively small and distinctiveness of the two taxa is not compromised by hybridization and introgression, it is best to treat them as separate species.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012) do not accept any varieties of S. exigua, treating S. interior at the rank of species.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015) treats S. interior Rowlee as S. exigua var. interior (Rowlee) Cronquist.

 

Other articles:
• Golden Checklist Flora:  Salix irrorata;  

Salix irrorata Andersson “Dewystem Willow”

 

Literature Cited:
- Andersson, Nils Johan, 1858.  

Andersson (1858, v. 15, p. 117) ...

14. S. (daphnoides) irrorata n. sp. amentis sessilibus, perulis maximis primo bracteatis, valde condensatis, masculis brevibus, femineis horizontalibus, elongatis, densifloris; capsulis sessilibus, crasse conicis, glaberrimis, stylo producto, stigmatibus integris; foliis lanceolatis, utrinque viridibus; ramis densissime glauco-irroratis.
Hab. in Mexico nova (Coll. Fendler, n. 812).
Nostrae S. daphnoidi ita est similis, ut nullis notis nisi amentis eximic condensatis et foliis (novellis!) integerrimis utrinque viridibus ab ea distingui possit. Squamae amenti masc. pilis brevibus, aureis vestitae; squamae amenti fem. pilis sat brevibus griseis ciliatae, unde amenta non ut in vera S. daphnoide longe pilosa conspiciuntur.

 

Other articles:
• Golden Checklist Flora:  Betula occidentalis;  

Betula occidentalis Hook. “Water Birch”

 

 

Other articles:
• Golden Checklist Flora:  Carpinus betulus;  

Carpinus betulus L. “European hornbeam”

There is only one collection of Carpinus betulus L. “European hornbeam” in Colorado, that made near the Coors plant in Golden in 1949.

Linnaeus (1753) described the tree as native to Europe and Canada, though POWO does not show it lives there, nor are there any Canadian collections in SEINet. The tree is native to Germany, and it is possibly planted on the Coors grounds for that reason.

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 998) ...

Original Text
Betulus. 1. CARPINUS fquamis ftrobilorum planis. Hort. cliff. 447. Fl. fuec. 786. Roy. lugdb. 79. It. fcan. 46. Dalib. parif; 294.
  Carpinus. Dod. pempt. 841. Cam. epit. 71.
  Oftrya ulmo fimilis, fructu in umbilicis foliaceis. Bauh. pin 427.
  Habitat in Europa, Canada. ♄

 

Other articles:
• Golden Checklist Flora:  Corylus cornuta;  

Corylus cornuta Marshall “Beaked Hazelnut”

 

 

Other articles:
• Golden Checklist Flora:  Quercus gambelii;  Nuttall, 1848, publication details;
Full Size ImageColl. No. 2288, Quercus gambelii  

Quercus gambelii Nutt. “Gambel Oak”

Gambel Oak — Quercus gambelii — is found mostly in southern Jefferson County and, until recently, not in Golden. However, the writer found it recently in a canyon in Apex Park. It is also found on Dinosaur Ridge just south of Interstate 70 and therefore just south of the Golden city limits. The Gambel oaks on Eagle Ridge Drive near Kinney Run were planted.

The oak is broadly distributed throughout the American Southwest. For example, the writer has also collected it in the Spring Mountains, Clark County, Nevada, about 45 km. northwest of Las Vegas.

Literature Cited:
- Jercinovic, Gene, n.d..
- Nuttall, Thomas, 1848b.

Locations: Rio Grande.  

William Gambel (June 1823 – December 13, 1849) was an American naturalist, ornithologist, and botanist from Philadelphia. As a young man he worked closely with the renowned naturalist Thomas Nuttall. At the age of eighteen he traveled overland to California, becoming the first botanist to collect specimens in Santa Fe, New Mexico and parts of California.

In March 1841, at the age of eighteen, Gambel set off on his own for California to collect plants and other specimens for Nuttall. He planned to take a more southerly route than that taken in 1834 by Nuttall and John Kirk Townsend. Upon reaching Independence, Missouri he joined a group of traders and headed for Santa Fe following the Santa Fe Trail. Gambel reached Santa Fe in June and spent the next couple months collecting plants. In September Gambel joined a party heading to California and accompanied them along the Old Spanish Trail, arriving in Mexican Alta California in early November, 1841, becoming the first botanist to enter California overland from the east.

Nuttall (1848) described an oak collected on the Rio Grande by William Gambel.

Original Text Comments and Interpretation
QUERCUS.
Q. Gambelii. Leaves obovate, shortly petiolate, narrowed below, sinuately lobed, dilated and somewhat 3-lobed at the summit, beneath pubescent, the lobes rather obtuse, the upper ones subdentate ; fruit sessile, small, the cup hemispherical, scales ovate-acute ; the glande ovate and acute, about half immersed in the cup ; the conic summit short.
HAB. On the banks of the Rio del Norte, but not abundant. With the aspect of our northern oaks, but very distinct ; in the leaf approaching a little to L. obtusiloba, but without any near affinity. [The Rio del Norte is now called the Rio Grande. – Ed.]

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

The collections around Golden may be the northernmost occurrences along the Front Range. Gambel oak has been collected in the vicinity of Boulder, though Weber & Wittmann (2012) note that they were planted.

 

Other articles:
• Golden Checklist Flora:  Ulmus americana;  

Ulmus americana L. “American Elm”

 

 

Other articles:
• Golden Checklist Flora:  Ulmus parvifolia;  

Ulmus parvifolia Jacq. “Chinese Elm”

 

 

Other articles:
• Golden Checklist Flora:  Ulmus pumila;  

Ulmus pumila L. “Siberian Elm”

 

 

Other articles:
• Golden Checklist Flora:  Celtis occidentalis;  

Celtis occidentalis L. “Western Hackberry”

 

Literature Cited:
- Gronovius, Jan Frederik, and John Clayton, 1738-1743.
- Linne´, Carl von, 1753.

Other articles:
• Golden Checklist Flora:  Gron. virg., 1738;  

occidentalis. 3. CELTIS foliis oblique, ovatis ferratis acuminatis.
  Celtis procera, foliis ovato-lanceolatis ferratis fructu pullo. Gron. virg. 195
  Lotus arbor virginiana, fructu rubro. Raj. hift. 1917.
  Habitat in Virginia. ♃
  Folia tenera, ovato-lanceolata, parum pubefcentia; adulta latopovata, acuminata, acumine & bafi integerrima, ceteram ferrata nuda, nervofo-venofa, latere-poftico duplo minore. Maxime affinis primae.

 

Other articles:
• Golden Checklist Flora:  Celtis reticulata;  

Celtis reticulata Torr. “Net-Leaved Hackberry”

 

Literature Cited:
- Torrey, John G., 1828.  

Torrey (1828, v. 2, p. 247) in his description of plants collected by Dr. Edwin James ...

Original Text
414. Celtis reticulata, folds lato-cordatis, subcoriaceis, subintegerrimis, obtusiusculis, basi inequalibus, supra papilloso-scaberrimis, subtus venis elevatis reticularis, pubescentibus ; pedunculis 1-floris.
Hab. Base of the Rocky Mountains.
Obs. A tall shrub. Branches compressed ; younger ones villous. Leaves about an inch and half long, obtuse, or a little acute, not acuminate, of a firm and almost coriaceous texture; margin with sometimes three or four serratures ; upper surface shining, but very scabrous ; beneath reticulated with prominent veins, pubescent and slightly scabrous ; petioles about three lines long. Fruit globose, solitary, on recurved pedicels one fourth of an inch in length.
This species is very distinct from C. occidentalis β. integrifolia of Nuttall.

   

Humulus L. “Hops”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 1028) ...

Original Text Translation and Comments
HUMULUS  
Lupulus. 1. HUMULUS. Hort. Cliff. 458. Fl. fuec. 818. Mat. med. 458. Dalib. parif. 301. Roy. lugdb. 222.
♀ Lupulus mas. Bauh. pin. 298.
  Lupulus falictarius Fuchf. hift. 124.
♂ Lupulus femina. Bauh. pin. 298. Cam. epit. 954.
Habitat in Europae fepibus & ad radices montium. ♃
 

Literature Cited:
- Nuttall, Thomas, 1848a.  

Nuttall (1848, p. 23) proposed Humulus americanus in describing some collections by Gambel.

Original Text
HUMULUS.
H. * Americanus. Leaves 3 to 5-lobed, the upper sometimes entire; inner divisions lanceolate-acuminate, denticulate along the apex ; scales of the cone ovate, acute, the lower ones acuminate.
Hab. Throughout the United States in alluvial situations. I have also most luxuriant specimens from the borders of streams (Ojito de Navajo) in the Rocky Mountains, near the line of New Mexico, collected by Mr. Gambel.

Literature Cited:
- Nelson, Aven, and T. D. A. Cockerell, 1903.  

Nelson & Cockerell (1903, v. 16, p. 45) ...

Original Text
Humulus Lupulus neomexicanus, n. var.
Leaves divided or sometimes parted, the segments varying from broadly lanceolate to nearly linear, acuminate, freely sprinkled with resin particles on the lower face; fruiting bracts ovate-lanceolate, usually acuminate, finely pubescent.

The hop indigenous in New Mexico seems to possess these characters in variance with the usual and more widely distributed form and may probably best stand as a variety. The type of the variety is No. 14, T. D. A. Cockerell, Beulah, N. M. (Canadian Zone) August, 1902. It is also abundant on the Valle Ranch, Pecos, N. M., and was collected by Professor Wooton in the White Mountains of that State (No. 294).

Literature Cited:
- Rydberg, Per Axel, 1917.  

Rydberg (1917) Flora of the Rocky Mountains …

2. HUMULUS L. Hops.
Perennial, twining herbaceous vines. Leaves opposite, 3-7-lobed, serrate. Stipule persistent, free. Staminate flowers in panicled racemes; sepals 5, imbricate; stamens 5; filaments short, erect. Pistillate flowers in ament-like, drooping spikes, 2 together, subtended by a bract; ovary 1-celled. Achenes a little flattened. Embryo spirally coiled.
Leaf-blades 3-7-lobed about half-way to the base, with ovate, acute or short-acuminate lobes: those of the inflorescence 3-lobed or undivided. 1. H. Lupulus.
Leaf-blades 5-7-divided to near the base, with lanceolate, long-acuminate divisions ; those of the inflorescence 5-cleft. 2. H. neomexicanus.
1. H. Lupulus L. A vine 5-10 m. high; leaf-blades cordate in outline, dark green, scabrous above, glabrous beneath except the pubescent veins; lobes coarsely toothed, with ovate teeth; bracts of the pistillate flowers broadly ovate, from obtuse to short acuminate. Common Hops. Rocky banks and copses: N.S. — Ga. — Kans. — Wyo. — Mont.; Eurasia; extensively cultivated. Plain — Submont.
2. H. neomexicanus (A. Nels. & Cockerell) Rydb. A vine 5-10 m. high; leaf-blades light green, minutely scabrous above, nearly glabrous beneath; bracts of the pistillate flowers narrower, lanceolate or ovate, acute or acuminate. H. Lupulus neomexicanus A. Nels. & Cockerell. Wild Hops. Among bushes: Wyo. — Utah — Ariz. — -N.M. Plain — Submont. Jl-Au.

Literature Cited:
- Harrington, H. D., 1964, 2nd ed..  

Harrington (1964, 2nd edition) ...

2 Humulus L. HOP
Perennial twining vines; stems and petioles retrorsely prickly (though weak); leaves opposite, palmately lobed; staminate flowers in loose panicles; pistillate flowers 2 together under a large persistent bract which with the others at maturity forms a large conelike “hop.”
1. Humulus americanus Nutt., Journ. Acad. Nat. Sc. Phil. ser. 2. 1:181. 1847.
H. lupulus var. neomexicanus Nels. & Ckll.; H. neomexicanus (A. Nels. & Ckll.) Rydb. — Vines 5-10 m. long; leaf blades 3- to 7-divided or parted usually over ½ way to base. with 3 main lobes, the divisions lanceolate or narrow, terminal one narrower at base than at middle, over twice as long as wide, acuminate or narrowly acute, somewhat scabrous above, lower surface usually copiously glandular, lobes of leaves serrate, petiole about as long or shorter than the blades, rough; bracts of pistillate flowers ovate or lanceolate, acute or acuminate. Our plants have rather variable leaves, some with few obtuse shallow lobes resembling the cultivated species, varying to the typical native form described above. It has been suggested that hybridization may be responsible for this variation. — Usually among bushes. Wyoming to Utah, south to New Mexico and Arizona. Our records scattered over the mountainous part of Colorado at 4000-9000 feet.

Literature Cited:
- Small, Ernest, 1978.  

Small (1978, ____) published H lupulus var. lupuloides. A numerical and nomenclatural analysis of morpho-geographic taxa of Humulus. — behind a paywall.

Literature Cited:
- Small Ernest, 1980.  

The relationships of hop cultivars and wild variants of Humulus lupulus

Literature Cited:
- Löve, Áskell, 1982a.  

Löve (1982, p. ___) published H. lupulus subsp. americanus (Nutt.) Á.Löve & D.Löve. — behind a paywall.

Literature Cited:
- Small, Ernest, 1993+.  

Key in FNANM

1 Leaf blades usually with fewer than 20 hairs per cm on length of midrib, fewer than 25 glands per 10 sq. mm between veins; nodes relatively limited in pubescence, usually fewer than 15 hairs per 0.1 sq. mm at most pubescent portion (excluding angle of petiole with stem). Humulus lupulus var. lupulus
1 Leaf blades usually with more than 20 hairs per cm on length of midrib, more than 25 glands per 10 sq. mm between veins; nodes relatively pubescent, usually more than 15 hairs per 0.1 sq. mm at most pubescent portion (excluding angle of petiole with stem). > 2
2 Leaf blades 10 cm or more usually having at least 5 lobes; smaller blades (ca. 5 cm) usually with more than 3 easily visible veins branching off midrib (excluding proximal branches). Humulus lupulus var. neomexicanus
2 Leaf blades 10 cm or more usually having fewer than 5 lobes; smaller blades (ca. 5 cm) often with no more than 3 easily visible veins branching off midrib (excluding proximal branches). > 3
3 Leaf blades conspicuously pubescent abaxially, more than 100 hairs per cm on length of medial midrib, hairs present between veins. Humulus lupulus var. pubescens
3 Leaf blades not conspicuously pubescent abaxially, usually fewer than 100 hairs per cm on length of medial midrib, hairs usually absent between veins. Humulus lupulus var. lupuloides

Literature Cited:
- Murakami, Atsushi, Branka Javornik, Paige Darby, and Maria Pais, 2006.  

Molecular phylogeny of wild Hops, Humulus lupulus L.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012) refer our local “hops” to Humulus lupulus var. neomexicanus Nelson & Cockerell.

HUMULUS L. 1753 [late Latin name of Teutonic origin]. Hops.
One species, H. lupulus var. neomexicanus Nelson & Cockerell [Lupulus, an early generic name.] 20B> This is a distinctly native race of the species, having been found fossilized in the upper Eocene formation at Florissant. Bases of talus slopes. The hops of commerce is a European form. (H. lupulus subsp. americanus apparently refers to an eastern North American race.)

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield, 2015, pp. 263 ...

HUMULUS L. — Hop
Twinning vines; leaves, simple, palmately lobed; Pistillate flowers in pairs, each pair subtended by a large, foliaceous bract, these imbricate in short spikes or drooping racemes; fruit an achene enclosed within a persistent bract.
Humulus neomexicanus (A. Nelson & Cockerell) Rydb. New Mexican hop. Vines with scabrous stems; leaves 5-15 cm ;ong, hirsute, with serrate margins; staminate flowers on pedicels 0.5-3.5 mm long, with sepals 1.5-3 mm long; pistillate flowers subtended by ovate, papery (at maturity) bracts 7-20 mm long; achenes 203 mm long smooth. Cool, moist, shaded canyons, rocky outcrops, 4800-8500 ft. June-Aug. E/W.

Literature Cited:
- Tembrock, L. R., J. M. McAteer, & T. M. Gilligan, 2016.  

Tembrock, et al., 2016, p. 19 ...

Key to the Native and Naturalized Species of Humulus
1. Petiole as long as or longer than lamina of leaf; pubescence on the abaxial side of the leaf midrib consisting of stiff spinulose trichomes ... *H. scandens
1. Petiole shorter than lamina of the leaf; pubescence on the abaxial side of leaf veins made up of soft silvery trichomes.
2. Glands on the abaxial leaf surface with fewer than 20 glands per cm2; 15 to 25 hairs per linear cm on the abaxial surface of the leaf midrib; nodes weakly pubescent ... *H. lupulus
2. Glands on abaxial leaf surface greater than 25 glands per cm2; 30 to 50+ hairs per linear cm on the abaxial surface of the leaf midrib; nodes with dense silvery pubescence.
3. Most leaves possess 5 or more lobes; glands on abaxial leaf surface in excess of 30 per cm2 ... H. neomexicanus
3. Most leaves possess 3 lobes or remain unlobed and cordate, rarely leaves of 4 lobes are found; glands on abaxial leaf surface fewer than 30 per cm2.
4. Abaxial surface of leaf midrib with more than 100 hairs per linear cm, these hairs spreading and silvery, abaxial surface of leaf evidently pubescent between veins ... H. pubescens
4. Abaxial surface of leaf midrib with 20 to 75 hairs per linear cm; these hairs closely appressed to the midrib, abaxial surface of leaf sparsely pubescent or glabrous between veins ... H. lupuloides
* Introduced species are indicated with an asterisk.

Literature Cited:
- Ackerfield, Jennifer, 2022.  

Ackerfield, 2022, pp. 282-283 ...

HUMULUS L. — Hop
Twinning vines; leaves, simple, palmately lobed; Pistillate flowers in pairs, each pair subtended by a large, foliaceous bract, these imbricate in short spikes or drooping racemes; fruit an achene enclosed within a persistent bract. (Tembrock et al., 2016)
Humulus neomexicanus (A. Nelson & Cockerell) Rydb. New Mexican hop. [Humulus lupulus L. var. neomexicanus A. Nelson & Cockerell] Vines with scabrous stems; leaves 5-15 cm ;ong, hirsute, with serrate margins; staminate flowers on pedicels 0.5-3.5 mm long, with sepals 1.5-3 mm long; pistillate flowers subtended by ovate, papery (at maturity) bracts 7-20 mm long; achenes 203 mm long smooth. Cool, moist, shaded canyons, rocky outcrops, 4800-8500 ft. June-Aug.

Literature Cited:
- SEINet, 2019+.
Full Size ImageNames applied to Colorado collections Humulus sp.  

 

 

Other articles:
• Golden Checklist Flora:  Humulus lupulus;  

Humulus lupulus L. “Wild Hops”

 

   

Humulus neomexicana (A. Nelson & Cockerell) Rydb. “New Mexican Hops”

 

 

Other articles:
• Golden Checklist Flora:  Parietaria pensylvanica;  

Parietaria pensylvanica Willd. “Pennsylvania Pellitory”

 

 

Other articles:
• Golden Checklist Flora:  Urtica dioica ssp. gracilis;  

Urtica dioica L. ssp. gracilis (Aiton) Seland. “Stinging Nettle”

 

 

Other articles:
• Golden Checklist Flora:  Arceuthobium vaginatum var. cryptopodium;  

Arceuthobium vaginatum (Willd.) J. Presl var. cryptopodium (Engelm.) Cronquist “Pineland Dwarf Mistletoe”

 

Literature Cited:
- Gray, Asa, 1850.  

Gray (1850, v. 6, p. 214) includes a long communication from George Engelmann regarding the mistletoes.

“4. A. cryptopodum (n. sp.) : caule ramisque acute quadrangulatis robustis articulis brevioribus ; squamis truncatis in vaginulas cupulatas connatis ; floribus in spicas densas compositas, foemineis ovatis in quavis axilla singulis ; baccis brevissime incluso-pedicellatis erectis. — Santa Fe, only on Pinus brachyptera, A. Fendler, No. 283. — Hooker's A. Oxycedri from the Hudson Bay country appears to belong here : the figure shows at least subsessile, erect fruits ; but the segments of the male flowers are broadly oval, whilte those of the New Mexican plant are lanceolate.” G. Engelmann.

 

Other articles:
• Golden Checklist Flora:  Comandra umbellata ssp. pallida;  

Comandra umbellata (L.) Nutt. ssp. pallida (A. DC.) Piehl. “Pale Bastard Toadflax”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753) first described Comandra umbellata from plants collected in Virginia and Pennsylvania.

Original Text Comments
umbellatum THESIUM floribus umbellatis, foliis oblongis.
Centaurium luteum afcyroides virginianum. Plnk. mant. 43. t. 342. f. 1.
Habitat in Virginiae, Penfylvaniae pafcuis ficcis Kalm.
Radix . Folia alterna, ovali-lanceolata, integerrima. Rami alterni, in fummitate caulis: Umbellulae terminales: involucro tetraphyllo, parvo: Flofculi 5.
“Kalm Petr. Plante canadenfes propediem edendae.” Although, I think his first name was actually “Pehr” and that his name is the source of the genus Kalmia.

Literature Cited:
- Nuttall, Thomas, 1818.  

Nuttall (1818) defined a new genus, Comandra, and places the North American Thesium into it. Nuttall also suggests that some of Linnaeus' eastern hemisphere Thesium might belong in Comandra. The genus Thesium L. remains accepted, and is native to Europe, Asia-Tropical, Africa, South America and Asia-Temperate.

Original Text Comments
233. *COMANDRA.† Thesium. L. Calix angular, tubular-campanulate, coalescing with an internal 5-toothed, glandulous disk. Petals 5, ovate, ingrafted upon the margin of the calix, persistent. Anthers attached to the petals by a tuft of filaments! Germ 3-seeded, immersed in the glandulous disk. Capsule valveless, 1-seeded, coated by the base of the calix.
Perennial, root ligneous, stem herbaceous; leaves simple, alternate, stipules none; radical gemmaceous scales numerous, persistent; flowers in a corymbulose terminal panicle.
Species. 1. C. umbellata
Thesium umbellatum. Linn. Willd. Sp. Plant.
Stem round and erect, sending out 2 or 3 infertile beanches below the panicle. Leaves approximating, erect, oblong-ovate, obtuse, smooth, reflected in the margin, and reticulately veined. Panicle short, ramuli axillary, corymbulose, corymbs about 5-flowered, with 4 involucrate bractes, uppermost peduncles fewer flowered. Calix uniting with the glandulous and nectariferous germinal disk; disk 5-toothed, obtuse. Petals 5, calycine, often 4 and 6, with the same number of stamina, ovate, acute, persistent, growing to the margin of the calix, white, internally villous (Seen through a lens), before expansion parallel. Stamina seated at the base of the petals, alternating with the dentures of the glandulous disk; filaments subulate, about half the length of the petals; anthers oval, 2-celled, connected at their summits to the petals near their base by a fascicle of yellow filaments — Style terete, simple; stigma round, entire; germ about 3-seeded, ovula pendulous, attached to the apex of a filiform contractile funiculus arising from the base of the capsule. Capsule nearly globular, and angular, 1-seeded, not opening, thin and brittle, not osseus, coated by the base of the calix. Seed round, about the size of a small pea, consisting almost entirely of a large carnose and oily perisperm, embryon inverted, small flat, nearly in the axis of the perisperm; radicle superior, thick and obtuse; cotyledons linear and acute.
Obs. The connecting fibres of the petals, appear to be a separation of a portion of the central vessels, for at that point the petal is greenish and callous, and the central nerves there commencing trichotomously, disappear above the connectile fibres, and the rest of the petal is then white.
This plant has some relation to the preceding, and they both appertain to the Natural Order Santalaceae of R. Brown, approaching at the same time very nearly to the Rhamnei of Jussieu. The genus here proposed may probably include some of the species of Thesium indigenous to the Cape of Good Hope.

Literature Cited:
- DeCandolle, Augustus Pyramus, 1857.
- Oliphant, J. Olin, 1934.
- Wikipedia contributors, 2020.

Other articles:
• Golden Checklist Flora:  Notes on Nothocalais cuspidata, Gray, 1884;
• Glossary:  Suffrutex;
• U. S. Highway 95:   in Lapwai;

Locations: Lapwai.  

DeCandolle (1857) proposed C. pallida from a collection in Oregon Territory.

Original Text Comments
3. C. pallida, foliis lividis inferioribus elliptico-oblongis acutis supremis lineari-acuminatis, lobis perigonii erecto-patentibus. Suffrutex Prope Clear Water, Oregon (Rev. Spalding ! Sub nom. Erroneo C. livide Hook. In h. meo et Boiss.). Caules semipedales, sublignosi, striati, erecti, apice solum ramosi. Folia non pellucida, ima squamaeformia, late ovata, 2 lin. Loga ; media 8-10 lin. Longa, 2-3 lin. Lata, mucronata-acuta, summa variabilia, pleraque 1-2 lin. Solum lata et 5-7 lin. Longa, quaedam latiora vel angustiora breviora. Cymae pauciflorae. Bracteae lineari-lanceolatae, 2 lin. Longae. Pedicelli graciles, lineam longi. Flos ut in C. umbellata, sed lobi magis papilloso-velutini praesertum ad marginem et intus. Differt a C. umbellata praecipue foliis angustis elongatis ramorum non florentium. (v. s.) I assume “Rev. Spalding” was Henry H. Spalding who travelled the Oregon Trail in 1836 and established a mission to the Nez Perce at Lapwai near present-day Lewiston, Idaho. Present-day Lapwai is several miles south of the Clearwater River, just up-river from Lewiston.

“The name C. livide Hook. is erroneous.” This refers to the currently named Geocaulon lividum (Richardson) Fernald.

“In my herbarium and that of Bossier.” The type is now at G: Conservatoire & Jardin botaniques de la Ville de Genève. Piehl (1965) states there is a photograph of this type at MICH, but I have been unable to find it online.

β. angustifolia, foliis mediis et superioribus omnibus lineari-acutis. Suffrutex In Novo Mexico (Wright ! N. 1783 in h. Boiss.). Folia ima ut in specimine oregonensi. (v. s.)  

Another first collection by Spalding was Nothocalais troximonoides.

Literature Cited:
- Oliphant, J. Olin, 1934.  

Oliphant (1934) published a paper on the botanical labors of the Reverend Henry H. Spalding. Spalding collected for one year and sent one box of dried plants east. He was apparently stimulated to this activity by contact with Charles A. Geyer, a German botanist, who was in Oregon Country in 1843-44. Spalding sent one box in 1846, which was received in Boston in 1848. By the time that Gray's encouragement for Spalding to continue collecting was sent to him in March 13, 1849, a massacre at Waiilatpu had sent Spalding and others fleeing to Lower Oregon ending his botanical career.

I don't know how Spalding's collections ended up in DeCandolle's herbarium, and why it was he and not Asa Gray who published Spalding's Comandra collections.

Literature Cited:
- Piehl, Martin A., 1965.  

Piehl (1965) wrote a natural history and taxonomy of Comandra. It is almost a monograph, but I don't think we do monographs about single species.

Original Text Comments
The name Comandra is from the Greek Kome (hair) and aner (man), alluding to the hairs superficially attached to the anthers, which actually are not staminal, but develop from the base of the sepals … … The name has frequently been misspelled “Commandra.”  
4. Comandra umbellata (L.) Nutt. Subsp. pallida (A. DC.) Piehl, comb. nov.

Comandra pallida A. DC., in DC. Prodr. 14:636-37. 1857
Comandra pallida β angustifolia A. DC., in DC. Prodr 14:637. 1857. Isotypes: New Mexico, in 1851-52, C. Wright 1783 (GH!, NY!).
Comandra umbellata (L.) Nutt. Var. angustifolia (A. DC.) Torr. Rept. U. S. Mex. Bound. Surv. (Bot.) 2 (Part 1): 185. 1859.
Comandra umbellata (L.) Nutt. Var. pallida (A. DC.) M. E. Jones, Proc. Calif. Acad., Ser. 2, 5:722. 1895.
Comandra linearis Rydb., Fl. Rocky Mts. 818, 1066. 1917. Holotype: Green River, Utah, 19 Aug. 1887, S. M. Tracy and Evans 716 (NY!).

 
Holotype: Clear Water, Oregon (now Idaho), s.d., Rev. Spalding s.n. (G!; photograph 1454 (MICH)) ; two probable isotypes (GH!). Another specimen labeled “plains — Oregon, May 7,” (GH!) could also be type material.  
… subsp. pallida intergrades with both subsp. californica and subsp. umbellata … is also quite variable, but more of the vaiation appears to be correlated with various habitats … contracts with the other species in that it characteristically sprouts from buds which are at or below the soil surface … the aerial part of the plant dying each season ….  
Selected Specimens Examined. COLORADO. Denver Co.: hilltops, Clear Creek near Denver, Clokey & Bethel 3260 (CAS, GH). Collection date: 8 June 1919, also at RM; Piehl does not cite any specimens from RM.

Literature Cited:
- Piehl, Martin A., 1965.  

 

Literature Cited:
- Der, Joshua P., and Daniel L. Nickrent, 2008.  

Der and Nickrent (2008) … phylogenetic analysis supports a Comandra clade consisting of Comandra and Geocaulon lividum, i.e., the C. livide rejected by DeCandolle. Their work also supports a Thesium clade distinct from a Comandra clade.

   

Eriogonum Michx. “Hairy Plant”

 

Literature Cited:
- Michaux, Andre, 1803.  

Michaux (1803, Vol. 1, pg. 246) published the first Eriogonum meaning “hairy plant.”

Original Text Comments
246 ENNEANDRIA. MONOGYNIA.  
  ERIOGONUM.  
  Εριον, lana, Γονυ, genu : planta lanata , geniculata.  
  Caulis herbaceus , dichotomus. Folia terna , verticillata , sessilia , basi subcommata. Flores singulorum fasciculorum e communi involucro subcampanulato erumpentes.  
  Cal. subcampanulatus, 6-partitus : laciniis ovalibus , obtusis ; tribus interioribus paulo majoribus.  
  Stam. 9 : filamenta capillaria , calyce paulo longiora : antheræ breves , ovate.  
  Pist. ovarium 3-quetrum : stylus brevissimus : stigmata 3 , longiora , subfiliformia.  
  Fruct. semem calyce tectum , acute triquetrum , apterum.  
  Obs. Genus inter Polygoneas defectu vaginæ stipularis insolitum.

Tomentosum.
tab. 24.
E. erectum ; partibus omnibus tomentosis : foliis ternis, cuneato-obovalibus.  
  Obs. Flores candicantes.
Hab. in aridissimis , pinetis Carolinæ et Georgiæ.
 

Literature Cited:
- Reveal, J. L., 2005.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum subgroup Eucycla;  Notes on Eriogonum subgroup Oligogonum;  Notes on Eriogonum subgroup Pterogonum;  

Subgroups of Eriogonum as described by Reveal (2005) in his treatment of the genus in Flora of North America.

  • Eriogonum
  • Oligogonum
  • Ganysma
  • Oregonium
  • Eucycla
  • Micrantha
  • Clastomyelon
  • Pterogonum

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum alatum;  Notes on Eriogonum arcuatum;  Notes on Eriogonum effusum;  Notes on Eriogonum umbellatum;  

Those that have been collected in Golden are found in three of the subgroups:

  • Oligogonum Nutt.
    • E. arcuatum Greene. Baker's Buckwheat.
    • E. umbellatum Torrey. Sulphur-Flower Buckwheat.
  • Eucycla Nutt.
    • E. effusum Nutt. Spreading Buckwheat.
  • Pterogonum H. Gross
    • E. alatum Torrey. Winged Buckwheat.

Two other wild buckwheats that are often of interest are both in subgroup Oligogonum Nutt.:

  • E. flavum Nutt. Golden Buckwheat.
  • E. jamesii Benth. James' Buckwheat.
Both of these are known from Jefferson County though not from Golden s.l.

 

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum, Reveal, 2005;  

Eriogonum subgroup Eucycla Nutt.

 

Literature Cited:
- Nuttall, Thomas, 1848b.

Other articles:
• Golden Checklist Flora:  Nuttall, 1848, publication details;  

Nuttall (1848b, p. 166) published Eucycla as a genus name for wild buckwheats he had collected, probably on his 1834 expedition to Oregon Territory.

Original Text Comments
*EUCYCLA.†  
Perianth membranaceous, coloured, petaloid, dimorphous, the three outer divisions orbicular, concave; the three inner linear-oblong, emarginate, connivent into a cylinder. Stamens nine; with short filaments, membranous at base. Styles three, of moderate length, with small, capitate stigmas. Achenium attenuated, triangular. Embryo excentric ; radicle superior ; cotyledons flat.  
E. *ovalifolia. — Leaves all redical, short and roundish-ovate, whitely tomentose; capitulum made up of several sessile, whitely tomentose involucres; outer segments of the yellow perianth rather narrower at base, the inner emarginate segments exserted.
Eriogonum ovalifolium. Nutt. Journ. Acad. Nat. Sci. Philad.
Hab. Sources of the Missouri. Flowers bright yellow.
Eriogonum ovalifolium Nutt. is an accepted name, common in south and east California.
E. *purpurea. Leaves all radical, short and roundish-ovate, whitely tomentose; capitulum made up of several sessile, smoothish, tomentosely margined involucres; outer segments of the purple perianth orbicular, sometimes emarginate at base; the inner emarginate, narrow, segments scarcely exserted.
Scape about a span high, arising from a multifid woody caudex; flower larger than in the preceding, and purple; filaments much shorter than the perianth, with a torn membranous margin, at base three stamens seated on each of the inner narow segments; embryo rather short.
Hab. Rocky Mountains.
This is a synonym of Eriogonum ovalifolium var. purpureum (Nutt.) Durand
† In reference to the circular figure of the perianth.  

Literature Cited:
- Flora of North America Editorial Committee, eds., 2005.  

The characters of subgroup Eucycla are primarily: perennial, stems not jointed, flowers without a stipe-like base.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum effusum;  

The one representative of subgroup Eucycla that is found in Golden s.l. is E. effusum Nutt. “Spreading Buckwheat.”

Other wild buckwheats in subgroup Eucycla in the author's experience are E. microthecum Nutt. “Slender Buckwheat” from the Mono Lake Basin and E. wrightii Torr. ex Benth. “Bastardsage” from the eastern Mojave Desert. All three of these wild buckwheats are very similar appearing.

 

Literature Cited:
- Nuttall, Thomas, 1848b.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum, Reveal, 2005;  

Eriogonum subgroup Oligogonum Nutt.

Nuttall (1848b, p. 165) discusses the differences between E. tomentosum on which Michaux (1803) founded the genus Eriogonum, noting some key differences and proposing a subgenus Oligogonum which contains some of our more common wild Buckwheats.

Original Text Comments
The character of the genus Eriogonum, of which there are so many species, perhaps deserves some additional consideration. The genus, as founded on E. tomentosum of Michaux, possesses yet some peculiarities not common to the rest of the genus; among the rest of its characters I may remark, that in place of the interior segments of the perianth being smaller, it is the reverse; the perianth increases in size with the perfecting of the fruit, and as in Rumex, the three inner segments are larger and erect, the three outer reflected ; the next discrepancy in the character, as given by authors, is in the condition of the embryo, which is placed in the centre of axis of the seed, and is not as described excentric.  
In E. longifolium, besides a remarkable difference in habit, the perianth is wholly herbaceous, very lanugunous, and the segments all so very equal, as to appear, at length, almost disposed in a simple series; the achenium is also lanuginous, and the seed presents, as in E. tomentosum, a concentric embryo. For E. longifolium I would therefore propose the name of Trachytheca, excluding every other species. It is the first species of the section Eriantha of Bentham, which name, in not being exclusive, and too near Erianthus, cannot properly be employed for the present plant. It is very peculiar in its solitary leaves and alternate beanches.  
In all the rest of this numerous genus, the outer segments of the perianth, which increase a little in size, are either larger and erect, or all nearly equal. In these, some of which have the embryo excentric and others concentric, the subgeneric name of Olygogonum may be applied, but whether any other subdivision may be made by the condition of the embryo is at present uncertain. In E. flavum, I believe the embruo is excentric, but how far that species connects itself to any others of similar form, remains to be examined. Olygogonum has become Oligogonum and is now the name of an Eriogonum subgroup that contains E. arcuatum, E. flavum, and E. umbellatum.

Oligogonum is the larger of the subgroups and contains many of the caespitose or mat-forming perennials, those with a stipe-like base on the flowers.

 

Literature Cited:
- Gross-Konigsberg, Hugo, 1913.

Other articles:
• Golden Checklist Flora:  Notes on Eriogonum, Reveal, 2005;  

Eriogonum subgroup Pterogonum Gross

Gross (1913, vol. 49, pg. 239, footnote 1) in his “Contributions to the knowledge of the Polygonaceae” proposed Pterogonum for some Eriogoneae but I cannot read the original German well enough to understand the basis for segregation from the remainder of Eriogoneae.

Original Text Comments
1) Pterogonum H. Gross nov. gen. Eriogonearum. Involucrum pluriflorum, gamophyllum, ± campanulatum, 5-dentatum. Perigonium profunde 6-partitum, tepalis ternis in 2 cyclis dispositis. Stamina 9. Ovarium trigonum, 1-loculare, ovula unico basilari orthotropo, stylis 3 capitato-stigmatosis. Achaenium maturum perigono longius, trialatum. — Herbae perennes Eriogonis simillimae, indumento subsericeo. Folia radicalia, caulina pauca, altrerna. Rami floriferi apice caulis in di-usque pleiochasium conferti nonnumquam etiam racemosi. Embryo fere rectus v. subexcentricus, cotyledonibus sat amplis.  
Species adhuc certae: Pterogonum alatum (Torr. sub Eriogono) Gross, P. atrorubens (Engelm. sub. Erigono) Gross, P. hieracifolium (Bth. sub. Eriogono) Gross.  

Literature Cited:
- Reveal, J. L., 2005.  

Reveal (2005) places six North American taxa in subgroup Pterogonum of which only one, E. alatum occurs in Colorado.

 

Other articles:
• Golden Checklist Flora:  Eriogonum alatum;  Notes on Eriogonum in Golden s.l.;  

Eriogonum alatum Torr. “Winged Buckwheat”

 

Literature Cited:
- Fremont, John C., 1845.

Other articles:
• Golden Checklist Flora:  Fremont, 1845, publication details;  

No mention of Eriogonum alatum in the report of Frémont's 1842 and 1843-'44 expeditions, though Torrey (1853) notes they were aware of it.

Literature Cited:
- Gray, Asa, 1852-1853.

Other articles:
• Golden Checklist Flora:  Gray, 1852-1853, Plantæ Wrightianæ;  

No mention of Eriogonum alatum in Gray's (1852) account of Wright's collections in Texas and New Mexico in 1849. In fact, there are no Polygonaceae in either part of Gray's account, although DeCandolle (1857) will say he has seen a collection of Eriogonum alatum by Wright. Gray's preface notes that the cacti were given to George Engelmann, of St. Louis, for examination. The seeds were divided between the Botanic Garden of Harvard University, under Gray's charge, and the Royal Botanic Gardens at Kew, under the direction of Sir Wm. Hooker. Sets of plants were distributed to Gray's herbarium, the Smithsonian Institution, John A. Lowell, a patron of the Wright's explorations, with a few distributions to others. If one assumes that Wright collected any Polygonaceae, what happened to them?

Literature Cited:
- Sitgreaves, L., 1853.

Other articles:
• Golden Checklist Flora:  Sitgreaves, 1853, publication details;  

Dr. John Torrey wrote the Botany chapter of the Sitgreaves Report on the expedition Sitgreaves led down the Zuni and Colorado Rivers in 1851.

Original Text Comments
E. alatum. (Torr., l. c.;) perenne ; caule erecto subflexuoso folioso, ramis alternis erectis paniculatis ; foliis spathulatis hirsutis ; pedunculis terminalibus ternis ; involucris solitariis campanulatis 5-fidis ; perigoniis glabris, laciniis aequalibus ; acheniis trialatis.

On the Zuñi river; September. Root stout and blackish, descending to a great depth ; stem 1--3 feet high, arising from a short thick caudex, which is clothed with the remains of leaves. Radical leaves 2--4 inches long, and 3--5 lines wide, almost villous, with long hairs, mostly obtuse ; stem leaves much smaller, and gradually diminishing in size upward, all of them erect. Branches solitary and distant, subdivided in a trichotomous manner, each division bearing a single involucre, which is about 2½ lines long, and pubescent. Pedicels glabrous, a little exserted, jointed close to the flower ; perigonium not enlarging after flowering ; the segments lanceolate ; filaments glabrous ; ovary oblong, triquetrous, longer than the styles ; achenium nearly four lines long, with three very conspicuous membranaceous wings ; seed ovate, triangular ; embryo straight.

Torrey's “l. c.” refers to the second previous entry, i.e., that for Eriogonum orthocladon for which the citation is “(Torr. mss., in D. C. Prodr. ined.:).” I interpret this as “I, Torrey, described this in a manuscript I gave to DeCandolle for his use in his Prodromo.” As it happens, though, Torrey's account in the Sitgreaves Report was published in 1853, four years before DeCandolle's Prodromo volume 14 containing Torrey's manuscript description.
This remarkable species was first detected by Colonel Frémont in upland prairies, at the sources of the Plata, in 1843, and again in 1845 in “Bahia Salada,” in the Rocky mountains. Lieutenant Abert found it on the Raton mountains in 1846.  

Literature Cited:
- DeCandolle, Augustus Pyramus, 1857.

Other articles:
• Golden Checklist Flora:  DeCandolle, 1857, publication details;  

DeCandolle (1857, vol. 14., p. 6) …

Original Text Comments
2. E. alatum (Torr.! in Frémont Rep.), herbaceum, elatum, subsericeo-villosum, foliis subradicalibus petiolatis oblongis lanceolatisve elongatis, caulinis paucis, pedrunculis axillaribus terminalibusque apice 2-3-chotomis umbellatisve, involucris pedicellatis campanulatis 5-dentatis, perigoniis parvis glabris, achaenio 3-4-plo longiore a basi trialato. ♃ In montibus Scopulosis in regione superiore fl. Platte meridionalis et Bahia Salada (Frémont !), in Novo-Mexico (Abert! Fendler ! n. 763 , Wright !). Habitus fere E. hieracifolii, sed præter corymbum terminalem adsunt etiam peduncili breves laterales. Perigonia multo minora, sub anthesi vix lineam longa, nec multum sub fructu aucta. Achænium 3-4 lin. longum, a basi as apicem late trialatum. (v. s.)  
β elatum. caule sub-3-pedali, foliis radicalibus 6-8 poll. longis. In montibus Scopulosis inter fl. Platte et Sweetwater (Geyer! n. 145). — Journ. 1853, p. 263.  

DeCandolle notes that he has seen Fendler's No. 763 and agrees it is E. alatum. Fendler's collections went to Gray, who in his Plantae Fendlerinae make no mention of it, or any other Polygonaceae. There are two vouchers of Fendler's 763, one at Harvard University (GH) and one at Brown University (BRU). The locality of both vouchers states “2 mi east of the Moro River,” which river does not now exist in New Mexico. However, there is a Mora River that is crossed by the Santa Fe Trail and given the August date of the GH voucher, it is likely that Fendler collected it on his way home. What I don't understand is why Torrey did not publish the name from Fendler's collection, but instead published the name from the Woodhouse collection on the Sitgreaves Expedition of 1851.

 

Other articles:
• Golden Checklist Flora:  Eriogonum arcuatum;  Notes on Eriogonum in Golden s.l.;
• Field Notes:   Coll. No. 1401.1, 8 Jun 2016;  Coll. No. 1695, 29 Jun 2017;
Full Size ImageInflorescence of Coll. No. 1401.1, Eriogonum arcuatum
Full Size ImageColl. No. 1695, Eriogonum arcuatum  

Eriogonum arcuatum Greene. “Baker's Buckwheat”

 

Literature Cited:
- Greene, Edward L., 1901.

Locations: Pagosa Springs.  

Greene (1901, v. 4., part 25, p. 319) described E. arcuatum from a collection on hillsides about Pagosa Springs, Colorado by C. F. Baker.

Original Text
Eriogonum arcuatum. Near E. flavum, about as large, more extensively caespitose, forming broad matted tufts : leaves oval, obtuse, an inch long or less, abruptly tapering to a rather slender petiole about as long, white-tomentose beneath, pale-green and thinly tomentellous above : scapiform peduncles 6 inches high, bearing a single large sessile involucre and a pair of long-peduncled ones arising from its base, these opposite each other and curving upwards to the length of 1½ to 2 inches : perianths yellow, very sparsely silky-villous : stamens long-exserted.
On hillsides about Pagosa Springs, Colorado, 17 July, 1899. C. F. Baker.

Literature Cited:
- Greene, Edward L., 1901.  

Six weeks later, Greene (1901a) published Eriogonum bakeri.

Original Text
Eriogonum Bakeri. Allied to E. flavum, rahter taller, the branches of the caudex very slender and only loosely leafy, the leaves thin, the elliptic-lanceolate blades 1/2 to 1 inch long, on slender petioles much longer, white-tomentose beneath, sparsely villous above: scapiform peduncles 5 to 8 inches high, erect slender; inflorescence of a sessile involucre and 1 to 3 dichotomous peduncles from its base, the whole number of involucres thus 7 to 9, all turbinate: perianths yellow, small very long-stipitate, silky villous, the inner segments much longer than the outer, all obovate, obtuse.
Black Cañon, 1 Aug., n. 696. Said to be caespitose in rather small tufts. The inflorescence is like that of E. Jamesii, though far less ample; and the real affinity is with E. flavum.

We now treat E. bakeri as a synonym of E. arcuatum.

Literature Cited:
- Reveal, James L., 2004a.  

Reveal (2004a, p. 157) reviewed all the available names in the subfamily Eriogonoideae in preparation for his contribution to FNANM. Reproduced below are his analyses of E. arcuatum and E. flavum.

Original Text
Eriogonum arcuatum Greene, Pittonia 4: 319. 1901. – T.: Hillsides about Pagosa Springs, Archuleta Co., Colorado, 17 Jul 1899, C. F. Baker 284. Holotype: NDG (sheet number 118); isotypes: B, BM, E, F, G, GH, K, MO, NDG, NMC, NY, P, POM, RM(2), UC, US. – E. jamesii Benth. Var. arcuatum (Greene) S. Stokes, Eriogonum: 118. 1936.  
Eriogonum bakeri Greene, Pl. Baker. 3: 15. 1901. – T.: Black Canyon, Montrose Co., Colorado, 1 Aug 1901, C. F. Baker 696. Holotype: NDG; isotypes: E, G, GH, K, LY, MIN, MO, NY, POM, RM(2), UC, US, VT, W. – E. jamesii Benth. Subsp. bakeri (Greene) S. Stokes, Eriogonum: 118. 1936. Elsewhere, Reveal noted that Greene published E. bakeri six weeks after E. arcuatum.
 
Eriogonum flavum Nutt., Cat. Pl. Upper Louisiana: [2,] no. 34. 1813. – T.: Variously stated as “in the vicinity of the rocky mountains” or “from the Arikare village,” most likely found in the vicinity of the site of Fort Mandan, Mercer or McLean counties, North Dakota, Jul-Aug 1811, T. Nuttall s.n. Holotype: PH; isotype: P. – A Nuttall sheet at PH with a label alluding to the Rocky Mountains was collected in 1834 and is not original material.  
Reveal lists eight synonyms of E. flavum, two of which are particularly interesting.  
 
Eriogonum laterifolium Raf. New Fl. N. Amer. 4: 53. 1838. – LT.: Upper Missouri River, probably in South Dakota, Jun 1811, J. Bradbury s.n. Lectotype: BM, designated here; isolectotype: LINN.

The original collection in Rafinesque's possession was probably destroyed, not found at P or PH.

This is one of the few published names I have seen that specifically refer to a Bradbury collection.
 
Eriogonum sericeum Pursh, Fl. Amer. Sept. 2:277. 1813. – T.: “On the prairies of the Missouri [River',” probably in the vicinity of the site of Fort Mandan, Mercer or McLean counties, North Dakota, Jul-Aug 1811, T. Nuttall s.n. Holotype: PH; isotype: P. – E. flavum Nutt. Var. sericeum S. Stokes, Eriogonum: 116. 1936. I assume this was Pursh's view of the same collection that Nuttall published as E. flavum.

Literature Cited:
- Ackerfield, Jennifer, 2015.
- Flora of North America Editorial Committee, eds., 1993+.
- SEINet, 2019+.  

The treatment of subfamily Eriogonoideae in Flora of North America was written by Jim Reveal. Eriogonum arcuatum, E. flavum, and E. jamesii are all in subgroup Oligogonum, and couplet 20 separates E. arcuatum and E. flavum. (I might add that the original online key contains an HTML coding error that will throw you off, and has been corrected below. To date, 22 Oct 2022, this HTML coding error has been propagated to the beta version of FNANM.)

Original Text
22 (21) Inflorescences compound-umbellate or, if umbellate or capitate, not of distribution of E. flavum; Colorado Plateau, s Rocky Mountains and w edge of Great Plains from s Wyoming to n Arizona and n New Mexico   129. Eriogonum arcuatum
+ Inflorescences subcapitate or umbellate; n Great Plains and n Rocky Mountains, Wyoming and Nebraska north to Canada and Alaska, west to e Oregon and Washington   133. Eriogonum flavum

The way I read this key, E. arcuatum and E. flavum should not be sympatric. Yet the map of collections determined one of the two taxa in Colorado are definitely geographically intermixed (SEINet, 2020). Similarly, the distribution maps in Ackerfield (2015) show the distributions of the two taxa are quite similar.

Full Size Image
Map of collections determined one of the two taxa of Eriogonum in Colorado

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012) do not distinquish between E. arcatum and E. flavum.

Original Text
7b. Flowers E. flavum (Nuttall) [yellow]. Plains to alpine. As we see it, this species consists of various distinctive local forms that probably evolved in isolation. Alpine plants tend to be of shorter stature, with congested umbels (E. arcuatum var. xanthum (Small) Reveal, E flavum subsp. chloranthum (Greene) Stokes). On the eastern plains the inflorescence varies from simple to compound umbels. In the foothills, taller forms have been called E. arcuatum Greene. One might be justified in considering the local races of no nomenclatural significance, and even treating E. jamesii as E. flavum subsp. jamesii.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015) distinguishes between E. arcuatum and E. flavum on the basis of pedunculate involucres.

Original Text

4a. Involucres elevated on evident peduncles ... E. flavum var. flavum
4b. Involucres sessile or nearly so ... E. arcuatum

 

Other articles:
• Golden Checklist Flora:  Eriogonum effusum;  Nuttall, 1848, publication details;  Notes on Eriogonum in Golden s.l.;  Notes on Eucycla, discussion.;  

Eriogonum effusum Nutt. “Spreading Buckwheat”

 

Literature Cited:
- Nuttall, Thomas, 1848b.  

Nuttall (1848) described Eriogonum effusum in his description of plants collected by William Gambel. It is, however, a Nuttall collection made on the “… Platte plains …” and therefore on his 1834 journey across the Rocky Mountains. An isotype is at the Gray Herbarium (GH) and can be seen through SEINet.

Original Text Comments
E. *effusum. Suffruticose ; leaves linear, oblong, obtuse, beneath whitely tomentose, above pubescent, greenish ; stem tomentose, two or three times trichotomous, divaricate ; bractes ternate, lanceolate-acute ; (flowers not seen.)  
Stem divided into many simple branches below ; flowering stem bearing bractes only, divided compoundly and numerourly, each division subtended by conspicuous trifid bractes.  
Hab. In the Rocky Mountains. (Nuttall.) I assume that “Nuttall” indicates this is a Nuttall collection rather than a Gambel collections.

In this same volume, Nuttall also published Eucycla as a generic name for the wild buckwheat we now call Eriogonum ovalifolium. This name has now become a subgroup name for wild buckwheats that are perennial, without jointed stems, and flowers without a stipe-like base.

 

Other articles:
• Golden Checklist Flora:  Eriogonum umbellatum;  Notes on Eriogonum in Golden s.l.;  

Eriogonum umbellatum Torrey, Ann. Lyceum Nat. Hist. New York. 2: 241. 1827. “Sulphur-Flower Buckwheat”

 

Literature Cited:
- Torrey, John G., 1828.  

Published in 1827 by John Torrey.

Original Text Comments
391. R. umbellatum, caule nudo, simplici ; pedunculis elongatis (sub-senis) umbellatis, apice involucrum singulum gerentibus ; calycibus glabris ; laciniis obovatis, basi angustatis, obtusis ; foliis obovato-spathulatis, subtus lanuginosis, supra glabriusculis.  
Desc. Perennial, herbaceous, cespitose. Caudex divided at the base into several branches ; most of the branches short, bearing fasciculi of leaves at the extremity, one of them elongated and resembling a scape ; covered with a copious loose tomentum. Leaves obovate-spathulate, about an inch long, and less than an inch broad, white tomentose beneath, smoothish above. Flowering stem a span high, bearing a simple umbel at the extremity. Peduncles or rays an inch and a half long, wooly, thick, furnished with several oblong-lanceolate leaves at the base. Involucrum about 20-flowered, campanulate, with lanceolate obtuse teeth. Florets distinctly pedicellate. Calyx very smooth ; segments obovate-oblong, obtuse, narrowed at the base. Stamens 9. Seed acutely triangular.  
Hab. With the preceding. The preceding appears to be 390. E. tenellum which is also described as being “… with the preceding …” E. tenellum is known from southeastern Colorado. The next preceding is
389. E. annuum, n. sp. Nutt. Mss. Near the Rocky Mountains. Found also on the Arkansa by Mr. Nuttall. Root annual !
E. annuum is known from eastern Colorado, but not Jefferson County.
Obs. Flowers larger than in E. tomentosum. It is remarkable that Humboldt and Bonpland did not observer any species of Eriogonum in Mexico, as they abound, accoding to Dr. James, about the sources of the Canadian.  

Published in 1827 by John Torrey.

 

Other articles:
• Golden Checklist Flora:  Eriogonum umbellatum var. ramulosum;  

Eriogonum umbellatum Torr. var. ramulosum Reveal “Buffalo Bill's Sulphur Flower”

 

Literature Cited:
- Reveal, James L., 2004.  

Reveal (2004) proposed variety ramulosum.

Original Text
Eriogonum umbellatum Torr. var. ramulosum Reveal, var. nov. TYPE: UNITED STATES. Colorado, Jefferson Co.: Along U. S. Hwy 6 adjacent to I-70, 0.4 mi E of Exit 256 to Buffalo Bill's Grave, N39°42'12", W105°14'32", T4S, R70W, sec. 18, 7100 ft elev., 25 Jul 1992, J. L. Reveal 7244. HOLOTYPE: NY. Isotypes: BY, CAS, COLO, GH, MARY, MO, RENO, RM, RSA, UC, US, UTC.
A Eriogono umbellato var. umbellato inflorescentibus divisis differt.
Plants low, often rather compact, mats 2-4 dm across; leaves in loose rosettes, the leaf-blades mostly elliptic to oval, 1-2.5 cm long, 0.5-1.5 cm side, densely white to gray tomentose abaxially, floccose and green adaxially; flowering stems erect, mostly 1-3 dm long, floccose; inflorescences compound umbellate and divided 2-3 times; involucral tubes 2-3 mm long, the lobes 1.5-3 mm long; flowers bright yellow, 4-7 mm.
Other specimens seen
UNITED STATES. Colorado, El Paso Co.: Colorado Springs, Jul 1892, Eastwood s.n. (F, MO, RM); Jefferson Co.: Mt. Vernon Country Club, Golden, 7 Aug 1941, Ehlers 8159 (COLO, MICH, TEX, WTU), Larimer Co.: Cherokee Park, 15 Jul 1934, R. J. Davis 394-W (IDS); Estes Park, 16 Jul 1978, Hampton s.n. (NY); 2.1 mi W of Drake, 9 Aug 1967, Neal & Neal 2 (ASU); Estes Park, Aug 1931, E. C. Smith s.n. (MONT); Big Thompson Canyon, 7500 ft elev., 3 Jul 1934, S. Stokes 201a (RM). Mineral Co.: Piedra, 12 Jul 1899, C. F. Baker 291 (RM).
Eriogonum umbellatum var. ramulosum (from the Latin ramulosus, meaning “with small branches”) is encountered mainly but infrequently along the Front Range of the Colorado Rocky Mountains mainly west of Fort Collins south to Colorado Spring then westward to Piedra. Buffalo Bill's sulphur flower is related to var. umbellatum, differing consistently in having a compound umbellate inflorescence.

 

Other articles:
• Golden Checklist Flora:  Eriogonum umbellatum var. umbellatum;  

Eriogonum umbellatum Torr. var. umbellatum. “Sulphur-Flower Buckwheat”

 

 

Other articles:
• Golden Checklist Flora:  Fallopia baldschuanica;  

Fallopia baldschuanica (Regel) Holub

The basionym is Polygonum baldschuanicum Regel published in Trudy Imperatorskago S.-Peterburgskago Botaniceskago Sada. Acta Horti Petropolitani. St. Petersburg 8: 684 (1884)

Literature Cited:
- Regel, E, 1883.
Full Size ImageRegel's illustration of Fallopia baldschuanica.  

POLYGONEAE. Polygonum L. POLYGONEAE. Polygonum L.  
81. Polygonum baldschuanicum Rgl. 81. Polygonum baldschuanicum Rgl.  
Tabula X. Tabula X.  
Glabrum; caule frutescente, lignoso, volubili, basi usque 13 Mm. in diametro, cortice fusco lenticellis crebris ornato. Ochreae brevissime cylindricae, ut bracteae membranaceo-hyalinae mox fissae, denique subevanescentes. Folia inferiora opposita, longe petiolata, hastato- cordata, acuta v. acuminata v. rarius obtusa, margine obsolete crenulato-scabra, petiolum superantia; folia suprema floraliaque valde minora, nunc caulinis similia, nunc lineari-lanceolata in petiolum attenuata. Florum paniculae axillares terminalesque, folium pluries superantes, fere aphyllae, laxae; rhachis angulata, ad angulos minute scabra. Flores ramulorum paniculae fasciculati, pedicellati; fasciculi pluriflori; pedicelli supra basin articulati, apicem versus hyalino-albido trialati. Calycis 5 partiti hyalino-albidi lobis tribus exterioribus ovatis, patentibus, dorso naviculari-alatis; alis in pedicellum decurrentibus; sepalis interioribus duobus erectis, obovatis, exalatis, rubescentibus. Stamina 8. Stigma capitatum, trilobum, in ovarii apice sessile. Calyx fructifer vix auctus, late alatus. Achaenium triquetrum, nigrum, nitidum.    
Folia caulina incluso petiolo usque 7 — 8 Cm. longa; lamina usque 4 Cm. lata et 5 Cm. longa. Flores 5 — 6 Mm. in diametro. Caulis 10—15 pedes altus.    
In Bucharae orientalis chanato Baldschuan ad fluvium Wachsch ad pedem orientalem montium Sevistan, 4—5000' alt., mense Julio anno 1883, leg. A. Regel. In the eastern part of Bukhara, the Baldschuan river runs to the Wachsch river the eastern foot of the Sevistan mountains, 4-5000' alt., in the month of July 1883, leg. A. Regel Bukhara is a city and a region of Uzbekistan located in the southwest of the country.

The Baldschuan River was suggested to be the Zarafshan River by ChatGPT, which I now think is incorrect.

The type of Primula baldschuanica B. Fedtsch. Consp. Fl. Turkestanicae 5:6 (1913) was named for the type location Baldschuan (Boldzhuan), Tajikistan. GoogleMaps spells it as Baldzhuvon at coordinates (38.307612,69.677265). I have also seen the name spelled Baljuan.

The Wachsch River might be the Vakhsh River.

Baldschuan is at the intersection of two rivers. However, I don't see that either of them flow into the Vakhsh River.

Species proxima P. multiflorum Thbrg. «caule herbaceo, floribus triplo minoribus rufescentibus, calyce fructifero valde aucto fuscescente alato» facile dignoscitur.    
Explicatio tabulae X.    
a. Ramus cum panicula, — flos a latere visus, — flos apertus a latere superiore visus. — d. e. stamina. — f. pistillum. — g. h. fructus maturi. — i. achaenium. — Figurae a, g, h, i magnitudine naturali, — b , c, d , e, f auctae. — k. floris digramma.    
   

 

Other articles:
• Golden Checklist Flora:  Fallopia convolvulus;  

Fallopia convolvulus (L.) A.Löve “Black Bindweed”

 

 

Other articles:
• Golden Checklist Flora:  Fallopia japonica;  

Fallopia japonica (Houtt.) Ronse Decr. “Japanese Knotweed”

(Syn: Polygonum cuspidatum Siebold & Zucc., Reynoutria japonica Houtt.)

Literature Cited:
- Houttuyn, Martinus, 1777.  

  Reynoutria
  Naar zekeren Heer van Reynoutre, waar van Lobel getuigt , dat dezelve aan de Kruidkunde zeer veel dienft gedaan heeft ; zo als ik bevoorens heb gemeld (*). To a certain Lord of Reynoutre, where van Lobel testifies that the same to Herbology has done much service; like me in advance have reported (*).

(*) II. D. VII. Stuk, bladz. 64.
II. Deel/. VIII. Stue.
   
IV. Afdeel. XI. Hoofdstuk. Driewyvige., Immers in dit Kruid heb ik de Bloempjes bevonden tien Meeldraadjes of liever Meelknopjes bevatten, en het Vrugtbeginzel is driekandg , met drie Siempels gekroond : uit welken hoofde, anders , dit Gewas tot het Duizendknoop betrokken zou kunnen worden. Het heeft den Kelk vyfbladig, zonder Bloemblaadjes. For in this Herb I have found the Flowers ten Stamen or rather Flour buds contain, and the Vrugtbeginzel is trikandg , crowned with three Siempels: on what head, otherwise, this Crop involved to the Knotweed could become. It has den Chalice with five petals, without petals.
I. Renoutria Japonica. japaniche. Pl. LI. Fig. 1. De eenigde Soort , my daar van bekend , heeft een vooze , ronde, doch eenigszins gegroefde of gellreepte , brumachtige Steng, die bogtig is en knoopig, overhoeks Takken uitgeevende, welke , zo wel als de Steng , overhoeks bezet zyn met lang gefteelde Piekswys' Hartvormige Bladen , de grootften wel vier Duimen lang en derdhalf Duim breed. Hier en daar komen aan de Knoopen , en in de Oxels der Bladen , drie of vier Ristachtige Bloem- en Zaadtrosjes voort, met zeer kleine Bloempjes , van gezegde hoedanigheid. De Bladen verkleinen niet naar boven , maar komen zelfs naar 't end der Takken veel grooter voor , dan om laag. The only kind, known to me, has a voodoo, round, but somewhat grooved or gellreepte, brumig Steng, that bogtig is and knotted, giving out branches diagonally, which, as well as the Steng, occupies across the corner are with long-stemmed Piekswys' Heart-shaped Blades, the sizes up to four inches long and three and a half inches wide. Coming here and there the Buttons , and in the Oxels der Bladen , three or four Ristle-like Flower- and Seed-trusses forth, with very small Flowers, from proverb capacity. The Sheets do not shrink to above , but come even to the end of the Branches much larger in front than down.
* II. D. v. Stuk, bladz. 116. Terwyl ik het Geflagt van Cotyledon , onder den naam van Navelkruid, als veele Heefterigen gen bevattende, reeds befchreeven heb * , zo komt hier thans eerst in aanmerking het. While I read the Geflagt of Cotyledon , below the name of Navelweed, like many of the Hasty gene containing, already described * , zo comes into consideration here first.

 

Other articles:
• Golden Checklist Flora:  Persicaria lapathifolia;  

Persicaria lapathifolia (L.) A. Gray “Pale Smartweed”

(Syn: Polygonum lapathifolium L.)

 

Other articles:
• Golden Checklist Flora:  Persicaria maculosa;  

Persicaria maculosa A. Gray “Lady's Thumb”

(Syn: Polygonum persicaria L.)

 

Other articles:
• Golden Checklist Flora:  Polygonum achoreum;  

Polygonum achoreum S. F. Blake “Leathery Knotweed”

 
 

There is one collection from South Table Mountain, Yeatts #808, 8/20/1983, that is labeled Polygonum erectum ssp. achoreum (COLO00613273,KHD21868).

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012) accept only P. erectum for Colorado, noting the differences are slight.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Treated at the species rank by Ackerfield (2015), P. erectum not accepted for Colorado.

 

Other articles:
• Golden Checklist Flora:  Polygonum aviculare;  

Polygonum aviculare L. “Prostrate Knotweed”

 

 

Other articles:
• Golden Checklist Flora:  Polygonum douglasii;  

Polygonum douglasii Greene “Douglas Knotweed”

 

Literature Cited:
- Greene, Edward L., 1885.  

Greene (1885, v. 1, n. 3, p. 125) ...

P. Douglasii.

Glabrous and somewhat glaucous, often a little scabrous about the nodes, 1-1½ feet high, with numerous, slender, divergent branches: leaves thinnish, oblong to lanceolate, 1-nerved, their margins smooth and more or less revolute; stipules entirely hyaline, the sheathing portion very short, or wanting, the upper part more or less lacerate: floral leaves reduced: flowers commonly more than one in each axil, their pedicels deflexed: akenes longer than in the preceding, shining or granular-roughened. — P. tenue, Watson, Bot. King. 315; Bot. Cal, II. 12, but not of Michx.
var. latifolium.
Leaves oblong : flowers numerous and crowded into a spike : face of akene rather oblong than ovate in outline. — P. tenue, var. latifolium, Engelm.
From the Saskatchewan to British Columbia, and southward everywhere in the mountains to the borders of Mexico. Clearly distinguishable from its eastern analogue by the characters indicated, of which the 3-parallel-nerved leaves and their almost saw-toothed margins are the most obvious.
In P. Douglasii, which we dedicate to perhaps its very earliest collector, the secondary veins, when apparent at all, are not parallel, bnt pinnate. The plant is variable, and yet the var. latifolium may be a distinct species; the shape of the akene is peculiar.

 

Other articles:
• Golden Checklist Flora:  Polygonum engelmannii;  

Polygonum engelmannii Greene “Engelmann's Knotweed”

(Syn: Polygonum douglasii Greene ssp. engelmannii (Greene) J.T.Kartesz & Gandhi )

Literature Cited:
- Greene, Edward L., 1885.  

Greene (1885, v. 1, n. 3, p. 126) ...

P. Engelmanni.
Erect-spreading, diffusely branched from the base, a span or more high, reddish, very minutely scabrous-puberulent throughout: branches slender and somewhat flexuous: stipules sparingly lacerate, short, with no tubular or herbaceous portion; leaves lanceolate, acute, with revolute margins, ½ inch or more long, the upper much smaller and remote: flowers in all the axils, solitary or in twos or threes, very small: pedicels strongly defiexed: sepals obtuse, shorter than the very small, ovate, shining akene, and but loosely investing it. — P. tenue, var. microspermum, Engelm.
Rocky Mountains of Colorado at considerable elevations. Very unlike any forms of the preceding species; differing not more remarkably in the minuteness of its flowers and fruit than in its peculiar erect-spreading habit and the fact of its flowering from the very base of the stems and branches.

 

Other articles:
• Golden Checklist Flora:  Polygonum ramosissimum;  

Polygonum ramosissimum Michx. “Bushy Knotweed”

 

 

Other articles:
• Golden Checklist Flora:  Rumex acetosella;  

Rumex acetosella L. “Sheep Sorrel”

(Syn: Acetosella vulgaris (K. Koch) Fourr., R. angiocarpus Murb.)

Literature Cited:
- Tabermaemontanus, Jacobus Theodorus, 1590.
Full Size ImageTabernaemontanus (1590) illustration of Oxalis ovina.  

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 338) ...

Original Text
Acetofella. 22. RUMEX floribus dioicis , foliis lanceolato-haftatis. Vir. cliff. 32. Hort. cliff. 139. Fl. lapp. 131. Fl. fuec. 296. Roy. lugdb. 231. Gron. virg. 153.
  Acetofa afvenfis lanceolata. Bauh. pin. 114.
  Oxalis ovina. Tabern. ic. 440.
  β. Acetofa lanceolata anguftifolia repens. Bauh. pin. 114. prodr. 55.
  γ. Acetofa arvenfis minima non lanceolata. Bauh. pin. 114.
  δ. Acetofa minor erecta, lobis multifidis. Bocc. muf. 164. t. 126.
  Habitat in Europae pafcuis & arvis arenofis. ♃.

 

Other articles:
• Golden Checklist Flora:  Rumex crispus;  

Rumex crispus L. “Curley Dock”

 

 

Other articles:
• Golden Checklist Flora:  Rumex salicifolius;  

Rumex salicifolius Weinm. “Willow Dock”

“The name R. salicifolius has been applied in a broad sense to nearly all species of subsect. Salicifolii, including even mostly Asian R. sibiricus. Rumex salicifolius appears to be most closely related to R. californicus and R. utahensis (Mosyakin, FNANM, v. 5)”.

 

Other articles:
• Golden Checklist Flora:  Rumex triangulivalvis;  

Rumex triangulivalvis (Danser) Rech. f. “Triangular-Valved Dock”

(Syn: R. salicifolius Weinm. var. triangulivalvis (Danser) J. C. Hickman )

 

Other articles:
• Golden Checklist Flora:  Rumex venosus;  

Rumex venosus Pursh “Veiny Dock”

 

Literature Cited:
- Pursh, Frederick, 1814.  

Descibed by Pursh (1814, v. 2, supplement, p. 733) from a collection by Bradbury in Upper Louisiana.

 

 

Other articles:
• Golden Checklist Flora:  Chenopodiaceae;  

Chenopodiaceae Ventenat

 

Literature Cited:
- Stevens, P. F., 2001 onwards.  

Given the obvious general relationships between the old Chenopodiaceae and Amaranthaceae, and the complexity of the details of the relationships between them, a classification of Amaranthaceae s.l., albeit incomplete, is suggested above, however, what the future holds in terms of ideas of relationships will determine its fate... `

 

Other articles:
• Golden Checklist Flora:  Atriplex canescens;  

Atriplex canescens (Pursh) Nutt. “Four-wing Saltbush”

 

Literature Cited:
- Pursh, Frederick, 1816.  

Pursh (1814, v. 2, p. 370) published the name as Calligonum canescens from a Lewis & Clark collection at the Big Bend of the Missouri.

Original Text Comments
  441. CALLIGONUM. Gen. pl. 834.
canescens. 1. C. dioicum, pulverulento-tomentosum ; foliis lanceolatis, floribus axillaribus glomeratis in apice ramulorum, subspicatus, frictibus alatis, alis venosis cristato-dentatis.
  In the plains of the Missouri, near the Big-bend. ♄. July, Aug. v. s. in Herb. Lewis flowers exceeding small. Goats delight to feed on this shrub.

Literature Cited:
- Nuttall, Thomas, 1818.  

Nuttall (1818, p. 197) moved it to Atriplex

Original Text Comments
283. ATRIPLEX. L. (Orache.)  
Flowers polygamous. — Calix 5-parted. Corolla none. Style bifid. Feminine flower; calix 2-parted, compressed. Seed vertical.  
Flowers glomerate, paniculate; bisexual; masculine and feminine flowers intermixed, or on separate plants; leaves alternate, rarely subopposite. Mostly annual, rarely shrubby.  
Species. 1. A. *canescens. (Calligonum canescens, Pursh, Flor. Am. Sept. 2. p. 370.) Dioicous; pulverulently furfuraceous and canescent; stem shrubby, diffuse; leaves linear-oblong, entire, obtuse, attenuated towards the base, younger leaves acute. — Obs. Stem much branched, and diffuse, about 3 or 4 feet high, with round gray branches. Leaves alternate, 15 to 20 lines long, about 3 wide, sometimes cuneate-oblong, obtuse and now and then emarginate, very entire, covered with the white branny scales common to this and the preceding genus. Flowers dioicous, with 4, 5, and 5 stamens, conglomerated towards the ends of the branches; male clusters (at least the lower ones) pedunculate. Calix of the female flowers 2-parted, becoming indurated, acute, with 4 unequal cristated or dentated angles. Style 1, deeply bifid, exserted.  
Nearly allied to A. portulacoides. Hab. On the denudated saline hills of the Missouri; commencing about 15 miles below the confluence of White river, and continuing to the mountains. Flowering in May.  

2. hortensis.
3. patula. …
4. laciniata.
5. *argentea. …
6. * arenaria. …
 
Chiefly an European genus, the above excepted, with 1 species at the Cape of Good Hope, 1 in Barbary, 1 in Siberia, 2 in Tartary, from whence A. hortensis is said to have originated, and 1 in Bengal.  

 

Other articles:
• Golden Checklist Flora:  Atriplex hortensis;  

Atriplex hortensis L. “Garden Orache”

 

   

Bassia All.

 

Literature Cited:
- Allioni, Carlo, 1766.  

The name Bassia was proposed by Allioni (1766) primariy from plants collected in Egypt.

Literature Cited:
- Kadereit, Gudrun, and Helmut Freitag, 2011.  

 

 

Other articles:
• Golden Checklist Flora:  Blitum nuttallianum;  

Blitum nuttallianum Schult. “Nuttall's Povertyweed”

 

Literature Cited:
- Nuttall, Thomas, 1818.  

Nuttall (1818, v. 1., p.4) ...

7. BLITUM. L. (Strawberry-spinage.)
Calix 3-parted. Corolla 0. Seed 1, covered by the calix, which enlarges and generally becomes a berry.
Flowers and berries in capitate clusters; the capituli resembling strawberries, and are both terminal and axillary.
Species. 1. B. capitatum. A doubtful native. 2. * Chenopodioides? Leaves almost hastate-riangular, somewhat dentate, at either end attenuate; glomeruli all axillary, leafy; seed distinct, punctate, not berried.
Hab. on arid soils near the banks of the Missouri. ☉
Obs. Stem erect, virgate; seeds naked, imricated, surrounded by a few linear leaves longer than the seed; proper calix, apparently none, style 1, deeply bifid. Seed coated, coverec with impressed punctures, oboval, slightly margined, beneath the outer envelope dark brown, a little rigose, emarginate below. Corculum curved round the perisperm, parallel with the margin of the seed; perisperm partly farinaceous and partly corneous.

Blitum chenopodioides L. was a validly published name for a plant found in Tataria, a name used for central Asia and European Russia east of the river Don. Nuttall's plant was not this, so the publication was a nom. illeg.

Literature Cited:
- Schultes, Josef A., 1822.  

Schultes (1822, v. 1, p. 65-66) ...

5. Blitum Nuttallianum; foliis subhastato-triangularibus, subdentatis, utrinque attenuatis; glomerulis omnibus axillaribus, foliosis; seminibus distinctis, punctatis, nec baccatis. Blitum chenopodioidis? Nuttall gener. P. 4.
Caules erecti, virgati; semina nuda, imbricata, foliolis aliquot linearibus circumdata, eadem excedentibus, cat. O; stylus 1, profunde bifidus. Semen obovatum, submarginatum, sub tegmine exterior fuscum, punctis impressis, subrugosum, basi emarginatum. Corculum circa perisperma farinoso - corneum curvatum. Nuttall. In aridis ad ripas Missouri. ☉.

Literature Cited:
- Greene, Edward Lee, 1891-97.  

Greene (1891-1897, p. 168) ...

3. MONOLEPIS, Schrader. Annuals, with the habit and foliage of Chenopodium, but the perianth consisting of a single scale-like or bract-like sepal (or this to be regarded as a mere bract subtending an achlamydeous flower). Stamen 1. Styles 2, filiform. Pericarp membranous, persistent upon the vertical compressed seed. Embryo annular; albumen copious. — In aspect wholly like Chenopodium, to which the genus may as well be united as Mungea to Amarantus.
1. M. Nuttalliana, Greene. Roem. & Schult. Mant. i. 65 (1822), under Blitum: M. chenopodioides, Moq. in DC. Prodr. xiii. 85 (1849). Blitum chenopodioides (?) Nutt (1818), not Linn. Branches many, decumbent or almost prostrate, 1/4-l ft. long ; herbage deep green, the growing parts mealy : leaves lanceolate-hastate, 1/2 - 1 in. long, entire or remotely sinuate-dentate, acute or obtuse, cuneate at base, the upper floral subsessile : flower-clusters axillary, dense, sometimes reddish : sepal foliaceous and fleshy, oblanceolate or spatulate, often exceeding the fruit : pericarp somewhat fleshy, becoming dry and favose-pitted, adherent : seed lenticular or reniform, 1/2 line broad. — Alkaline soils along the eastern base of the Sierra.

Literature Cited:
- Fuentes-Bazan, Susy, Pertti Uotilla, and Thomas Borsch, 2012.  

Fuentes-Bazan, Susy, Pertti Uotilla, and Thomas Borsch (2012, p. 18) …

3. Blitum nuttallianum Schult., Mant. 1: 65. 1822 ≡ Blitum chenopodioides Nutt., Gen. N. Amer. Pl. 1: 4. 1818 [non L. 1771] ≡ Monolepis chenopodioides Moq. in Candolle, Prodr. 13(2): 85. 1849, nom. illeg. ≡ M. nuttalliana (Schult.) Greene, Fl. Francisc.: 168. 1891. — Described from the banks of the Missouri river; type not designated.
= Chenopodium trifidum Trev., Ind. Sem. Hort. Bot. Vratislav. 1829 [n.v.] ≡Monolepis trifida (Trev.) Schrad., Ind. Sem. Hort. Goett: 4. 1830. — Lectotype (designated here): “C. trifidum Trev., m[isit] Trevianus” [later added:] “M. trifida Schrad.” [both, manu Ledebour] in herb. Ledebour (LE!).

 

Other articles:
• Golden Checklist Flora:  Chenopodium album;  

Chenopodium album L. “Lambsquarters”

 

 

Other articles:
• Golden Checklist Flora:  Chenopodium atrovirens;  

Chenopodium atrovirens Rydb. “Pinyon Goosefoot”

 

Literature Cited:
- Rydberg, Per Axel, 1900a.  

Rydberg (1900, p. 131) ...

Original Text
* Chenopodium atrovirens.
Stem 3-5 dm. high, branched, striate, obtusely angled ; leaves slender-petioled, broadly ovate, obtuse, truncate, or the upper mucronate, entire, sometimes slightly hastately lobed at the base, three-nerved, very dark green, only sparingly mealy, rather thick and somewhat fleshy, 1-3 cm. long and 5-15 mm. wide; flowers in small spike-like glomerules in the axils of the leaves and in compound interrupted spikes at the ends of the branches, very small and sparingly meally; seeds lenticular, 1 mm. long, almost black, smooth, easily separating from the pericarp.
It much resembles C. olidum, but differs in the darker green color of the leaves and stem, the sparser mealiness, the smaller flowers and the easily separated seeds, which are not pitted as are those of C. olidum. The type was growing on a dry hillside together with Mentzelia tenerrima and Symphoricarpos vaccinioides, under some trees of Pseudotsuga mucronata.
Montana: Foothills of Electric Peak, August 18, 1897, Rydberg & Bessey, 3948 (type).
Yellowstone Park: Stevenson Island, 1885, Tweedy, 439.


* Watson, Rev. N. A. Chenopodiaceae in Proc. Am. Acad. 9: 82-126.

 

Other articles:
• Golden Checklist Flora:  Chenopodium berlandieri;  

Chenopodium berlandieri Moq. “Pitseed Goosefoot”

 

Literature Cited:
- Moquin-Tandon, Alfred, 1840.  

Moquin-Tandon (1840, p. 23) ...

Original Text Comments
9. C. Berlandieri , caule fruticoso ascendente ramoso, foliis alternis ovatis vel ovato-oblongis aculis submucronulatis integerrimis pulverulentis glaucis, inferioribus subrliombeo-ovatis interdum rhombeis (angulis lateralibus prominulis), spicis subpaniculaiis laxis aphyllis , perigonio fruclifero non carinato , semine punclulato-rugoso haud nitido margine acuto.  
Circa Mexico. Berlandier 1906. (v. s. in herb. Moricand.) Seen in the dried state in the herbarium of Moricand.

Moricand is probably Moïse-Étienne Moricand, a Swiss botanist and malacologist (1779–1854). In 1833-1846, he published Plantes nouvelles d'Amérique, based chiefly on collections of J.L. Berlandier, J.S. Blanchet and J.A. Pavón. Moricand's herbarium, numbering more than 54,000 specimens at the end of his life, contained material from all over the world, again collected by him and those with whom he had built relationships. It was given to the Conservatoire botanique de la ville de Genève by his son in 1908.

   

Dysphania R. Br. “Wormseed”

 

Literature Cited:
- Uotila, Pertti, et al., 2021.  

… placed Cycloloma in Dysphania

 

Other articles:
• Golden Checklist Flora:  Dysphania botrys;  

Dysphania botrys (L.) Mosyakin & Clemants “Jerusalem Oak Goosefoot”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Chenopodium botrys L., Sp. Pl. 1: 219 (1753).

 

Dysphania botrys (L.) Mosyakin & Clemants, Ukrayins'k. Bot. Zhurn. 59(4): 383 (2002).

   

Kochia Roth

 
 

IPNI (2021) provides this reference, “Kochia Roth, J. Bot. (Schrader) 1800(1): 307 (1801),” which I think is this publication: https://hdl.handle.net/2027/nyp.33433011408956. Unfortunately, I am unable to find it there.

I think it is pretty clear that Bassia Allioni (1766) has priority over Kochia Roth (1801).

 

Other articles:
• Golden Checklist Flora:  Kochia scoparia;  

Kochia scoparia (L.) Schrad. “Common Red Sage”

The basionym is Chenopodium scoparia L.

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 221) ...

Original Text Interpretation and Comments
15. CHENOPODIUM foliis lineari-lanceolatis planis integerrimis. Hort. cliff. 86. Hort. upf. 55. Roy. lugdb. 220. Scoparia.  
Linaria fcoparia. Bauh. pin. 212.    
Ofyris. Dod. pempt. 151    
Habitat in Græcia, Japonia. ☉    

Literature Cited:
- Schrader, Heinrich Adolph, 1809.  

Schrader (1809, p. 85) placed our plant in Kochia Roth. There may be some discussion on the preceding pages explaining the reason. However, my German abilities are insufficient to find it.

Literature Cited:
- Kadereit, Gudrun, and Helmut Freitag, 2011.  

Excerpts that relate to Kochia scoparia:

Original Text Comments
Morphological studies … frame of the study did not allow the full study of critical groups, such as Kochia scoparia s.l. and K. prostrata s.l., in detail to solve their inherent taxonomic problems.  
Bassia and Kochia are highly polyphyletic, with the former distributed among all clades, and the latter in most of them.  
Scott (1978) already reduced the number of non-Australian genera of Camphorosmeae to five by including Kochia, Londesia and Chenolea into Bassia.  
The Bassia hyssopifolia subclade contains Bassia hyssopifolia and the Kochia scoparia group, a clade consisting of four closely related and morphologically similar taxa; Fig. 5. We also treat the latter here under Bassia. The subclade consists of annual species and has a wide distribution area in Eurasia which ranges from the Iberian Peninsula to Japan.  
The Kochia scoparia group is well-known for its large variation in shape, size and indumentum of leaves, indumentum of leaf axils (very dense and long in K. densiflora), size and shape of wings on the fruiting perianth etc. which were treated in various ways by different authors. Our sampling included some conspicuous forms but was not large enough to resolve the taxonomy of the group. For morphological and ecological reasons, most likely only K. scoparia, K. indica and K. littorea deserve species rank.  
Bassia scoparia (L.) A.J. Scott in Feddes Repert. 89: 108. 1978 = Chenopodium scoparium L., Sp. Pl.: 221. 1753 (‘scoparia’) = Kochia scoparia (L.) Schrad. in Neues J. Bot. 3(4): 85. 1809.
= K. sieversiana (Pall.) C.A. Mey. in Ledebour, Fl. Altaic. 1: 415. 1829.
= K. scoparia var. densiflora Moq. in Candolle, Prodr. 13(2): 131. 1849.
= K. densiflora auct.
= K. alata Bates in Amer. Bot. (Binghamton) 24: 52. 1918.

Further studies are required to elucidate the infraspecific grouping.

 

I should look up this reference:

Scott, A.J. 1978. A revision of the Camphorosmoideae (Chenopodiaceae). Feddes Repert. 89: 101–119.

 

Other articles:
• Golden Checklist Flora:  Salsola tragus;  

Salsola tragus L. “Tumbleweed”

(Syn: S. australis R. Brown, S. iberica (Sennen & Pau) Botschantzev ex Czerepanov, S. kali, S. kali var. tenuifolia Tausch., S. pestifer Nels.)

 

Other articles:
• Golden Checklist Flora:  Suckleya suckleyana;  Torrey & Gray, 1855. Pac. R. R. Rep.;  Turland, et al., 2018;  

Suckleya suckleyana (Torr.) Rydb. “Poison Suckleya”

 

Literature Cited:
- Gray, Asa, 1860.

Locations: Milk River.
Full Size ImageIllustration of Suckleya suckleyana.  

Torrey in Gray (1860, v. 12, Bk. 2, pt. 2, p. 47) … the "Pacific Railroad Reports" through Washington Territory.

Original Text
OBIONE SUCKLEYANA, Torr., n. sp. (Plate IV.) "Annual, stem branching, prostrate; leaves suborbicular on long petioles, acutely repand-dentate, pale-green both sides, nearly glabrous ; glomerules axillary, monoecious bracts of the sessile fruit deltoid, united to the summit, the margin narrowly winged, crenate-denticulate. Very distinct from every other North American species of Obione, but having some resemblance to 0. argentea. It is remarkable for the roundish leaves, very long petioles, and the large and much compressed nearly glabrous fruit. The male flowers were tetramerous." — TORREY. This was collected in the Milk River valley, August 19.

Literature Cited:
- Gray, Asa, 1876b.  

Gray, Asa, 1876, pp. 101-103 described Suckleya as a taxon that fits between Atriplex and Grayia. In that process, Gray proposed a new name — Suckleya petiolaris — and placed Obione Suckleyana Torr. in synonomy. Now, of course, we would describe Gray's specific epithet as superfluous. Not sure what Gray was thinking, perhaps that Suckleya suckleyana was an redundant name. Certainly, he knew about priority of names, as the first “code” was published in 1867.

V.
MISCELLANEOUS BOTANICAL CONTRIBUTIONS.
By Asa Gray.
Presented, Oct. 12, 1875.
The following notes and characters relate mainly to Californian botany, the writer having been engaged in the preparation of the Gamopetalae for Professor Brewer's Botany of California, now printing. Some of the observations are such as could not well be recorded in that work ; and the characters of certain new genera and species may appropriately be introduced to the botanical world in a continuation of the " Contributions " which have from time to time been communicated to the Academy, and published in its Proceedings. My first note has reference to two plants of the Atlantic United States, which have long been confounded.
Grayia Brandegei. Inermis, sesquipedalis, leviter fuifuraceo-cinerea ; foliis spathulato-linearibus ; thecis minoribus flavidulis oblato-orbiculatis quandoque trialatis basi latissime retusis, alis subundulatis ; ovario basilari papuloso. — Hillsides, among fragments of cretaceous sandstone, on the San Juan River, near the boundary between Colorado and Utah, T. S. Brandegee in Hayden's Exploration, August, 1875. — While pleased with an accession to this genus, and with the opportunity of associating it with the name of an excellent correspondent who discovered it, I must add that it does not much strengthen the genus. The small thecae, as far as seen only 3 lines broad, and with some furfuraceous puberulence (but they are far from mature, and mainly unfertilized), and the papulose cellular ovary too much remind us of Atriplex (incl. Obione). A. Endolepis of Watson, Rev. Chenop., p. 111 (of which I should like to form a distinct section), has as thin and complete a sac; but there are two minute teeth at its apex, and their position, along with the venation, shows that the sac is compressed laterally, i.e., formed of two flat bracts. I agree with Mr. Watson's view, that the sac of Grayia is obcompressed, or formed of a pair of conduplicate bracts, completely united to the very tip ; and on this character (along with the inferior radicle) the genus actually rests. But this view demands the separation from Atriplex of a species which has always appeared like a stranger in the genus, and which I propose to establish by itself, between Atriplex and Grayia, under the name of its discoverer, Dr. George Suckley, U. S. A., one of the naturalists of the exploration across the Continent under Governor Stevens.*

* Subtribus EUROTIEAE. Theca, e bracteis pl. m. conduplicatis coalitis con stans, obcompressa, rarissime triptera.
1. Grayia. Theca nuda, integerrima, scariosa, orbiculata, plana, samaroidea, alato-marginata. Radicula infera. Flores dioici.
2. Suckleya. Theca nuda, subhastata, complanata, marginibus herbaceo-cristatis, apice bidentato. Radicula supera. Flores monoici. — S. petiolaris. Obione Suckleyana Torr. Atriplex Suckleyana Watson, l. c.
3. Eurotia. Theca villosissima, turgida, nec marginata nec aristata, apice bifida. Radicula infera. Flores dioici.
4. Ceratocarpus. Theca cuneata, obcompresso-plana, biaristata. Radicula infera. Flores monoici.

Literature Cited:
- Rydberg, Per Axel, 1900a.  

Rydberg (1900, p. 133) corrected Gray's (1876) error in Catalogue of the flora of Montana and the Yellowstone National Park without comment, placing Gray's superfluous name in synonomy along with Torrey's Obione suckleyana.

Original Text
Original Text
Suckleya Suckleyana (Torr.) ; Obione Suckleyana Torr. Pac. R. R. Rep. 12: 47, 1860; Suckleya petiolaris Gray, Proc. Am. Acad. 11: 103, 1876 [Man. R. M. 311].
Montana: Milk River, 1853 (Stevenson Exped.), Suckley.

 

 

Other articles:
• Golden Checklist Flora:  Amaranthaceae;  

Amaranthaceae Juss.

 

Literature Cited:
- Morales-Briones, Diego F., et al., 2021.  

Despite the strong signal of gene tree discordance, both nuclear and plastid data sets strongly supported five major clades within Amaranthaceae s.l.: Amaranthaceae s.s., “Chenopods I,” “Chenopods II,” Betoideae, and Polycnemoideae … These five clades are congruent with morphology and previous taxonomic treatments of the group.

Chenopods I includes Atriplex, Grayia, Chenopodium, Dysphania, and Krascheninnikovia.

Chenopods II includes Suaeda, Salicornia, Allenrolfia, Salsola, and Kochia.

 

Other articles:
• Golden Checklist Flora:  Amaranthus albus;  

Amaranthus albus L. “Prostrate Pigweed”

 

 

Other articles:
• Golden Checklist Flora:  Amaranthus arenicola;  

Amaranthus arenicola I. M. Johnst. “Sandhill Pigweed”

 

Literature Cited:
- Johnston, Ivan M., 1948.  

Original Text
Amaranthus arenicola sp. nov.

Herba erecta dioica annua: ... [... Latin diagnosis omitted ...]


KANSAS: sandhills, Hamilton County, 1895, Hitchcock 428A (type, Gray Herb.) ; sandy soil, southwestern Kansas, Aug. 3, 1895, Hitchcock 609.
COLORADO: South Fork of the Platte, 1856, H. Engelmann; indefinite, 1861, Parry 323; Ft. Lupton, Weld. Co., 1914, Johnston 275; Gilcrest, Weld Co., 1916, Johnston 275A.
OKLAHOMA: bottom of Cimarron River, Cimarron Co., 1936, Demaree 13306.
TEXAS: Limpia Canyon, 8.7 miles northeast of Ft. Davis, 1942. Cory 40520.
INDEFINITE: Powell Colorado Exploring Exped., 1868, no. 509 ; Upper Missouri River, Hayden; sandhills, 1862, E. Hall.
A species most closely related to A. myrianthus Standley of northeastern Mexico and southern Texas. The latter is a loosely and much branched plant with usually sprawling branches and has the bracts subtending the female flowers broad, cuspidate, and distinctly shorter than the perianth lobes. The plant here described grows in sandy places on the high plains. It has passed as A. Torreyi Gray and is treated under that name in Standley's revision of the genus, No. Am. Fl. 21: 107 (1917). The name "A. Torreyi Gray," however, properly belongs to a very different plant growing along the Mexican boundary. Cf. Johnston, Jour. Arnold Arb. 25: 155 (1944).

 

Other articles:
• Golden Checklist Flora:  Amaranthus blitoides;  

Amaranthus blitoides S. Watson “Mat Amaranth”

 

Literature Cited:
- Watson, Sereno, 1871.
- Watson, Sereno, 1877.

Other articles:
• Golden Checklist Flora:  Watson, 1871, 40th Parallel, publication details;  

Sereno Watson (1879, v. 12, new ser. v. 4, p. 274) proposed Amarantus blitoides.

Original Text
Amarantus (Pyxidium) blitoides. Prostrate or decumbent, the slender stems becoming a foot or two long, glabrous or nearly so : leaves broadly spatulate to narrowly oblanceolate, attenuate to a slender petiole, an inch long or usually less : flowers in small contracted axillary spikelets : bracts nearly equal, ovate-oblong, shortly acuminate, 1 to 1½ lines long, little exceeding the oblong obtuse and mucronulate or acute sepals : utricle not rugose, slightly longer than the sepals : seed nearly a line broad. — Frequent in the valleys and plains of the interior, from Mexico to N. Nevada and Iowa, and becoming introduced in some of the Northern States eastward. It somewhat resembles the A. Blitum, Linn., of the Old World, and has been mistaken for it ; but that species is usually erect, with shorter and more scarious bracts, and a smaller seed more notched at the hilum. The allied A. albus, Linn., also common and indigenous throughout the interior, is distinguished by its usually erect diffusely branched habit … [… Description of A. albus omitted …]
Of the collections currently determined A. blitoides with data available online, the oldest are:
  • F. J. Lindheimer #513, 1846, Texas, GH1928850, A. graecezans L
  • J. M. Bigelow, s.n., 1853, NY3363450, Texas, Canadian River, Fort Smith to the Rio Grande, Amaranthus albus -- Whipple 35th parallel.
  • F. V. Hayden, s.n., 1858, NY3363419, Nebraska
  • I. Burk, s.n., 1864, PH20866791, Pennsylvania, Philadelphia, Grenwich Point.
  • S. Watson, 1004, July 1868, NY2281426, Utah (Nevada), Diamond Valley.
  • Pomell, s.n., 1868, SJNM2064, Nebraska, Douglas County, Omaha.
  • George Vasey, 1868, SJNM, Nebraska, Douglas County, Omaha.

I think Watson's Coll. No. 1004 is interesting. When Watson collected and first reported it (Watson, 1871, p. 297), he applies Amaranthus alba L., while noting it is a prostrate form.

Original Text
Amarantus albus, L. Reported from the Upper Missouri, Northern Texas, and Menzies Island in the Columbia River. Truckee River bottom, and roadsides in the low valleys of Nevada and Utah. Midvein of the leaf terminating as usual in a short awn ; erect, the lower branches ascending ; ½-2 high. (1,003.) With it was also found a wholly prostrate form, the stems 1-2° long; leaves obovate or nearly orbicular. (1,004.)

In 1877, Watson will propose A. blitoides for a prostrate form of Amaranthus that is otherwise similar to A. albus. While Watson labeled his collection A. albus, it has since been annotated A. blitoides. Watson's Coll. No. 1003, to which he initially applied A. albus has since been annotated A. californicus.

 

Other articles:
• Golden Checklist Flora:  Amaranthus powellii;  

Amaranthus powellii S. Watson “Powell's Pigweed”

The earliest collection of this pigweed would appear to be Fendler #735, which was in 1894 designated the type of A. bracteosus Uline & W. L. Bray, who thought that Fendler's collection was different from the plants grown from Powell's seed. However, Watson's (1875) A. powellii has priority.

Literature Cited:
- Watson, Sereno, 1875.  

Watson (1875, v. 10, p. 347) published Amarantus powellii from plants that were garden-grown from seeds brought from Arizona by Col. Powell.

Original Text Interpretation and Comments
Amarantus Powellii. Slender, 4-5 feet high, glabrous, the stem becoming bright red, branches erect ; leaves small, oblong-lanceolate, 2 inches long, cuneate at base, the slender petiole shorter ; flowers in close narrow compound spikes, the lateral ones erect ; seed nearly black, shining, less than half a line broad, lenticular, very minutely and slightly tuberculate. — Likewise cultivated from seeds brought from Arizona by Col. Powell, and apparently a very distinct species. A. albus and retroflexus were also raised from the same collection of Arizona seeds, used as food by the Indians. The only other species known from our western territory and apparently indigenous are A. chlorostachys and Blitum. The other seeds were A. leucocarpus that is now treated as a synonym of A. hypochondriacus L.

 

Other articles:
• Golden Checklist Flora:  Froelichia gracilis;  

Froelichia gracilis (Hook.) Moq. “Slender Snakecotton”

 

Literature Cited:
- Hooker, William Jackson, Sir, 1837-1841.  

Hooker (1840, v. 3., Tab. CCLVI) notes a distinct Oplotheca collected by Drummond in Texas. He also mentions a genus name of Froelichia of Moench.

Original Text
… and a very distinct one also exists in Mr Drummond's 2d Coll. from Texas, n. 244, which may be thus characterized :—
Oplotheca gracilis ; pubescenti-sericea, caulibus gracillimis basi decumbentibus geniculatis dein erectis, foliis anguste lineari-lanceolatis, spiculis parvis paucifloris remotis, perianthio tomentoso, fructifero cristis brevibus crassis profunde dentatis.
Hab. Texas, Drummond.

Literature Cited:
- DeCandolle, Agustin Pyramus, 1849.  

DeCandolle (1849, v. 13, pt. 2, p. 419) accepted Froelichia Moench and placed Oplotheca Nutt. in synonomy.

Original Text Interpretation and Comments.
XLIII. FROELICHIA Moench meth. 1794, p. 50, Endl gen. p. 302, n 1959, non Froehlichia Vahl (1), nec Wulf (2). — Oplotheca Nutt. gen. am. 1818. v. 2, p. 78, …  
Sectio I. Hoplotheca  
1. F. gracilis, caule tereti striato villoso ... [... Latin diagnosis omitted ...] (I) In Texas (Drummond! n. 244), Oplotheca gracilis Hook.! in herb. (v. s. in h. Hook. et Mus. vindob.) Seen in the dry state in Hooker's herbarium and the herbarium of the Natural History Museum of Vienna.

 

Other articles:
• Golden Checklist Flora:  Abronia fragrans;  

Abronia fragrans Nutt. ex Hook. “Snowball Sand Verbena”

 

Literature Cited:
- Hooker, Sir William J., 1853.

Other articles:
• Golden Checklist Flora:  Hooker, 1849-1857, publication details;  

Hooker (1853, p. 161) described this taxon from a description by Nuttall.

Original Text
2. Abronia fragrans, Nutt.

Hab. On loamy, sandy, firm banks, within the high drift-sand hills of the Lower Platte. Two feet long. Umbels large. Flowers porcelain-coloured, opening only at night, very fragrant ; growing with “Rumex venosus” and Psoralea arenaria, Ph. June n. 157.

Nuttall collected the plant on the sand hills of the Lower Platte, though he does not identify the expedition or the year. It is assumed (by me and others) that the collection was made on his trip to Oregon Territory in 1834. Otherwise, the earliest know collection would be on Fremont's expedition to the Rocky Mountains, 1842. This voucher (NY3370444) was in Torrey's Herbarium, so it would seem that knew the plant and the name but deferred to Nuttall to publish it.

 

Other articles:
• Golden Checklist Flora:  Mirabilis linearis;  

Mirabilis linearis (Pursh) Heimerl “Narrowleaf Four O'Clock”

(Syn: Oxybaphus linearis (Pursh) B. L. Robinson )

Literature Cited:
- Nuttall, Thomas, 1813.  

Nuttall (1813, p. 2) ...

Original Text Comments
21. [Calymenia] angustifolia. ‡ M.  

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, Supplementum, p. 728) ...

Original Text Comments
p. 97 Allionia linearis. — A. caule erecto teretii, foliis linearibus subcarnosus, floribus racemosis, pedicellis aggregatis, staminibus longissime exertis.  
  Calymenia angustifolia. Fraser Catal.. 1813.  
  In Upper Louisiana. Bradbury. v. s. in Herb Bradbury.  

Literature Cited:
- Heimerl, Anton, 1901.  

Heimerl (1901, p. 186) ...

Original Text Translation and Comments
10. Mirabilis linearis (Pursh). — Das Herbarium Delessert besitzt von dieser, bekannutlich in Nordamerika sehr gewohnlichen Pflanze eine neue Form, welche von dem verbreiteten Typus durch rauhe Behaaruug (besonders der unteren Stengelstucke) abweicht und als forma subhispida bezeichnet werden soll. Sie stimmt vollig mit dem Typus im Habitus, in den sitzenden linealeu bis lineallanzettlichen, ganzrandigen oder undeutlich uod entfernt gezahnelteni Blattern, in der ziemlich reichblutigen, opponiert und decussiert verastelten, drusenlos behaarten Rispe, in den 2-3 blutigen Involucren, mit ziemlich ansehnlichen Bluten u. s. f. uberein — unterscheidet sich aber leicht folgendermassen : caulis internodia imprimis inferiora et media (paniculae regione excepta!) pilis haud densis, saepius sparsis, patentibus, asperis, usque 2 mm. lg. subhispida; folia inferiora et media iisdem pilis imprimis in margine et infra in nervo mediano parce setoso-hirta, folia in paniculae regione brevius pubescentia. Das Exemplar wurde von Earle in New-Mexico gesammelt und mit der Bezeichnung ausgegeben: N. Mex., north of El Capitan Mts., Lincoln County (Exsicc. no. 383). The Herbarium Delessert possesses of this, notably in North America ordinary plant a new form, different from the common one Type by rough hair (especially the lower stem pieces) differs and is called forma subhispida target. ... The specimen was obtained from Earle in New Mexico collected and issued with the designation: N. Mex., north of El Capitan Mts., Lincoln County (Exsicc. no. 383).

There of five vouchers of Earle's Coll. No. 383 listed on SEINet: three at NMC, and one each at NY and RM.

 

Other articles:
• Golden Checklist Flora:  Mirabilis nyctaginea;  

Mirabilis nyctaginea (Michx.) MacMill. “Heartleaf Four O'Clock”

 

Literature Cited:
- Michaux, Andre, 1803.  

Michaux (1803, v. 1, p. 100) ...

Original Text
ALLIONIA. L.
Cal. communis oblongus , simplex , triflorus ; proprius obsoletus , superus. Corollulae irregulares. Recept. nudum.
nyctaginea. A. erecta, glabriuscula : foliia latocordatis, acutis : pedunculis solitariis ; involucro quinquefido , 3 - floro : calycibus fructiferis brevissima pube hirsutulis, involucro ampliato multo minoribus.
Obs. Habitus omnino Nyctaginis hortensis (Mirabilis Jalapae. Linn.)
Hab. ad ripas fluminis Tennassee.

 

Other articles:
• Golden Checklist Flora:  Claytonia lanceolata;  

Claytonia lanceolata Pursh “Lanceleaf Springbeauty”

 

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 1, p. 175) ...

Original Text Comments and Interpretation
3. C. foliis lanceolatis : caulinis ovatis sessilibus, racemo solitario elongato, calycis foliolis brevibus obtusissimis, petalis cuneatis bifidis, radice tuberosa. — Pall. Mss. lanceolata.  
On the Rocky-mountains. M. Lewis. ♃ June. v. s. in Herb. Lewis. Flowers white, nearly the size of the first species, without veins. In the collection of A. B. Lambert, Esq. I found a specimen collected by Pallas in the eastern parts of Siberia, perfectly agreeing with the present species.   The “first species” was C. virginica.

 

Other articles:
• Golden Checklist Flora:  Claytonia rosea;  

Claytonia rosea Rydb. “Rocky Mountain Springbeauty”

 

Literature Cited:
- Rydberg, Per Axel, 1904.  

Rydberg (1904, v. 31, n. 7, p. 404) ...

Original Text
Claytonia rosea sp. nov.
Scape about 1 dm. high, slender, from a small corm 10-15 mm. diameter ; basal leaves rare, long-petioled ; blade 1-2 cm. long, spatulate ; stem-leaves linear or narrowly linear-lanceolate, sessile, 2-5 cm. long, 1-ribbed or faintly 3-ribbed, acute, rather fleshy ; sepals rounded ovate, rounded at the apex, about 5 mm. long, half as long as the pink obovate petals; inflorescence 5-10-flowered, short, little exceeding the leaves, bractlets lanceolate ; capsule shorter than the sepals ; seeds about 2 mm. long, black and very glossy.
The few specimens of this species that are found in our herbaria bear a variety of names, as C. caroliniana, C. caroliniana lanceolata, C. caroliniana sessilifolia and C. lanceolata sessilifolia. The original C. lanceolata Pursh is a much larger plant with broad, strongly 3-ribbed stem-leaves, elongated inflorescence and large flowers, the sepals being about 6 mm. long. The type of C. caroliniana sessilifolia Torr., is a plant somewhat resembling the present species, but with somewhat broader leaves abruptly contracted at the sessile base and with acutish sepals. Neither in the herbarium of the New York Botanical Garden nor in that of Columbia University is found any other specimen matching the original collected by Bigelow. Most species of Claytonia are found near water or in damp places. The type of C. rosea was collected on rather dry hills, at an altitude of 2200-2300 m.
Colorado : La Veta, 1900, Rydberg & Vreeland, 6300 (type) ; "Colorado," G. C. Woolson ; 1875, W. A. Henry ; Graham's Peak, 1899, C. F. Baker, 303.
Wyoming: Pole Creek, 1894, Aven Nelson, 27.

 

Other articles:
• Golden Checklist Flora:  Phemeranthus parviflorus;

Locations: North Table Mountain.
Full Size ImagePhemeranthus parviflorus on top of North Table Mountain.  

Phemeranthus parviflorus (Nutt.) Kiger. “Sunbright”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1838-1843) proposed Talinum parviflorum from a manuscript by Nuttall.

Original Text
2. TALINUM. Adans. (in part.); Sims, bot. mag. t. 1357.
Sepals 2, ovate, concave deciduous. Petals 5, sessile, hypogynous. Stamens 10-20, inserted with the petals, and often coherent with them at the base. Style trifid. Capsule subglobose, 3-valved, many-seeded.
§ Stigmas or lobes of the style short, connivent, Perennial herbs, with a short thick and firm stem, and terete subulate fleshy stems : flowers in a terminal dichotomous cyme, expanding for a single day. — Phemeranthus, Raf.
1. T. teretifolium (Pursh): ...
2. T. parviflorum (Nutt. ! mss.): “small; leaves slender; stamens 5?-10.”

On rocks, Arkansas; with the preceding species, Nuttall! — A distinct species, according to Nuttall, with muvh smaller flowers than T. teretifolium.

Literature Cited:
- Kiger, Robert W., 2001.  

Kiger (2001) described new combinations to be used in his treatment of Portulacaceae in Flora of North America.

Original Text Comments
... recent molecular data ... congruent with morphological evidence ... indicates that Phemeranthus is phylogenetically distinct from the mainly Old World Talinum ... ... morphological differences between Phemeranthus and Talinum ...
Phemeranthus parviflorus (Nuttall) Kiger. comb. nov. Basionym: Talinum parviflorum Nuttall, in J. Torrey & A. Gray, Fl. N. Amer. 1: 197. 1838. TYPE: U.S.A. Arkansas: n.d., T. Nuttall s.n. (possible isotypes or syntypes, K [in type folder, on sheet with Drummond 34], K [in general herbarium]) There follows some discussion of whether the specimen at K is a type or, if not, further search should be conducted before designating a lectotype.

 

Other articles:
• Golden Checklist Flora:  Portulaca oleracea;  

Portulaca oleracea L. “Little Hogweed”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 445) ...

Original Text
1. PORTULACA foliis cunciformis, floribus feffilibus. Roy. lugdb. 473. Gron. virg. 59. Hort. upf. 146. Mat. med. 260. Dalib. parif. 158. Hall. helv. 392. Sauv. monfp. 168. oleracea.
Portulaca foliis cuneiformibus verticillatis feffilibus, floribus feffilibus. Hort. cliff. 207.  
Portulaca anguftifolia fylveftris. Bauh. pin. 288.  
β. Portulaca latifolia fativa. Bauh. pin.288.  
Portulaca domeftica. Lob ic. 388.  
Habitat in Europa auftrali, India, Inf. Afcenfionis, America, ☉  

 

Other articles:
• Golden Checklist Flora:  Cerastium arvense strictum;  

Cerastium arvense L. ssp. strictum Gaudin “Field Chickweed”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 439) ...

Original Text
8. CERASTIUM foliis lanceolatis, pedunculis ramofis, capsulis fubrotundis. ftrictum.
Ceraftium caule perenni procumbente, foliis lanceolatis tomentofis. Roy. lugdb. 450.  
Myofotis arvenfis, polygoni folio. Vaill. parif. 141. t. 30. f. 5.  
Lychnis incana repens. Bauh. pin. 206.  
Habitat in Gallia, Italia. ♃  

 

Other articles:
• Golden Checklist Flora:  Cerastium brachypodum;  

Cerastium brachypodum (Engelmann ex A. Gray) B. L. Robinson “Short-stalked mouse-ear chickweed”

 

Literature Cited:
- Gray, Asa, 1867.  

Cerastium nutans var. brachypodum Engelm. ex A.Gray, Manual (Gray), ed. 5. 94 (1867).

Gray (1867, p. 94) ...

Original Text
3. C. nutans, Raf. Annual, very clammy-pubescent; stems erect, slender, grooved, diffusely branched (6'- 20' high); cyme loose and open, many-flowered; leaves oblong-lanceolate, acute, the lowest spatulate ; peduncles mostly elongated ; petals longer than the calyx ; pods nodding on the stalks, curved upwards, thrice the length of the calyx. — Moist places, Vermont to Minnesota and southward. May-July. — Var. brachypodum, Engelm., W. Illinois and southwestward, has pedicels shorter than the pods.
 

Cerastium brachypodum (Engelm. ex A.Gray) B.L.Rob., Mem. Torrey Bot. Club v. (1894) 150; et in Proc. Am. Acad. xxix. (1894) 277.

 

Other articles:
• Golden Checklist Flora:  Dianthus armeria;  

Dianthus armeria L. “Deptford Pink”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 410) ...

Original Text
Armeria. 3. DIANTHUS floribus aggregatis fafciculatis, fquamis calycinis lanceolatis villofis tubum aequantibus. Hort. cliff. 165. Fl. fuec. 345. It. gotl. 301. Roy. ludb. 443. Sauv. monfp. 144.
  Caryophyllus barbatus fylveftris. Bauh. pin. 208.
  Armeria fylveftris altera. Lob. ic. 448.
  Habitat in fterilibus Gotlandiae, Germaniae, Galliae, Italiae. ☉

 

Literature Cited:
- Zeise, Larry Steven, 1976.

Other articles:
• Golden Checklist Flora:  Eremogone fendleri;  

Eremogone fendleri (A. Gray) Ikonnikov “Fendler's Sandwort”

 

Literature Cited:
- Gray, Asa, 1849.

Other articles:
• US Interstate 25:   at Las Vegas;

Locations: Las Vegas.  

Gray (1849, p. 13) …

Original Text
† 57. Arenaria (Eremogone) Fendleri (sp. nov.) : caudice polycephalo; turionibus imbricato-polyphyllis ; foliis praelongis erectis setaceis planiusculis margine serrulato-scabris cauleque slmplici glabris, caulinis ochreato-connatis ; symis strictis sparsifloris et calycibus glanduloso-pubescentibus ; pedicellis gracilibus ; sepalis ovato-lanceolatis sensim acuminatis cuspidatis inferne late scariosis medio viridibus trinerviis petala obovata subaequantibus ; stjlis exsertls. — Prairies, five miles west of Las Vegas ; August. — A group of grassy-leaved species, belonging to a group not before known in the New World, and to the subdivision Chromolemmae of Fenzl. My specimen is eleven inches high, just in flower, but destitute of fruit. The leaves of the sterile radical tufts are 3 to 4 inches long; the cauline pairs (3 or 4) successively shorter. Petals white, 4 lines in length.

 

Other articles:
• Golden Checklist Flora:  Gypsophila elegans;  

Gypsophila elegans M. Bieb. “Showy Baby's Breath”

There is one collection of this species of Baby's Breath, made on Lookout Mountain. One other collection in Jefferson County was made at Rocky Flats. The few (eight) Colorado collections have been made along the base of the Front Range.

The species was first described from the region of Mount Kafbek (Kazbek) in the Caucasus Mountains, in the gravels of the Terek River that drains the north side of the mountains, and in the Alps of Kafbek (Bieberstein, M., 1808). It is frequently found in wildflower mixes.

Literature Cited:
- Bieberstein, Marschall von, Freidrich August, 1808.  

Bieberstein (1808, v. 1, p. 319) ...

Original Text Comments
789. GYPSOPHILA elegans.  
G. foliis lanceolatis fubcarnofis paniculaeque ramis dichotomis patentiffimis glabris; corolla emarginata calyce genitalibusque duplo longiore.  
Habitat in glareofis torrentis Terek, circa portas caucalicas et alpem Kafbek. Septembri hinc inde adhuc florentem legi. ☉ “She lives in the gravel valley of Terek, near the caucal gates and the Alps of Kafbek. I picked up on it in September and it is still flowering.”
Radix exilis annua. Caulis paniculatus: ramis floriferis ad angulum rectum patentibus per iteratam dichotomiam dinaricatis. Pedicelli filiformes. Flores forina, colore et magnitudine praecedentis. Affinis G. vifcofae; feci foliis utrinique aculis anguftioibus, paniculae ramis glaubris longioribus et calycibus breuioribus diftincta.  

 

Other articles:
• Golden Checklist Flora:  Holosteum umbellatum;
Full Size ImageColl. No. 2246, Holosteum umbellatum  

Holosteum umbellatum L. “Jagged Chickweed”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 88) ...

Original Text Comments and Interpretation
umbellatum. 4. HOLOSTEUM floribus umbellatis.  
  Spergula foliis oppofitis, pedunculis umbellatis. Guett. ftamp. 298. Dalib. parif. 134.  
  Holofteum caryophyllaeum arvenfe. Tabern. ic. 233.  
  Lychnis graminea hirfuta umbellifera. Morif. hift. 2. p. 546. f. 5. t. 22. f. 46.  
  Caryophyllus arvenfis umbellatus, folio glabro. Bauh. pin. 210.  
  Caryophyllus arvenfis. Bauh. hift. 3. p. 361.  
  Habitat in Germaniae, Galliae arvis.  Lives in German and French fields.

 

Other articles:
• Golden Checklist Flora:  Paronychia jamesii;  

Paronychia jamesii Torr. & A. Gray “James' Nailwort”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1838, v. 1, p. 170-171)

Original Text
4. P. Jamesii: minutely scabrous-pubescent, caespitose, much branched from the base ; leaves linear-subulate, obtuse (the uppermost mucronate) ; cymes dichotomous, few-flowered, crowded, with a central subsessile flower in each division ; sepals minutely hairy at the base, linear-oblong, obscurely 3-ribbed or even, with a very short cusp, arched at the summit within ; sterile setae as long as the filaments. — P. dichotoma ? Torr. ! in ann. lyc. New York, 2. p. 290.
β depressa: dwarf, densely dichotomous; leaves and stipules imbricated on the short branches ; flowers nearly immersed in the leaves. — P. depressa, Nutt.! mss.
Rocky Mountains, lat. 41°, Dr. James! Nuttall ! β “On the barren plains of the Rocky Mountains (lat. 41°), and on the plains of the Oregon.” Nuttall ! — Stems 4-6 inches high. Leaves on the flowering branches about half an inch long, slightly 2-sulcate, about the length of the internodes. Stipules shorter than the leaves, ovate-lanceolate, acuminate, or setose ; the point much shorter than in P. dichotoma ; the flowers smaller, fewer, and more crowded ; the cusp stouter and confluent with the arched inner portion. Calyx obpyramidal at the base. Style 2-cleft ⅓ of its length. The β. depressa has the branches crowded with leaves to the summit; the stipules are nearly the length of the leaves, and the flowers are scarcely cymose.

Literature Cited:
- Gray, Asa, 1849.

Locations: Big Sandy Creek.  

Gray (1849, p. 14) ...

Original Text
69. Paronychia Jamesii, Torr. & Gray, Fl. 1. p. 170. High prairies, in gravely soil, east of Big Sand Creek (between the crossing of the Arkansas and Bent's Fort), Sept., 1846 ; and from San Miguel, New Mexico, to Council Grove, Aug. – Sept.

Big Sand Creek may now be Big Sandy Creek, and is the same creek where the Sand Creek massacre occurred.

 

Other articles:
• Golden Checklist Flora:  Pseudostellaria jamesiana;  

Pseudostellaria jamesiana (Torr.) W. A. Weber & R. L. Hartm. “Tuber Starwort”

 

Literature Cited:
- Torrey, John G., 1828.  

Published as Stellaria jamesiana Torrey (1827) who noted the habitat was moist situations within the Rocky Mountains.

Literature Cited:
- Weber, William A., and Ronald Hartman, 1979.
Full Size ImageColl. No. 2787, Pseudostellaria jamesiana  

Weber & Hartman (1979) transferred our plant to Pseudostellaria saying,

“... realignment of Stellaria jamesiana with Pseudostellaria reinforces the emerging pattern of the Southern Roky Mountain Flora as one having a strong Asiatic element probably dating back to the Tertiary, a feature which greatly impressed Sir Joseph Hooker a century ago when he visited Colorado with Asa Gray.”

 

Other articles:
• Golden Checklist Flora:  Saponaria officinalis;  

Saponaria officinalis L. “Bouncingbet”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1., p. 408-409) ...

Original Text
officinalis. 1. SAPONARIA calycibus cylindricis, foliis ovato-lanceolatis. Hort. cliff. 165. Hort. upf. 106. Mat. med. 214. Roy. lugdb. 444. Gron. virg. 160. Gort. gelr. 245.
  Saponaria major laevis. Bauh. pin. 206.
  Saponaria vulgaris. Cam. epit. 152.
hybrina. β. Saponaria concava anglica. Bauh. pin. 206. Morif. hift. 2. p. 548. f. 5. t. 22. f. 52.
  Gentiana folio convoluto. Bauh. hift. 3. p. 521.
  Habitat in Europa media.

 

Other articles:
• Golden Checklist Flora:  Silene antirrhina;  

Silene antirrhina L. “Sleepy Catchfly”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1., p. 419) ...

Original Text Comments
17. SILENE foliis lanceolatis glabris, pedunculis trifidis, petalis emarginatism calycibus ovatis, Roy. lugdb. 447. antirrhina.
Silene corymbo dichotomo, floribus pedunculatis, ramis alternis erectis, foliis lanceolatis integerrimis. Gron. virg. 50.    
Vifcago americana noctiflora, antirrhini folio. Dill. elth. 442. t. 313. f. 403.   Leaves like Antirrhinum.
Habitat in Virginia, Carolina. ☉   Known at the time of publication from Virginia and (North and South) Carolina, now described as rangeing from Canada to Mexico (POWO, 2021).

 

Other articles:
• Golden Checklist Flora:  Silene drummondii;  

Silene drummondii Hook. “Drummond's Campion”

 

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.  

Hooker (1830, v. 1, p. 89) ...

Original Text
Sect, IV. Stachymorpha. Otth. MSS. Caulescentes, Flores spicati (vel racemosi) axillares, non (aut rarius) oppositi. Calyx 10-striatus. DC.
6. S. Drummondii ; tota pubescenti-glandulosa viscida, caulibus erectis simplicibus strictis, foliis remotis lineari-lanceolatis, racemo laxo paucifloro, pedicellis elongatis plerumque alternis, calycibus oblongo-cylindraceis erectis. — S. nicaeensis. Cham. et Schlecht. in Linnaea, v. 1. p. 41.?
Radix perennis, crassiuscula. Caules simplices, erecti, strictissimi, teretes, pedales ad sesquipedalem, ubique, ut et tota plauta, praeter corollam, pubescenti-glandulosi, viscidi, superne prsecipue. Folia lineari-lanceolata, radicalia latiora, in petiolum attenuata; caulina remota sursum sensim minora, opposita, basi connata, uninervia. Flores pauci, 3-5 in racemum laxum, raro spicatum, strictissimum dispositi. Bracteae subulatce. Pedicelli alterni, nunc, sed raro, oppositi, unciam sesquiunciam longi, simplices, nudi vel, nunc, bibracteati. Calyx oblongo-cylindraceus, pubescenti-viscidus, 5-dentatus, dentibus parvis, striis decem viridibus, demum, fructiferus, paulo latior, nunquam inflatus nee clavatus. Petala alba, parva, vix calyce longiora. Capsula cylindracea, sessilis, calycis longitudine, apice 6-dentata.
Hab. Plains of the Saskatchawan. Dr. Richardson ; Drummond. Common on the elevated, gravelly soils near Fort Vancouver, and skirting the Blue Mountains. Douglas. — I am doubtful whether this species should be referred to the division “Stachymorpha” or “Siphonomorpha” of the Prodromus. It accords in many respects with S. gallica of the former division, but it has the flower-stalks vastly more elongated, sometimes opposite, and the calyx more cylindrical; while, on the other hand, it approaches very near to the S. nicaeensis in the latter division, but is far less glutinous, and never has clavate calyces, as in my specimens of nicaeensis from the south of France. Still I suspect it may be the Californian nicaeensis of Chamisso and Schlechtendal, which those authors refer, doubtfully, to the plant of Allioni and De Candolle. From both the species now mentioned ours differs in its remarkably strict habit and small petals.

 

Other articles:
• Golden Checklist Flora:  Silene latifolia;  

Silene latifolia Poir. “Bladder Campion”

(Syn: Lychnis alba Mill. , Melandrium dioicum (L.) Cosson & Germain ) Bladder Campion

 

Silene latifolia Poir., Voy. Barbarie ii. 165. (1789).

 

Other articles:
• Golden Checklist Flora:  Stellaria media;  

Stellaria media (L.) Vill. “Common Chickweed”

(Syn: Alsine media L. )

 

Other articles:
• Golden Checklist Flora:  Actaea rubra;  

Actaea rubra (Aiton) Willd. “Red Baneberry”

 

Literature Cited:
- Aiton, William, 1789.  

Aiton (1789, v. 2, p. 221) ...

ACTAEA. Gen. pl. 644.
Cor. 4-petala. Cal. 4-phyllus. Bacca 1-loculatis. Sem. femiorbiculata.  
1. A. racemo ovato, fructibus baccatis. Sp. pl. 722. fpicata.
α baccis nigris. nigra.
Common black-berries Herb-chriftopher.  
β baccis niveis. alba.
White-berried Herb-chriftopher.  
γ baccis rubris. rubra.
Red-berried Herb-chriftopher.  
Nat. α. of Britain ; β. and γ. of North America.  
Fl. April and May. H. ♃.

Literature Cited:
- Willdenow, Carl L., 1797-1830.  

Willdenow (1799, v. 2, pt. 2, p. 1039) ...

1. ACTAEA fpicata.  
α. Actaea fpicata nigra. ...  
β. ...  
γ. Actara fpicata rubra.  
A. baccis rubris. Ait. Kew. 2. p. 221. I think this is the publication of our name by Aiton.
Houttuyn Lin. Pfl. Syft. 7. p. 181.  
Aehrentragendes Chriftophskraut. W.  
Habitat in nemoribus Europae, β. γ. Americae. ♃ (v.v.)  

Literature Cited:
- Willdenow, Carl Ludwig, 1809.  

Willdenow (1809, v. 1, p. 561) ...

★ 2. ACTAEA rubra.
A. racemo ovato, petalis stamininus brevioribus, fructibus baccatis.
Actaea spicata γ. rubra. Sp. pl. ed. W. 2. p. 1139.
Habitat in Canada. ♃. D.
Baccae rubrae et Petala staminibus breviora. Non est varietas praecedentis. Actaea baccis albis ex America boreali mihi est ignota.

 

Other articles:
• Golden Checklist Flora:  Anemone canadensis;  

Anemone canadensis L. “Canada Anemone”

(Syn: Anemonidium canadensis (L.) A. Löve & D. Löve)

 

Other articles:
• Golden Checklist Flora:  Anemone cylindrica;  

Anemone cylindrica A. Gray “Candle Anemone”

 

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.  

Hooker (1840, v. 1, p. 7) ....

9. A. multifida; pilosa, foliis ternatim divisis, segmentis cuneatis tripartitis laciniatis, laciniis linearibus acutis, involucri involucellorumque breve petiolatis conformibus, sepalis 5-8 subcoriaceis ellipticis extus sericeis. — Poir. in Encycl. Meth. Suppl. v. 1. p. 364, De Less. Ic. v. 1. t 16, De Cand. Prodr. v. 1. p. 21. — A. Hudsoniana. Herb. Banks. Rich. in Frankl. 1st Journ. ed. 2. App. p. 22.
β. caule unifloro. — A. multifida, γ. uniflora. De Cand. Prodr. v. 1. p. 21. De Less. Ic. v. 1. t. 17.
γ. flore sanguineo. — A. sanguinea. Pursh, in Herb. Lamb. — A. Hudsoniana, β. sanguinea. Rich. in Frankl 1st. Journ. ed. 2. App. p. 22.
Hab. Gulf of the St. Lawrence. Goldie. From the shores of Hudson's Bay to the western declivity of the Rocky Mountains, and from the United States to near the shores of the Arctic Sea : common. Dr. Richardson. Drummond. West side of the Rocky Mountains, near the sources of the Columbia. Douglas. — Variable in size and in the number of peduncles arising from each stem, from 1 to 3, and agreeing with the figures of De Lessert (from plants gathered in the Straits of Magellan) in every particular, except that in those the segments of the leaves are shorter in proportion to the breadth ; so that their single-flowered variety has very much the appearance of a large state of A. Baldensis. Heads of pericarps, as in that species, roundish-oval, woolly. Flower white, yellow, purple, and deep red.
The species has a very extended range, from the mouth of the St. Lawrence to the sources of the Columbia, west of the Rocky Mountains. It was found at Conception in Chili, during Capt. Beechey's Expedition, at the Straits of Magellan, and Dr. Gillies has gathered it on the Andes of Chili.

 

Other articles:
• Golden Checklist Flora:  Anemone multifida var. multifida;  

Anemone multifida Poir. var. multifida “Pacific Anemone”

 

   

Aquilegia

 

Literature Cited:
- Ray, John, 1686.
Full Size ImageRay's (1686) description of Aquilegia.  

Latin Automated Translation to English My Interpreted English
Aquilegia recens vocabulum est, quasi Aquilina ob florum mucrones aduncos ut sunt Aquiline ungues. Anglis Columbine dicitur, quia florum calcaria recurva columbarum cervicem cum capite & rostro tum figura tum colore referunt. An veteribus Graecis vel Latinis cognita, aut quo nomine dicta nondum constat. Aguilegia recent designation is, as Aquilina of flowers blades that are AQUILINE hooked claws. English Columbine called for bits of flowers and beaks and curved shape and color the dove's neck and head. Is it known to the ancient Greeks, or to the Latins, the name of which it was said, or not yet been agreed. … flower petals that are like an eagle's hooked claws …
Notae illius genericae sunt folia Umbelliferarum modo divisa, flores penduli cum pluribus corniculis recurvis, femina nigra lucida. Note that the generic Umbelliferarum newspapers are now divided, with many little flowers hanging bend, female black cloud.
Aquilegiam stellatam, quae caret illis corniculis recurvis in flore, pro specie degenere habeo. Aquileia the Star, which has no unto them, little curves that bend in the flower, here standing for species degenerates I have.

Notes:

  1. “aquiline” like an eagle
    • (of a person's nose) hooked or curved like an eagle's beak.

 

Other articles:
• Golden Checklist Flora:  Aquilegia coerulea;  

Aquilegia coerulea E. James “Colorado Blue Columbine”

 

Literature Cited:
- James, Edwin, ed., 1823.  

James (1823, v. 2, p. 15) ...

[July] 11th. [1820] From our encampment, we travelled nearly south, and, crossing a small ridge dividing the waters of the Platte from those of the Arkansa, halted to dine on a tributary of the latter. In an excursion from this place we collected a large species of columbine, somewhat resembling the common one of the gardens. It is heretofore unknown to the Flora of the United States, to which it forms a splendid acquisition. If it should appear not to have been described, it may receive the name of Aquilegia coerulea.* Our road, during the morning, lay for about twelve miles, along ...

* A. coerulea. — Leaves twice ternate; flowers terminal, remote; nectaries strait and very long. It inhabits sandy woods of pine, and spruce within the mountains, rising sometimes to the height of three feet.

 

Other articles:
• Golden Checklist Flora:  Aquilegia saximontana;  

Aquilegia saximontana Rydb. “Rocky Mountain Columbine”

 

Literature Cited:
- Gray, Asa, 1878-1895.  

Gray (1895, v. 1, pt. 1, p. 43) ...

A.* saximontana, P. A. Rydberg. Much lower, scarcely a span high : stems slender, several from a scaly rootstock, quite glabrous : leaves small, twice ternate, even the upper slender-petioled, smooth : flowers much as in the preceding, but carpels glabrous. — Rydberg in ms. A. vulgaris, var. brevistyla. Gray, Am. Jour. Sci. ser. 2, xxxiii. 242 ; Porter & Coulter, Fl. Col. 4. A. brevistyla. Coulter, Man. Rocky Mt. Reg. 10; Jones, Zoe, iv. 258. — Rocky Mountains of Colorado, first collected by Parry.

Literature Cited:
- Rydberg, Per Axel, 1896.  

Rydberg (1896, v. 3, p. 482) ...

A. saximontana Rydberg ; Gray, Syn. Fl, i, pt. 1, 43 (1895); A. vulgaris brevistyla Gray, Amer. Journ. Sci. ser. 2, xxxiii, 110, and Proc. Acad. Phila, 1863, 57 (1863), name only; Porter, Port. & Coult. Fl. Col. 4 (1874), description; A. brevistyla Coulter, Man. Rock. Mount. Reg. 10 (1885) ; Jones, Zoe, iv, 258 (1893). Pl. XIX.
Stem 1 to 2 dm. high, densely tufted, scarcely exceeding the leaves, perfectly smooth; leaves twice-ternate, all on slender petioles thin, the upper a little smaller; leaflets 8 to 15 mm. long, with long petiolules, pedicels slender, upright; sepals greenish and obtuse or blue and acute; limb of the petals yellow, longer than the blue, curved spur, and the stamens and pistils; ovary smooth; pod 1.5 to 2 cm., smooth.
Rocky Mountains of Colorado. Specimens examined : Colorado, Dr. James (labeled A. caerulea, var. ?); 1861, C. C. Parry, No. 90; 1862, Hall & Harbour, No. 23; 1869, Scoville ; Argentine Pass, 1878, M. E. Jones, No. 875 ; Gray's Peak, 1895, P. A. Rydberg and C. L. Shear.

 

Other articles:
• Golden Checklist Flora:  Clematis columbiana;  

Clematis columbiana (Nutt.) Torr. & A. Gray. “Rock Clematis”

 

Literature Cited:
- Graustein, Jeannette E., 1967.
- Nuttall, Thomas, 1834a.

Other articles:
• Golden Checklist Flora:  Nuttall, 1834a, publication details;
• US I-90:   near Warm Springs;   near St Regis;

Locations: Clark Fork. Clark Fork.  

Nuttall (1834, p. 7) describes Atragene columbiana from collections by Nathaniel Wyeth on the Flat-Head river on the return trip between the Falls of the Columbia and the first navigable waters of the Missouri.

Original Text
2. Atragene *Columbiana. Pedunculis unifloris, foliis oppositis ternatum sectis, foliolis ovatis acutis, obsolete crenulatis, sepalis ovatis acuminatis, staminibus vix suplo longioribus.
Hab. Flat-Head river. In flower by the first of March, forming an intricate mat of branches so as to appear almost like a bush. Readily distinguishable from A. verticillaris by the flowers, which are scarcely half as large and of a dull palish blue. The leaflets are also cuneate rather than cordate at base, and the lateral ones apparently always entire.
Clark's Fork receives the Flathead and Bitterroot Rivers, but Wyeth, and consequently Nuttall, called Clarks Fork and all its branches the “Flathead” River (Graustein, 1967, p. 261).

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.

Other articles:
• Golden Checklist Flora:  Torrey & A. Gray, 1838-1843, publication details;  

Torrey & A. Gray (1838) placed Nuttall's Atragene columbiana into Clematis.

Original Text Comments
§ 2. Involucre none : sepals 4 : petals several, minute. — Atragene, DC. Here, Torrey & A. Gray credit DeCandolle as the author of section Atragene
19. C. Columbiana : peduncles 1-flowered ; leaves ternate : leaflets ovate, acute, obscurely crenulate ; sepals ovate, acuminate, nearly twice the length of the stamens. — Atragene Columbiana, Nutt. in jour. acad. Philad. 7. p. 7.  
Rocky Mountains, Mr. Wyeth. March. — Flowers smaller than in C. verticillaris, pale blue. Nuttall.  

Literature Cited:
- Pringle, James S., 1993+.  

James S. Pringle (1993+) writing in Flora of North America places C. columbiana in subgroup Atragene giving Torr. & A. Gray (1838) authorship of the name.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012) treat Atragene at the rank of genus.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015) in Flora of Colorado accepts Clematis columbiana (Nutt.) Torr. & A. Gray, placing Atragene columbiana Nutt. in synonomy.

 

Other articles:
• Golden Checklist Flora:  Clematis hirsutissima;  

Clematis hirsutissima Pursh “Hairy Clematis, Sugar Bowls”

(Syn: Coriflora hirsutissima (Pursh) W. A. Weber )

Literature Cited:
- Pursh, Frederick, 1814.

Other articles:
• Golden Checklist Flora:  Notes on Anemone patens var. multifida, Nuttall, 1818;  

Pursh (1814, v. 2, p. 385) ...

Original Text Comments
9. C. erecta, simplex ; foliis hirsutissimis bipinnatifidis incisis, laciniis linearibus acutis, caulinis composite-multifidis, pedunculo terminali solitario, flore erecto campanulato, petalis 4. erectis ovatis obtusiusculis apice reflexis. hirsutissima. Pursh states that the flower is erect, as might be deduced from the Lewis & Clark specimen. However, the flower is nearly always nodding. This may have led to confusion with Nuttall's Anemone ludoviciana, for which the flowers are erect.
On the plains of Columbia river. ♃. May. v. s. in Herb. Lewis. The whole plant is covered with a close coat of long hairs like Anemone Pulsatilla, which it very much resembles in several respects. I consider all the division of Anemones with caudated seeds to belong to this genus, or at least to one separate from Anemone.   The original label states, “One of the most common plants on the plains of the Columbia — May 27th 1806,” see Moulton (1999).

 

Other articles:
• Golden Checklist Flora:  Clematis ligusticifolia;
• Kinney Run Trail:   near Eagle Ridge Drive;
Full Size ImageColl. No. 1970, Clematis ligustifolia  

Clematis ligusticifolia Nutt. “Western White Clematis”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Published in Torrey & Gray (1838-1843, v. 1, p. 9) from a Nuttall manuscript.

Original Text Comments
8. C. ligusticifolia (Nutt. ! mss.): “plant somewhat pubescent ; flowers in paniculate corymbs, dioecious ; leaves pinnate ternate ; leaflets oblong, acute, mostly somewhat lanceolate-cuneate, incisely toothed and trifid ; petals and stamens equal in length ; carpels with long plumose tails. — C. Virginiana, Hook. fl. Bor.-Am. 1. p. 1. (in part). It appears that this entire description is a quote from Nuttall's manuscript.
β. brevifolia : leaves smoother, shorter and broader.  
“Plains of the Rocky Mountains, in open and in bushy places, near streams. β in the Blue Mountains and on the borders of the Oregon. — Very similar to C. Virginiana, but the leaves are mostly 5-foliolate, and almost lucidly coriaceous ; They are also much smaller, and in the var. α much narrower and longer. The tails of the carpels are also longer and more densely plumose in C. Virginiana. Flowers white and fragrant.” Nutt.  

There are two vouchers of Nuttall collections of Clematis ligusticifolia at NY. One of them (NY233134) gives the location of “R. Mts. Lewis' River” in Nuttall's hand and was designated the holotype by Arnold Tiehm, December 1985. The Lewis River of the time is today's Snake River. The other (NY233159) is from the Torrey Herbarium and gives the location as the Columbia River. This has been designated as the holotype of C. ligusticifolia var. brevifolia Torr. & A. Gray found in the Blue Mountains and on the borders of the Oregon.

 

Other articles:
• Golden Checklist Flora:  Clematis orientalis;  

Clematis orientalis L. “Oriental Virginsbower”

 
 

Linnaeus (1753, v. 1, p. 543) ...

4. CLEMATIS foliis compofitis : foliolis incifis angulatis lobatis cuneiformibus. orientalis.
Clematis orientalis, apii folio, flore e viridi-flavefcente pofterius reflexo. Tournef. cor. 20.  
Flammula fcandens, apii folio glauco. Dill. elth. 144. t. 119. f. 145.  
Habitat in Oriente.  

 

Other articles:
• Golden Checklist Flora:  Delphinium carolinianum;
• Field Notes:   Coll. No. 1440, 15 Jun 2016;  Coll. No. 2359, 12 Jun 2020;
Full Size ImageColl. No. 2359, Delphinium carolinianum subsp. virescens
Full Size ImageColl. No. 1440, Delphinium carolinianum subsp. virescens  

Delphinium carolinianum ssp. virescens (Nutt.) R.E. Brooks. “Plains Larkspur”

 

Literature Cited:
- Walter, Thomas, 1788.

Other articles:
• Golden Checklist Flora:  Walter, 1788, Flora Caroliniana, publication details;  

Walter (1788, p. 155)

Original Text Comments
T R I G Y N I A  
220. DELPHINIUM. Cal. nullus. Petala 6. Nectarium 2-phyllum, cornutum. Silique 3.  
carolinianum 1. nectario diphyllo, quam flos longiori, labellis integris, floribus fpicatis purpureis macula lutea, petalis duobus bifidis barbatis.  

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Nuttall, 1818, publication details;  

Nuttall (1818) described the Plains Larkspur from his collection on the plains of the Missouri.

Original Text
371. DELPHINIUM. L. (Larkspur.)
Calix none. Petals 5. Lepanthium (nectary) bifid, cornutely produced behind. Siliques 3 or 1.
Herbaceous, lower leaves digitate or palmate, upper ones often undivided; flowers spiked or paniculate, blue, violaceous or yellowish.
Species. 1. D. tricorne 2. azureum 3. exaltatum. 4. consolida. Naturalized.
5. *virescens. Perennial, pubescent; lepanthium 4-leaves, shorter than the 5 calicine petals, interior laminae densely bearded; leaves 3-parted, segments linear, subtrifid, lower ones divaricate.
Hab. On the Plains of the Missouri.
Obs. Stem simple about 8 to 12 inches, pubescent; leaves upon long petioles, partly digitate or 5-parted, 10 to 15 lines wide; spike few-flowered, flowers greenish white, petalois calix, 5-leaved, leaves oblong, spur longer than the flower, nearly straight. Petals of lenanthium 4; the 2 internal ones irregularly concave, small, the 2 lateral larger, flat, and unguiculate, bearded, claw sending out a short spur at the base; capsules 3. Flowering in June.
A genus almost equally divided betwixt Siberia and the south of Europe.

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.

Other articles:
• Golden Checklist Flora:  Torrey & A. Gray, 1838-1843, publication details;  

Torrey & A. Gray (1838, v. 1, p. 32) accepted Nuttall's D. virescens.

Original Text Comments
8. D. virescens (Nutt.): pubescent ; petioles scarecely dilated at the base ; leaves 3-5-parted, the middle division mostly undivided, lateral ones 2-3-cleft ; lobes lanceolate ; raceme loose, few-flowered ; sepals oblong or lanceolate ; spur longer than the sepals, ascending ; lower petals deeply 2-cleft ; claw gibbous at the base. — Nutt. ! gen. 2. p. 14; DC. prodr. 1. p. 53.  
Plains of Missouri and Arkansas, Nuttall ! North Carolina, Schweinitz ! Georgia, Le Conte ! June. — Stem 8-12 inches high. Raceme simple. Pedicels longer than the flowers. Bracts subulate. Flowers large, yellowish or greenish white, minutely pubescent. Sepals marked with a brownish spot near the apex, much longer than the petals. Spur straight or somewhat incurved. Lower petals rather densely bearded. Ovaries 3. It is possible that the collections of D. virescens should carry the name D. carolinianum.

Torrey & A. Gray (1838, v. 1, p. 32) treat D. carolinanum Walt. as a synonym of D. azureum Michx.

 

Rydberg (1899) supports acceptance of D. virescens, although on somewhat flimsy evidence. Not sure why he did not mention the taxon as found in Colorado or Wyoming.

Literature Cited:
- Brooks, Ralph E., 1982.
- Warnock, M. J., 1981.  

Original Text Comments
Delphinium carolinianum Walt. subsp. virescens R. E. Brooks, comb. nov. Basionym: D. virescens Nutt., Gen. N. Amer. Pl. 2: 14. 1818.  
Warnock (1981) treated virescens as a synonym of D. carolinanum subsp. pernardii (Huth) Warnock. Field examinations of numerous individuals combined with mor[hological evaluations, including SEM studies of the seeds, indicate that virescens is sufficiently distinct from pernardii to warrant taxonomic recognition of virescens. Warnock's (op. cit.) concept of subspecies in this group is new, however, and one with which I agree. Subsp. virescens has mostly equally distributed cauline leaves with a few basal leaves and the upper stem and rachis are covered with basally yellow, pustualte (sic) trichomes. It occurs in the eastern Great Plains and adjacent eastern areas from North Dakota south to northeast Texas and Missouri. Subsp. pernardii usually has a distinct basal rosette of leaves with a few cauline leaves, the upper spem is canescent and sparsely pustulate hairy, and the rachis is canescent. The subspecies occurs in the western Great Plains from western Nebraska south to Texas.  

Literature Cited:
- Warnock, M. J., 1995.  

 

Literature Cited:
- Jabbour, Florian, and Susanne S. Renner, 2012.  

Jabbour and Renner (2011) studied the global phylogeny of tribe Delphineae that comprises Aconitum and Delphinium. The only studied taxon found in Golden s.l. was Delphinium nuttallianum, so there is nothing to say about D. carolinianum subsp. virescens. The study showed that some re-arrangement of names in Delphineae is required though none that will affect names in Colorado. Molecular clock dating suggests that Delphinium arrived in North American from Asia in the Pliocene.

 

Other articles:
• Golden Checklist Flora:  Delphinium geyeri;  

Delphinium geyeri Greene “Geyer's Larkspur”

 

Literature Cited:
- Greene, Edward L., 1894b.  

Greene (1894, v. 2, p. 189-190) ...

Delphinium Geyeri. Rootl woody-fibrous : stem stoutish, 10 to 20 inches high: leaves mostly near the base of the stem and forming a considerable tuft, but in taller specimens many cauline also, these diminishing upwards; the whole plant, especially the calyx, pedicels, and upper part of stem, canescently tomentulose: leaves slightly fleshy, cut into many linear segments, each of these tipped with a white callosity : flowers rather large, very deep azure, in a narrow simple and strict raceme; spur stoutish, horizontal or ascending, curving downwards from near the tip.
Common plant of the very high plains toward the head-waters of the Platte River in Wyoming and northern Colorado. First collected by Nuttall, whose specimen (Herb. Brit. Mus.) is ticketed "D. hicolor? collected in Rocky Mts. by myself," though it is far enough from being his D. bicolor. It is also Geyer's n. 163 of “Slopes of undulating plains between the Kansas and Platte rivers, with Oenothera serrulata.” It is one of several easily distinguishable plants, from widely sundered, and very different climatic regions, which Dr. Gray referred to D. azureum.

 

Other articles:
• Golden Checklist Flora:  Delphinium nuttallianum;

Locations: Schweich Hill.  

Delphinium nuttallianum Pritz. “Twolobe Larkspur”

One our two common larkspurs in Golden s.l., Delphinium nuttallianum Pritz. “Twolobe Larkspur” in every open space except Schweich Hill. Found mostly along the base of the Front Range in Jefferson County, with a few collections up into the foothills. Colorado state collections are from the Front Range to the west, sparingly in the large valleys and parks, such as San Luis Valley and South Park.

The first name applied was D. pauciflorum Nutt. From a manuscript by Nuttall in Torrey & Gray (1838-1843). However, this name was previously used for a larkspur in Nepal and therefore nom. Illeg. This was corrected by Pritzel in a manuscript published by Walpers (1843).

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1838-1840, v. 1, p. 33) ...

11. D. Pauciflorum (Nutt.l mss.) : “somewhat hirsutely pilose ; petioles scarcely dilated ; leaves reniform, lobes bifid or trifid, linear and entire; bracts simple, minute ; raceme 3-5-flowered ; spur subulate, straight, about the length of the oblong acutish petals ; stigmas and styles smooth; root grumous. “petals” has been crossed out, and “sepals” written in the margin.
“Rocky Mountains and Blue Mountains of the Oregon. — Scarcely a foot high, slender ; the lower part and the stem more or less minutely and roughly pubescent. Leaves nearly smooth on the upper surface ; two or three divided ones on the stem, the uppermost beneath the flowers simple. Flowers 2-3, large, blue. Lower petals with a central line of pubescence ; upper ones hirsute externally. Carpels pubescent.” Nutt.  

Literature Cited:
- Walpers, Wilhelm Gerhard, 1842-1847.  

Walpers (1843, t. 2. p. 744) ...

13. *D. PAUCIFLORUM Nutt. mss. in Torr. et Gr. Fl. of North Am. I. p. 33. — (Wlprs. Rep. 1. p. 56.) accipiat nomen : D. Nuttallianum Pritz. mss.

 

Other articles:
• Golden Checklist Flora:  Myosurus minimus;  

Myosurus minimus L. “Tiny Mousetail”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Myosurus minimus was first published by Linnaeus (1753), and some other previous works. It's still an accepted name.

Original Text
POLYGYNIA.
MYOSURUS
minimus. 1. MYOSURUS.
Myofurus foliis integerrimis. Hort. Cliff. 177. Fl. Fuet. 261. Roy. Lugdb. 492.
Holofteo affinis Cauda mirus. Bauh. Pin. 190.
Cauda muris. Dod. Pempt 112.
Habitat in Europae collibus apricis aridis. ☉

 

Other articles:
• Golden Checklist Flora:  Pulsatilla nuttalliana;  

Pulsatilla nuttalliana (DC.) Spreng. “Nuttall's Pasque Flower”

 

Literature Cited:
- DeCandolle, Augustin Pyramus de, 1818.  

DeCandolle (1818, v. 1, p. 193) ...

7. Anemone Nuttalliana.
A. foliis palmati-sectis , segmentis tribus multifidis , lobis linearibus elongatis , involucralibus in lobos lineares multi-sectis, flore erecto, sepalis 6 erecto-conniventibus.
Hab. in Louisiana. Nuttall. ♃ ( v. s. sp. in H. Lamb.).
Species distinctissima ob habitum oninino Puisatiiiae et folia radicalia palmatim trisecta , nec pinnatim multisecta ; petioli adpresse pubescentes , 3 poll. longi; limbus trisectus, segmentis petiolulatis multifidis, lobulis linearibus , elongatis , acutis , minime dentatis , superne et imprimis subtus subvillosis ; scapus 6-12 poll. longus, plus minusve villosus , interdiim glabratus ; involucri folia in lobos lineares multisecta, basi hirsuto-villosa ; pedicellus iongitudine varins; flos purpurascens erectus , sepalis 6,oblongis, acutis, erecto-conniventibus , extus villosis, 8-10 lin. longis ; fructus A. pulsatillae !

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Notes on Clematis hirsutissima, Pursh, 1814;  

Nuttall (1818, v. 2, p. 20) ...

1. A[nemone] * ludoviciana.
Hab. Commencing near the confluence of the river Platte and Missouri; on gravelly hills; flowering about April. Clematis hirsutissima. Pursh 2. p. 385. Obs. A species related to A. patens and A. Pulsatilla, and much about the size of the latter. ...

While Nuttall was describing Pulsatilla nuttalliana, he placed Pursh's Clematis hirsutissima in synonomy, an error, thus rendering his name of A. ludoviciana illegitimate.

Literature Cited:
- Berchtold, Bedrich (Friedrich) Všemír (Wssemjr) von, and Jan Svatopluk (Swatopluk) Presl, 1823.
Full Size ImageBrechtold & Presl (1823) publication of Pulsatilla nuttalliana.  

Bercht. & J.Presl (1823) published Pulsatilla nuttalliana without much comment. They did, however, establish a specific epithet for our plant as a North American entity.

Literature Cited:
- Nuttall, Thomas, 1825.  

Nuttall (1825, v. 5, p. 158) ...

Observations on a species of Anemone, of the section Pulsatilla, indigenous to the United States. By Thomas Nuttall. Read October 11, 1825.
That the genus Anemone, as constituted by Linnaeus and his successors, even after the judicious separation of Hepatica by Willdenow, still requires revision, must, I conceive, be apparent to all who have ever compared the Pulsatillas with such other assumed species of the genus, as A. virginiana, A. caroliniana, or A. thalictroides : the last, indeed, with strict propriety, has been referred by De Candolle to Thalictrum, and the Pulsatillas are sectioually divided by Persoon, from his Anemonoides, being distinguished remarkably from the latter by the long plumose appendages of their spreading carpelles, in which particular, though remote in habit, they bear so marked an affinity to the genus Clematis, that Pursh, deceived by imperfect specimens of our plant, named it Clematis hirsutissima, (2. p. 385) and by this name he marked the specimen in Mr. Lambert's herbarium which I had collected. In the work on the Genera of North American plants and the rarer species, Vol. II. p. 20, I have attempted to describe it as Anemone *ludoviciana, having a 1 -flowered involucrate scape, remarkably soft and lanuginous (no ways hirsute) with the segments subulately divided ; the leaves pseudo-digitate, multifid, and smooth on the upper surface, the segments entire, linear and acute; the petals 6, oblong-ovate, and nearly erect.
This species has also been introduced into the Systema Vegetabilium of De Candolle, by the name of Anemone Nuttallii ; and the better to illustrate it, I have now added a figure from a sketch made on the spot (vid. Pl. viii.) Like most of the section to which it appertains, it grows on exposed bare hills, and flowers early in the month of April. The colour of the flower is a grayish sombre blue. We found it growing in considerable tufts, on hills near the confluence of the Missouri and Platte rivers. It would be highly acceptable to the flower garden from its early and showy appearance.

Unfortunately, Plate viii is missing from the online volume on Biodiversity Heritage Library.

 

Hooker (1840, v. 1, p. 4-5) ...

1. A. patens; molliter sericea, foliis serotinis tripartitis ternatisve, segmentis cuneatis trifidis incisis, lobis lineari-lanceolatis, involucro lineari-multifido, flore erecto, sepalis 5-6, aristis longis barbatis. — Linn. Sp. Pl. p. 759. Gmel. Siber. v. 4. p. 195. De Cand. Prodr. v. 1. p. 16. — (β. ochroleuca.) Sims, in Bot. Mag. t. 1994. — Clematis hirsutissima. Pursh, Fl. Am. v. 2. p. 285. — Anemone ludoviciana. Nutt. N. Am. Fl. v. 2. p. 26. — A. Nuttalliana. De Cand. Prodr. v. 1. p. 17. Nutt. in Journ. of Acad. of Nat. Sc. of Philad. v. 5, p, 158. t 8. (A. Nuttallii,) excellent. Rich. in Frankl. 1st Journ. ed. 2. App. p. 21.
Hab. Profusely in the eastern prairie district ; and, more scattered, in the central limestone tracts from lat. 45° to 67° on the Mackenzie. Dr. Richardson. Valleys in the Rocky Mountains. Drummond. Douglas. — There is no difference whatever between this American plant and the A.patens which I possess from the Russian Empire, gathered by Dr. Goldbach, and from Silesia on the borders of Poland, by Professor Treviranus. Both are liable to vary in the breadth of the segments of their leaves, and in the colour of their flowers. Mostly, however, these are purple. The pale yellow-flowered variety from Siberia, is cultivated in England; and the fresh juice of that kind, Gmeliu tells us, is employed by the inhabitants of Irkutsck to cure deafness, and newly gathered leaves for pain in the head, as vesicatories. The plant affects sandy soils, and its blossoms appear among the earliest of the season. The young buds are eaten by the Marmots, inhabitants of the plains of N. America. — A. Halleri comes too near to this species. Its flowers I have often gathered in Switzerland before the leaves have appeared; but since, in both, the flowers continue a long time, they and the leaves are sometimes gathered in perfection together, I possess specimens from Moscow, in which the former year's leaves are present with the vernal flower ; quite dead, indeed, and brown, but perfect in form and shape.

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1840, v. 1, p. 11) ...

§ 1. Carpels with long bearded tails : leaves of the involucre sessile, palmately divided, with linear lobes. — Pulsatilla, DC.
1. A. patens (Linn.): silky-villous ; leaves 3-parted or ternate; segments cuneiform, 3-cleft, incised; lobes linear-lanceolate; involucre linearly many-cleft; sepals 5-6.— DC. prodr. 1. p. 16. – (β. ochroleuca); Hook ! f.. Bor.-Am. 1. p. 4. A. Ludoviciana, Nutt. ! gen. 2. p. 26. A. Nuttalliana, DC. prodr. l. c. p. 17; Nutt. in jour. acad. Philad. 5. p. 158. t. 8. and 7. p. 7 ; Richards. ! app. Frankl. journ. (ed. 2.) p. 21. Clematis hirsutissima, Pursh, fl. 2. p. 385.
British America as far north as lat., 67° ! Valleys of the Rocky Mountains, Drummond, Nuttall! On the Missouri and Platte, Nuttall! Galena, lllinois ! — About a span high. Sepals an inch or more in length, dull blue or purple. Tail of the carpels nearly two inches long. — Appears to be identical with the European plant.

Literature Cited:
- Pritzel, Georg A., 1842.  

Anemone patens var. multifida Pritz., Linnaea 15(6): 581 (1842). Georg August Pritzel (2 September 1815, Carolath – 14 June 1874) was a German librarian and botanical writer. He studied in Breslau, graduating with a dissertation titled Anemonarum revisio. In 1851 he began work as a Hülfsarbeiter at the royal library in Berlin, a post which eventually led to curator duties. From 1855 onwards, he served as archivist at the Prussian Academy of Sciences.

Pritzel (1842, v. 15, n. 6, p. 581) …

A. patens L. sp. 759. … Hook. Fl. Bor. Am. I. p. 4 Torrey & Gray Fl . Of North-Am. I. p. 11. …  
Anemone Nuttalliana DC. Syst. I. 193. Prodr. I. 17. Pulsatilla Nuttalliana Spr. Syst. 11. p. 663. (Clematis hirsutissima Pursh, Anemone Ludoviciana Nutt. Gen. am. Et Journ. Of the Academy of nat. sc. Of Philadelphia vol. V. (a. 1825.) p. 158. et icon tab. 8. ex auctoritate Hookeri, Gray et Torrey! (Hab. as confl. Platt et Missouri, 39° lat. — secundrum DC., quod tacent Illustr. Viri Hooker, Torrey et Gray, in Louisiana; 29°-33° lat. (?!) — ad flum. Columbia, 33° lat. Sprengel!)  
 
4. var. multifida, segmento intermedio petiolato, lateralibus subsessilibus, omnibus bi-tripartitisva ; laciniis profunde incisis: lacinulis saepe iterum incisis. (var. 3. ft. alt.) — in Sibiria frequenter. While Pritzel publishes the the variety name of multifida, he does not make a connection to the North American plants.
In America boreali copiose in planitiebus orientalibus, rarius in interioribus regionibus calcareis inter lat. 45-67° (1) ad Mackenzie flumen, Dr. Richardson! Vallibus montium rupestrium (Rocky-Mountains, Felsengebirge), Drummond et Dougas! — ad confl. Platte et Missouri, 39° lat. Nuttall! — prope Galena civitatis Illinois, Torrey et Gray! 42° 50' lat. — In Louisiana, (29-33° lat.!) DC. Teste Nuttall! (locus, scriptoribus Americanis non laudatus!). Si recta res se habet. A. patens in American a lat. 29° (?!) ad lat. 67° distributa est! —  

Literature Cited:
- Hoot, Sara B., Kyle Meyer, and John C. Manning, 2012.  

The only reference made by Hoot, et al. (2012) to A. patens is to place it in section Pulsatilla (Mill.) DC. 1817:193.

Literature Cited:
- Mosyakin, Sergei L., 2016.  

Mosyakin (2016) proposed ...

... a nomenclatural and taxonomic update to the treatment of Anemone in the Flora of North America North of Mexico (Dutton et al., 1997). The taxa placed in Anemone in that treatment should not be placed in at least four genera: Anemone sensu stricto, Hepatica, Pulsatilla, and Anemonastrum.

That explains the proposed treatment as Pulsatilla. What about adoption of the specific name of nuttalliana in preference to multifida. Mosyakin (2016) summarizes ...

Native North American plants of the Pulsatilla patens aggregate were recognized by [FNANM] as Anemone patens var. multifida Pritzel, which was described from Siberia (Pritzel 1841). The North American taxon has a very complicated synonymy and it is definitely not identical with the typical European–Western Asian P. patens sensu stricto (subsp. patens). Its correct name as a species of Pulsatilla is P. nuttalliana (DC.) Bercht. & J. Presl.

Literature Cited:
- Sramko, Gabor, Leventa Laczko, Polina A. Volkova, Richard M. Bateman, and Jelena Mlinarec, 2019.  

Sramko, et al. (2019) ...

Genus Pulsatilla Mill.
...
Subgenus Pulsatilla
....
Section Pulsatilla
Series Patentes (Aichele & Schweg.) Juz. ex Tamura
species: P. integrifolia (Miyabe & Tatew.) Vorosch., P. patens (L.) Mill. [incl. subsp. flavencens (Zucc.) Zamelis, subsp. multifida (E. Pritz.) Zamelis, subsp. nuttalliana (DC.) Grey-Wilson], P. vernalis Mill.
...

 

Other articles:
• Golden Checklist Flora:  Ranunculus acriformis;  

Ranunculus acriformis A. Gray “Sharpleaf Buttercup”

 

Literature Cited:
- Gray, Asa, 1886.  

A.Gray, Proc. Amer. Acad. Arts 21: 374 (1886) ...

R. acriformis. A foot high, strict, with pubescence in good part appressed : leaves all palmately or pedately and deeply 3-5-parted or even divided, and often again 2-3-cleft into narrower lanceolate or linear segments and lobes : petals orbicular-obovate, one fourth inch long, hardly double the length of the spreading calyx : akenes over a line long; beak of half their length. — R. acris, Hook. Fl. i. 18, partly, & Lond. Jour. Bot. vi, 66, not L. — Northern Rocky Mountains, lat. 58°, Drummond. Wyoming, Parry (distrib. as R. affinis). Wind River, Dr. Forwood, and near Cheyenne, Greene.

 

Other articles:
• Golden Checklist Flora:  Ranunculus acris;  

Ranunculus acris L. “Tall Buttercup”

 

 

Other articles:
• Golden Checklist Flora:  Ranunculus aquatilis var. diffusus;  

Ranunculus aquatilis L. var. diffusus With. “Threadleaf Crowfoot”

(Syn: Ranunculus aquatilis L. var. capillaceus (Thuill.) DC.)

Literature Cited:
- Withering, William, 1796.  

Ranunculus aquatilis var. diffusus With., Arr. Brit. Pl., ed. 3. 2: 507 (1796).
Withering, William (1741-1799)
An Arrangement of British Plants ..., Edition 3

 

Other articles:
• Golden Checklist Flora:  Ranunculus cymbalaria;  

Ranunculus cymbalaria Pursh “Alkali Buttercup”

(Syn: Halerpestes cymbalaria (Pursh) Greene , R. cymbalaria Pursh var. saximontanus Fernald )

 

Other articles:
• Golden Checklist Flora:  Ranunculus glaberrimus ellipticus;  

Ranunculus glaberrimus Hook. var. ellipticus (Greene) Greene “Sagebrush Buttercup”

 

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.

Locations: Kettle Falls (historical).  

Hooker (1829, v. 1, p. 12) ...

6. R. glaberrimus ; foliis omnibus petiolatis, radicalibus subrotundis integerrimls vel grosse tridentatis, caulinis subcuneatis trifidis, calyce patente petalis dimidio breviore, fructibus globosis. (Tab. V. A.)
Radix fasciculato-fibrosa. Tota planta hirsutie destituta. Caulis erectus, subspithamieus, superne uni-bitriflorus. Folia omnia (nisi supernum quod subsessile,) petiolata, subsucculenta : radicalia subrotunda, integra, atque oranino iutegerrima vel apice dentibus trlbus grossis obtusis : caulina cuneata, fere ad medium trifidum : segmentis lanceolatis, obtusis, integris. Pedunculus et calyx etiam glaberrimi. Sepala ovalia, concava, patentia, non reflexa, corolla duplo breviora. Petala 5, ovalia, flava. Fructus, vix maturus, globosus.
Hab. Common on the mountains around the Kettle Falls, and on the Rocky Mountains, near the limits of perpetual snow. Douglas. — This differs from all its congeners by its entirely glabrous stem, leaves, and calyx, by its entire or only 3-toothed, rounded, radical leaves, and by the petiolated, never more than trifid, cauline ones. The whole plant has a succulent appearance, and turns almost black in drying. I have seen it in no collection but that of the indefatigable Douglas.

Literature Cited:
- Greene, Edward L., 1890.  

Greene (1890, v. 2, p. 110-111) ...

R. ellipticus. R. glaberrimus, Gray, Am. Journ. Sci. 2d Ser. xxx. 241; Port. & Coult. Fl. Colo. 7; Coult. Man. 7; Brew. & Wats. Bot. Calif. i. 7 mainly; Gray, Proc. Am. Acad. xxi. 369 in part, not of Hooker. R. brevicaulis, Hook. Lond. Journ. vi. 66, not of Fl. Bor.-Am. (teste Gray, l. c.). R. alismaefolius, Gray, Am. Journ. l. c. 404, not of Geyer. Very glabrous : stems several, 2 or 3 inches. high, from a large fascicle of perennial fleshy-fibrous roots : radical leaves elliptical, entire, acutish at both ends, the petiole equalling the blade ; cauline narrower, often cleft into two or three linear divisions : petals often wanting, sometimes 1 only, or 5, large, broadly obovate or more rounded, bright yellow : achenes very numerous, plump, smooth, tipped with a short curved style and disposed in a large globose head.
Lower and middle mountain districts of Colorado, Utah and Nevada to eastern California. Very nearly allied to R. glaberrimus, though a smaller plant, and inhabiting a different climatic belt. There is a difference in the achenes ; those of the true glaberrimus having a slender beak. The order of leaf-division is, moreover, inverted in the two species. In R. ellipticus the cauline are 3-cleft while the radical are entire and even acute. In R. glaberrimus the radical ones are broad and broadly 3-lobed while the cauline are entire. The herbage of the latter invariably blackens in drying. In the former it undergoes no change of color.

Literature Cited:
- Greene, Edward Lee, 1891-97.  

Greene (1892, p. 298) in Flora Franciscana ...

8. R. glaberrimus, Hook. Fl. i. 12 t. 5. A (1829). Glabrous, flaccid but rather fleshy, 3—6 in. high: leaves all petiolate; radical rounded, 3-lobed or coarsely toothed; cauline subcuneate, trifid or entire: fi. several, large ; sepals spreading; petals 3-4 lines long, obovoid: achenes plump, smooth, puberulent, with a short curved beak, and disposed in a large globose head. Var. ellipticus. R. ellipticus, Greene, Pitt. ii. 110 (1890). Radical leaves elliptical, acute, entire: stems shorter; fl. fewer, often apetalous. — The type, a plant of the far north and east, reaches our borders on the eastern slope of the Sierra northward. The variety, a plant of different aspect, and with very dissimilar foliage, is found not far from Truckee, Mr. Sonne, where it appears as if confluent with the type.

 

Other articles:
• Golden Checklist Flora:  Ranunculus ranunculinus;  

Ranunculus ranunculinus (Nutt.) Rydb. “Tadpole Buttercup”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.

Locations: Independence Rock.  

Torrey & A. Gray (1838-1840, v. 1, p. 26) worked from Nuttall's manuscript.

7. CYRTORHYNCHA. Nutt. mss.
" Sepals 5, petaloid, narrow, spreading. Petals 5, narrow and unguiculate ; the claw nearly the length of the lamina, with a projecting scaly callosity at its summit. Stamens rather numerous: anthers rounded. Stigmas short and subulate, strongly incurved. Achenia oblong-cylindrical, somewhat conspicuously grooved (not carinated), collected into a spheroidal head. Seed suspended. — A small perennial herbaceous plant. Leaves mostly arising from a short caudex, ternate and bipinnately divided. Panicle loose and cymose. Calyx petaloid and, like the corolla, bright yellow. In the fruit it resembles Thalictrum; in the flower, both Anemone and Ranunculus."
C. ranunculina (Nutt.! mss.)
" By the sides of gravelly brooks in the eastern range of the Rocky Mountains, around the place known by the name of Independence Rock on the banks of the Sweet Water of the Platte, but not further to the westward. Flowers in June. — Caudex clothed with numerous brown vestiges of sheathing petioles. The whole plant quite smooth. Leaves somewhat coriaceous and shining ; radical ones on long petioles, the subdivisions pinnatifid ; laciniae entire or 2-3-toothed. Stem, or scape, about a span high, cymosely branched above ; bearing at the lowest division a single sessile 3-parted leaf, and at the upper divisions minute and undivided leaves. Sepals oblong-ovate, spreading but not reflexed. Petals somewhat longer than the sepals, oblong, obtuse, very conspicuously narrowed below into a long claw, (almost like the nectaries of Coptis) ; the upper part of the claw thickened bv a scale-like process. Stamens 20 or more : anthers adnate. Carpels 10-15, quite glabrous, cylindrical-oblong, grooved (as in Thalictrum). Stigma subulate, shorter than the ovary, inflexed so as to be almost concealed in the mature fruit."
 

Rydberg (1984, p. 23) ...

Ranunculus ranunculinus (Nutt.)
Cyrtorrhynca ranunculina Nutt. in Torrey & Gray's Fl. N. Am. I., 26. (1838)

Literature Cited:
- Emadzade, Khatere, Carlos Lehnebach, Peter Lockhart, and Elvira Horandl, 2010.  

Emadzade, et al. (2010) ...

Clade II-a, Arcteranthis-Beckwithia-Cyrtirhyncha-Halerpestes-Oxygraphis-Trautvetteria clade.
...
Achenes of Cyrtorhyncha have long triangular hooked beaks and almost parallel longitudinal veins (Fig. 4H) which are unique within the tribe. Although there are no obvious morphological synapomorphic characters shared between Cyrtorhyncha ranunculina and Beckwithia andersonii, these two taxa form a clade with 100% BS in tree topologies based on combined nuclear and chloroplast data (Figs. 1, 2) and have a similar distribution area.

 

Other articles:
• Golden Checklist Flora:  Ranunculus repens;  

Ranunculus repens L. “Creeping Buttercup”

 

 

Other articles:
• Golden Checklist Flora:  Ranunculus sceleratus;  

Ranunculus sceleratus L. “Cursed Buttercup”

(Syn: Hecatonia scelerata (L.) Fourr.)

 

Other articles:
• Golden Checklist Flora:  Ranunculus testiculatus;  

Ranunculus testiculatus Crantz “Bur Buttercup”

 

 

Other articles:
• Golden Checklist Flora:  Berberis repens;  

Berberis repens Lindl. “Creeping Barberry”

 

Literature Cited:
- Lindley, John, 1828.
Full Size ImageLindley's (1828) figure of Berberis repens  

Lindley (1828, t. 1176) ...

1176
BERBERIS* repens
Creeping-rooted Berberry.

HEXANDRIA MONOGYNIA
Nat. ord. Berberideae.
BERBERIS. — Supra, vol 6. fo. 487.

B. repens; foliis pinnatis 2-3-jugis; foliolis subrotundo-ovatis opacis spinoso-dentatis glaucis, fasciculis diffusis, radice repente.
Frutex humilis, ramosus, erectus. Folia sempervirentia, nunc ternata, saepius bi-trijuga, cum impare ; foliolis glabris, ovato-subrotundisy spinoso-dentatis, utrinque glaucis, nulla modo lucidis. Racemi terminales. fasciculati, diffusi, e gemmis squamaceis orti. Flores lutei.

A native of the north-western part of North America, where it was originally found by the party accompanying Captains Lewis and Clarke in their expedition across the continent of America.
From seeds procured on that occasion plants were raised in America, which have lately been sold into Europe at the rate of twenty-five dollars each. One of these now growing in the Garden of the Horticultural Society afforded our figure and the opportunity of examining the species : it had been purchased of Mr. Michael Floy, Nurseryman at New York, under the name of Berberis aquifolium.
It appears, however, from the researches of Mr. Douglas, that this is not the true B. aquifolium. That species was described by Pursh, in part from an inspection of specimens in the collection of Captain Lewis, but chiefly from the Banksian Herbarium, in which it had been placed by Mr. Menzies, who discovered it on the north-west coast of America. From this last source the drawing in the Flora Americae Septentrionalis was also taken. It is probable that the specimens in Captain Lewis's Herbarium were of the plant now under consideration; but it is also certain that those of Mr. Menzies belong to very distinct species. Hence it seems that Pursh founded two plants under the same name That he intended to call Captain Lewis's plant B. aquifolium, there can be no doubt; but it is equally certain, that in consequence of his having figured Menzies' species, the world now applies the name to the latter. This being the case, it has become necessary to distinguish the former by a new name, which has been suggested by its singular property of creeping at the root; a habit peculiar to this species among Berberries.
A hardy, evergreen shrub, flowering in April ; propagated, but with difficulty, by its creeping roots. Branches short, stiff, erect. Leaves evergreen, sometimes ternate, more frequently of two or three pairs, with an odd one ; leaflets ovate, roundish, with spiny teeth, glaucous on each side, in no degree shining. Racemes terminal, fascicled, diffuse, arising out of scaly buds. Flowers yellow.
J. L. [John Lindley]

* Berberys, according to Golius, as quoted by De Theis, is the Arabic name of the fruit.

 

Other articles:
• Golden Checklist Flora:  Argemone hispida;  

Argemone hispida A. Gray “Rough Prickly Poppy”

 

Literature Cited:
- Gray, Asa, 1849.  

Gray (1849, p. 5) ...

16. Argemone hispida (sp. nov.) : radice perenni ; caule crasso foliisque profunde pinnatifidis pube brevi cinerea undique tectis et (nervis marginibusque praesertim) setosissimis ; calyce aculeato ; corolla alba maxima ; capsula cylindrica (2-unciali) acutata spinis validis setisque horrida. — Low, sandy places around Santa Fe ; the stems 1 to 2 feet high, growing socially in great numbers ; June, July. (Also on the Upper Arkansas, &c., Fremont, Wislizenus.) — The flower is 3 or 4 inches in diameter, and accords with Dr. Lindley's figure of A. grandiflora, excepting the prickly calyx. That is a glabrous plant, while ours is not only densely setose, but is hoary throughout with a short and close hirsute pubescence. The pod is covered with very strong spines, of which the larger are often branched, and also with smaller prickles and a hoary and bristly pubescence. A. Mexicana was also collected, in two forms.

 

Other articles:
• Golden Checklist Flora:  Argemone polyanthemos;  

Argemone polyanthemos (Fedde) G.B. Ownbey “Crested Prickly Poppy”

 

Literature Cited:
- Horneman, Jens Wilkin, 1815.  

Hornemann (1815, p. 489) is the first publication of A. albiflora

Original Text
621. ARGEMONE.
1. A. mexicana Lin.: capsulis sexvalvibus sessilibus, foliis spinosis.
Hab. in Mexico, Charibaeis. ☉ {the Sun}. D.
2. A. albiflora mihi: capsulis 5-6-valvibus pedunculatis, foliis subspinosis.
Hab. — — ☉. D. intr. 1812.

Hornemann does not explain his use of the sun symbol (☉). However, most publications of this age use it to mean annual, or monocarpic. Also, it is unclear the meaning of Hornemann's two dashes, whether they mean “ditto” or “unknown.”

Literature Cited:
- Rafinesque, C. S., 1817.  

Rafinesque (1817, p. 83, no. 266) is the first publication of Argemone alba. It is now treated as a synonym of A. mexicana. So the name was not available to James (1823), see below.

Original Text
Order II.—Papaveracea.
266. Argemone alba Raf. Caule foliisque spinosis, foliis runcinatis, capsulis oblongis pentagonis spinosis semi-5-valvis Raf.—Argemone Rob. p. 464. A beautiful plant introduced in the gardens of Louisiana from Mexico, with snow-white petals, yellow stamina and violet stigma, stem three feet high. The variety of A. mexicana with white flowers mentioned by Pursh, is perhaps this species.

Literature Cited:
- Goodman, George J., and Cheryl A. Lawson, 1995.
- James, Edwin, ed., 1823.

Other articles:
• Interstate 80:   at Exit 211;

Locations: Gothenburg.  

James (1823, p. 460-1), describes a new prickly poppy encountered just west of the Sand Hills of Nebraska.

Original Text
On Monday [June] 19th, we moved on, and ascending the Platte about 30 miles, arrived in the evenong at a place where the hills on the north side close in, quite to the bed of the river. On both sides they became more broken and elevated, and on the north, they approached so near to the bed of the Platte, that we were under the necessity of travelling across them. We were glad, however, of any change of scene. The monotony of a vast unbroken plain, like that in which we had now travelled, nearly one hundred and fifty miles, is little less tiresome to the eye, and fatiguing to the spirit, than the dreary solitude of the ocean.
With this change of the surface, some change is observed in the vegetable products of the soil. Here we first saw a new species of prickly poppy,* with a spreading white flower, as large as that of the common poppy of the gardens. The aspect of this plant is very similar to that of the common poppy, except that the leaves are covered with innumerable large and strong prickles. When wounded it exudes a thick yellowish sap, intensely bitter to the taste. ...

* Argemone alba, a large plant very distinct from A. mexicana.

Goodman and Lawson (1995, p. 14) place the Long party camp on June 20th (erroneously dated June 19th) slightly west of Gothenburg, Dawson County, Nebraska, just beyond the area where the Nebraska Sand Hills make their easternmost approach to the Platte River.

Literature Cited:
- Sweet, Robert, 1830, 2nd ed..  

Argemone intermedia was published in Sweet's Hortus Britannicus (Sweet, 1838, 2nd ed., p. 585)

Original Text
PAPAVERACEÆ. p. 18.
ARGEM`ONE. p. 19.
5 intermèdia. (wh.)
Argem`one.
intermediate.
Mexico. 1828. 7. 10. H.♃

Literature Cited:
- Watson, Sereno, 1878.  

Watson (1878, part 1, p. 41) summarizes the bibliography of Argemone and places our plant as A. mexicana var. albiflora.

Original Text
A. Mexicana. Linn. Spec. 508. See syn. in DC. Syst. 2. 85. Pursh, 366. Nutt. Genera, 2, 9. James, Catalogue, 183. Elliott, 2. 13. Hook. Jour. Bot. 1. 189. Torr. & Gray, Fl. 1. 61; Pac. R. Rep. 159. Gray, Genera, 1. 112, t. 47; Pl. Fendl. 5; Manual, 59. Engelm. Wisliz. Rep. 28. Tottey, Emory's Rep. 406; Marcy's Rep. 280; Pac. R. Rep. 4. 64; Mex. Bound. 31. Scheele. Roemer's Texas, 436. Chapman, 21. Porter, Fl. Col. 6.
Var. Albiflora. DC. Syst. 2. 86; Prodr. 1. 120. Torrey, Ann. Lyc. N. Y. 2. 166; Frem. Rep. 87. Torr. & Gray, Pac. R. Rep. 2. 125. Gray, same, 12. 40.
A. albiflora. Hornem. Hort. Hafn. 489. Bot. Mag. t. 2342.
A. alba. Raf. Fl. Lud. 83. James, Long's Exp. 2. 149.
A. grandiflora. Sweet, Brit. Fl. Gard. t. 226. Lindl. Bot. Reg. T. 1264.
A. Georgiana. Croom. Am. Jour. Sci. 1. 25. 75.
A. vulgaris, var. albiflora. Spach, Hist. Veg. 7. 26.

Literature Cited:
- Fedde, Friedrich Karl Georg, 1909.  

Fedde (1909, heft (fascicle) 40, p. 283) described polyanthemos as a new variety of A. intermedia Sweet.

Original Text
Var. polyanthemos Fedde nov.var. — Herba valde, at noncompacte, sed diffuse ramosa, non corymbosa. FOlia obovata vel obovato-oblonga irregulariter et subpinnatim dentata, ad apicem late cuneata, superiora ad basim semiamplexicaulia. Flores minores valde numerosi cymis non corymbosis dispositi. Petala 2-2,5 cm longa.
Oklahoma: Kiowa Reservation (James Mooney 1895!).
Nota. Sine dubio sepalorum et imprimis fructum conditione A. intermediae valde affinis, sed differt inflorescentiis multifloria et foliis minus incisis, quibus notis valde ad var. corymbosam appropinquat.

Fedde also placed A. alba James in Long's Exp. II. (1823) 149 sec. Prain. in synonomy with A. intermedia Sweet.

Literature Cited:
- Ownbey, Gerald B., 1958.  

Ownbey (1958, p. 131) explains why A. intermedia is a confused name and a new name is introduced for our species.

Original Text
It is regrettable that a new name must be introduced for the well-known plant of the western plains which for many decades has been called A.intermedia Sweet by American botanists. The exact identity of A. intermedia has yet to be determined, but it seems certain that it is not con-specific with A. polyanthemos. The former was described from plants grown from seeds from Mexico; the latter is not found outside the United States. The presence in Mexico of argemones which in the past have been lumped with A. polyanthemos under the binomial A. intermedia, even when the original application of A. intermedia was unclear has led to much confusion. The important fact to be emphasized here is that the name A. intermedia is a nomen confusum which cannot be applied to any species of Argemone with assurance. Even though its true identity may ultimately be determined the name cannot be applied, on distributional grounds, to the taxon native to the western Great Plains and foothills of the Rocky Mountains.

Ownbey (1958, p.131) also considers and rejects A. alba James, placing it in synonomy with A. polyanthemos (Fedde) Ownbey.

Original Text
A. alba James is validly although very casually published. It applies without doubt to the same taxon as A. polyanthemos. A. alba James is, fortunately, a later homonym of A. alba Raf. which in turn is a synonym of A. albiflora Hornem. We are thereby freed from any eventuality which might require the revival of A. alba James, a step which would unavoidably lead to confusion.

 

Other articles:
• Golden Checklist Flora:  Corydalis aurea;  

Corydalis aurea Willd. “Scrambled Eggs”

 

Literature Cited:
- Willdenow, Carl Ludwig, 1809.  

Corydalis aurea was published by Willdenow (1809), in an enumeration of plants growing in the Berlin garden.

Original Text Interpretation and Comments
* 8. CORYDALIS aurea.  
C. caule ramoso erecto, siliquis linearibus pedunculo triplo longioribus, foliis glaucis, caulinis biternatis.  
Fumaria aurea. Muhlenberg. nom. inval., does not appear that Muhlenburg published this name.
Habitat in Canada. ♂ D. The Mars sign ♂ indicates the plant is a biennial. D indicates “planta semper sub dio vegetans” — the plant always grows in the open air (?)
Folia glaucescentia bipinnata, pinnis inferioribus alternis, pinnulis alternis sessilibusque simplicibus et partitis, brevibus angustis lineari-lanceolatis utrinque acutis. Flores aurei spicato-racemosi approximati.  

 

Other articles:
• Golden Checklist Flora:  Fumaria vaillantii;  

Fumaria vaillantii Loisel. “Earthsmoke” in Desv. Journ. Bot. ii. (1809) 358.

 

Literature Cited:
- Loiseleur-Deslongchamps, J. L. A., 1809.  

Original Text Translation
FUMARIA VAILLANTII N. Fumaria vaillantii nov. sp.
F. caule ramoso erectiusculo , foliis bipinnatis , pinnis 2-5-laciniatis , laciniis linearibus planis , racemis brevibus oppositifoliis , pericarpiis globosis subtuberculatis vix mucronulatis monospermis. F. lobis longioribus et angustioribus sparsis. Vaill., Bot. Par. 56, tab. 10, fig. 6. F. stem branched erectiusculo leaves bipinnatis, 2-5-fringed wings, lobes linear planes on short stalks oppositifoliis, pericarpiis spherical subtuberculatus Scarcely mucronulate monospermis. f lobes a longer and narrower spread.
Cette plante a par ses caractères principaux beaucoup de rapports avec la Fumaria parviflora ; mais ses rameaux redressés au lieu d'être étales et presque couchés sur la terre , les découpures de ses feuilles plus alongées , planes et non canaliculées; enfin, ses fleurs reugeâtres au lieu d'être blanches , la font facilement distinguer. Je l'ai trouvée cette année , en herborisant avec M. Merat , entre Chanteloup et Poissy , à sept lieues de Paris , où elle est très - commune dans les champs sablonneux ; et après l'avoir recueillie , examinée et comparée avec la F. parviflora , qui étoit aussi très-abondante dans ce canton , nous reconnoissions sans nous tromper , et rien qu'au port, les deux plantes à plus de vingt pas de distance. Cette espèce fleurit en mai et juin (.). This plant has by its main characters many links with Fumaria parviflora; but its twigs straightened instead to be spread out and almost lying on the ground, the cutouts of its more elongated leaves, flat and not canaliculate; finally, her flowers red instead of white, make it easily distinguish. I found this year, while botanising with M. Merat, between Chanteloup and Poissy, seven leagues from Paris, where it is very - common in the fields sandy; and after having collected it, examined and compared with the F. parviflora, which was also very abundant in this canton, we recognize without being mistaken, and in the port alone, the two plants are over twenty paces away. This species blooms in May and June (.).
A l'article Fumaria offïcinalis , page 437 de ma Flore , les synonymes de Vaillant doivent être rayés, puisque je les ai rapportés aux deux nouvelles espèces que je viens de décrire. To the article Fumaria offïcinalis , page 437 of my Flora, the synonyms of Vaillant must be scratched, since I brought them back to the two new species that I come from to describe.

Sébastien Vaillant (1669-1722) was French botanist who was born at Vigny in present-day Val d'Oise. While a surgeon in 1691, he was in Paris when he took as his master of botany Joseph Pitton de Tournefort (1656–1708). Vaillant devoted himself to the study of plants and worked for a time in the Jardin du Roi in Paris. He became ill and too poor to publish his Botanicon parisiensis (alphabetically or Enumeration of plants that grow in and around Paris) illustrated by Claude Aubriet. A fruit of 36 years of work, he left his work at Herman Boerhaave's home, Oud Poelgeest. The work contained engraved illustrations and was published in 1727. It is a work of particular importance in the history of botany and one of the first to describe the known flora. The standard author abbreviation Vaill. is used to indicate this person as the author when citing a botanical name.

 

Other articles:
• Golden Checklist Flora:  Alliaria petiolata;  

Alliaria petiolata (M. Bieb.) Cavara & Grande “Garlic Mustard”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Erysimum alliaria L., Sp. Pl. 2: 660 (-661) (1753).

Linnaeus (1753, v. 2, p. 660-661) ...

Original Text
Alliaria. 3. ERYSIMUM foliis cordatis. Hort. cliff. 338. Fl. fuec. 558. Mat.med. 334. Roy. lugdb. 342. Dalib. parif. 201.
  Alliaria. Bauh. pin. 110. Fuchf. hift. 104. Cam. epit. 589.
  Habitat in Europae fepibus, cultis, umbrofis. ♂
 

Arabis petiolata M.Bieb., Fl. Taur.-Caucas. 2: 126 (1808).

 

Alliaria petiolata (M.Bieb.) Cavara & Grande, Bull. Orto Bot. Regia Univ. Napoli 3: 418 (1913).

 

Other articles:
• Golden Checklist Flora:  Arabis pycnocarpa;  

Arabis pycnocarpa M. Hopkins var. pycnocarpa “Cream Flower Rockcress”

 

   

Boechera Á.Löve & D.Löve. “Rockcress”

 

 

Other articles:
• Golden Checklist Flora:  Boechera fendleri;  

Boechera fendleri (S.Watson) W.A.Weber. “Fendler's Rockcress”

 

Literature Cited:
- Gray, Asa, 1849.
- Gray, Asa, 1878-1895.

Other articles:
• Glossary:  syntopical;  

Sereno Watson in his work on volume 1 of Gray (1878) Syntopical Flora of North America described our Boechera fendleri as Arabis holboellii var. fendleri.

Original Text
164 CRUCIFERAE Arabis.
A. Holboellii, Hornem. Biennial …
Var. Fendleri, Watson, n. var. Stems often several and ascending from a biennial root, a foot high, hirsute below with simple of branched hairs, glabrous above : lower leaves roughly stellate-pubescent and petioles ciliate ; the upper glabrous : pods somewhat curved.
— From Colorado, Parry, no. 94, Hall & Harbour, no. 36 ; N. Nevada to New Mexico, Fendler, no. 27, Palmer, Rusby ; and California, Tulare Co., Coville & Funston, no. 1388. (Chihuahua, Wright, no. 1313.)

Fendler no. 27 is not listed in Gray (1849) Plantae Fendlerianae Novi-Mexicanae.

Literature Cited:
- Gray, Asa, 1895-1897.  

The variety fendleri was repeated in Robinson's edition of Gray (1895-1897, p. 186) Syntopical Flora of North America.

Original Text
A. Holboellii, Hornem. Biennial, …
Var. Fendleri, Watson, n. var. Stems often several and ascending from a biennial root, a foot high, hirsute below with simple or branched hairs, glabrous above; lower leaves roughly stellate-pubescent and petioles ciliate; the upper glabrous; pods somewhat curved. — From Colorado, Parry, no. 94, Hall & Harbour no. 36; N. Nevada to New Mexico, Fendler no. 27, Palmer, Rusby ; and California, Tulare Co., Coville & Funston, no. 1388. (Chihuahua, Wright, no. 1313.)

Literature Cited:
- Greene, Edward L., 1897.  

Two years later, Greene (1897) published Arabis fendleri differnentiating it mostly on the different pubescent compared to A. holbollii

Original Text
Arabis Fendleri. More slender than the above and less tall, mostly or wholly destitute of stellate hairs, only the lowest entire oblanceolate leaves very distinctly hirsute-ciliate, and with scattered trifurcated hairs on the lower face, all the upper parts of the plant, including all but the lower cauline leaves, glabrous and glaucus ; the sessile cauline leaves barely auricled, not sagittate : pods mostly less than 2 inches long, scarcely curved, acutish, on spreading or scarcely deflexed pedicels of 1/2 inch or more : seeds in 2 rows, small marginless.
A somewhat variable plant of subalpine situations in the Rocky Mountains from New Mexico to Wyoming ; but in none of its phases does it exhibit the pubescence of A. Holbollii, but always its own, which is chiefly conspicuous as a ciliation of the leaf-margin. The description here drawn mainly from Colorado specimens of my own collecting, which exactly match Fendler's n. 27 from New Mexico, which is the type of Mr. Watson's A. Holbollii Fendleri, in Gray, Syn. Fl. i. 164.

Literature Cited:
- Weber, William A., 1982.  

Weber (1982) published Boechera fendleri after taking Rollins to task.

Original Text
Love & Love (1976) proposed the genus Boechera to accommodate species of Arabis having the chromosome base number x=7, the type species of Arabis (alpina) having x=8. Rollins (1977) certainly had the right to criticize the Love's for not pointing out morphological evidence to support their separation on cytogenetic grounds. But at the same time, the difference in basic chromosome number, especially when it continues to be borne out on examination of other species, does represent a divergent phylogenetic line, and the genetic barrier that it presents to interbreeding of the units is sufficient justification for thinking in terms of discrete genera. However, Rollins is not scientifically objective when he says that “their describing a new genus to accommodate perfectly ordinary species of Arabis has no merit and should not be followed.” Posterity, rather than appeal to authority, should be allowed to decide the wisdom of this.
Rollins himself says earlier that “those species of North America most closely to Arabis of Eurasia have the same basic chromosome number pattern, i.e., x=8, whereas those species with a somewhat different circle of close affinity [my italics] are based on x=7.” Even a cursory examination of Arabis in the herbarium results in fairly easy separations: the loose slender root systems, large and numerous cauline leaves, most commonly toothed in Arabis, versus the short clustered caudices, small or absent and almost always entire cauline leaves of Boechera. The often very dense indument of forked or stellate trichomes of Boechera is not a characteristic feature of Arabis.
Even with Boechera removed, Arabis in America remains a genus in need of additional fragmentation. One group in particular deserving attention is the purple-flowered species endemic in the ancient Siskiyou area of southwestern Oregon and northwestern California (cf. Rollins 1977). At present, however, I am confining transfers to those taxa in the Colorado flora.
BOECHERA FENDLERI (S. Wats.) W. A. Weber, comb. Nov. Arabis holboellii var. fendleri S. Wats. In Gray, Syn. Fl. N. Am. 1:164. 1895.
Love, Askell & Doris Love. 1976. Nomenclatural notes on Arctic plants. Bot. Not. 128:497-523.
Rollins, Reed C. 1973. Purple-flowered Arabis of the Pacific Coast of North America. Contrib. Gray Herb. 204:149-154.
Rollins, Reed C. & Lily Rudenberg. Chromosome numbers of Cruciferae III. Contrib. Gray Herb. 207:101-116. 1977,

 

Other articles:
• Golden Checklist Flora:  Descurainia incisa;  

Descurainia incisa (Engelm.) Britton “Mountain Tansy Mustard”

 

Literature Cited:
- Gray, Asa, 1849.  

Gray (1849, n.s. v. 4, p. 8) from a manuscript by George Engelmann ...

Original Text
29-31. Sisymbrium incisum (Engelm. Mss.) : "annuum vel bienne ; caule glabriusculo seu puberulo seu glanduloso-piloso ; foliis pinnatisectis, segmentis lanceolatis vel lineari-lanceolatis inciso-serratis ; petalis flavis lanceolato-spathulatis calycem superantibus ; pedicellis calyce ter-quaterve longioribus, fructiferis (racemo fructifero elongato) patentibus capillaribus siliquas lineari-filiformes erecto-patentes subaequantibus ; valvis indistincte uninerviis." — Banks of streams in New Mexico ; Santa Fe Creek and Mora River ; June to August. Plant 1 to 2 feet high, blanching, at length almost glabrous. Silique about 5 lines long ; the valves with one indistinct middle nerve. Seeds linear-oblong, yellow, almost smooth, in one row. — Distinguished from S. Sophia by the longer petals, shorter silique on proportionally longer pedicels, and coarser, much less divided leaves. No. 29 and No. 31 are forms with large leaves, their lanceolate segments coarsely serrate or incised. No. 30 is a very imperfect and dubious specimen. — From Clear Water, Oregon, by Mr. Spalding, I have another form (β filipes) of the same species, with the divisions of the cauline leaves narrowly linear, sparingly incised or incisely pinnatifid, or some of them quite entire, and with fructiferous pedicels three fourths of an inch long and longer than the pods. The species would appear to stand between S. Sophia and S. tanacetifolium, L. (Hugueninia, Reichenbach), with which, like its allies, it accords in the barely one-nerved valves of the silique.

 

Other articles:
• Golden Checklist Flora:  Descurainia pinnata;  

Descurainia pinnata (Walter) Britton “Western Tansymustard”

 

 

Other articles:
• Golden Checklist Flora:  Draba reptans;  

Draba reptans (Lam.) Fernald “Carolina Whitlow Grass”

 

Literature Cited:
- Lamarck, Jean-Baptiste, 1783.
- Plukenet, Leonard, 1696.

Other articles:
• Glossary:  paronychia;  

Lamarck (1783, t. 1, p. 222) ...

Original Text Translation and Comments
21. Arabette rampante , Arabis reptans. Arabis foliis subrotundis . integerrimus , hirsutis , surculis reptantibus. N. B. Paronychia myosotis Virginiana, foliis subrotundis. Pluk. Alm. 281. Tab. 51. f. 5. Arabis with roundish, entire, hirsute leaves, with young branches creeping.

“N. B.” — unknown meaning.

“Pluk. Alm.” — Plukenet, Leonhart. 1696. Almagestum botanicum. London.

Cette plante, encore peu connue des Botaniftes, nous paroit devoir etre rapportee a ce genre. Sa racine eft fibreufe, & pouffe de fon collet quelques jet fteriles, feuilles & rampans. Ses feuilles font ovales-arrondies, retrecies vers leur bafe, tresentieres, & velues ou hifpides. Ses tiges font greles, nues, ou chargees d'une ou deux fleurs difpofees en grappes, auxquelles fuccedent des filiques pedonculees, d'une longueur mediocre. Cette plante paroit originaire de la Virginie. This plant, still little known to Botaniftes, seems to us to be related to this genus. Its root is fibrous, and its root pod has a few sterile streams, leaves & crawlers. Its leaves are oval-rounded, narrowed towards their base, presentieres, & hairy or hispid. Its stems are slender, bare, or loaded with one or two flowers arranged in clusters, to which follow pedunculate threads of mediocre length. This plant appears to be native to Virginia.

Literature Cited:
- Fernald, M. L., 1934.  

 

 

Other articles:
• Golden Checklist Flora:  Erysimum asperum;
• Glossary:  divaricate;  

Erysimum asperum (Nutt.) DC. “Prairie Wallflower”

 

Literature Cited:
- Pursh, Frederick, 1816.  

Original Text
527. ERYSIMUM. Gen. pl. 1090.
lanceolatum. 3. E. foliis lanceolatis inferioribus dentatis, unguibus calyce longioribus,' laminis orbiculato-obovatis, stigmate sessili. Ait. Kew. Ed. 2. t. 4. p. 116.
Chieranthus erysimodes. Willd. Sp. Pl. 3. p. 514.
Icon. Jacq. Fl. Austr. 74.
On the banks of the Missouri. ♂. June. v. s.

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Nuttall, 1818, publication details;
• Glossary:  asperate;

Locations: White River.  

Nuttall (1818, v. 2., p. 69) …

Original Text
462. CHEIRANTHUS. L. (Wall-flower. Stock.)
Calix closed; 2 of the leaves gibbous at the base. Petals dilated. Disk of the germ biglandulous. Silique compressed or terete. Stigma bilobed. Seeds flat, sometimes marginated.
Herbaceous or suffruticose; leaves more or less pubescent, entire or pinnatifid; flowers yellow or purple, large, and often odorous. A genus very nearly allied to Hesperis and to the entire leaved species of Erysimum, excluding E. Alliaria.
Species. 1. C. Pallasii. Ph.—North West Coast. 2. *asper. Stem simple, and acutely angular; leaves canescently polise, sublinear, entire, margin acculeately and retrorsely asperate, radical fasciculated attenuated-sublanceolate, acute, retrorsely toothed; silique very long quadrangular and divaricate; claws of the petals longer, than the calix. Cheiranthus erysimoides. Ph. Hab. On the plains of the Missouri, commencing near the colfluence of the White river. Fl. June. Obs. Biennial. Stem 12 to 18 inches high, mostly simple, but now and then branching towards the summit. Leaves every where covered with white short, retrorse, strigose and appressed hairs; margin aculeolate, stem leaves crowded, 2 or 3 lines wide and 2 inches long. Flowers very similar to those of C. Cheiri, and almost equally odorous. Calix oblong, 2 of the leaves distinctly gibbous at the base. Petals dilated, claws long, limb broud obovate, bright yellow. Stigma bilobed. Silique 2 or 3 inches long, spreading, 4-sided, 2 of the angles asperate.
A genus of about 40 species, indigenous to Europe and the temperate and colder parts of Asia and Africa in both hemispheres.

Literature Cited:
- DeCandolle, Augustin Pyramus de, 1818.

Other articles:
• Glossary:  asper;  

DeCandolle (1821, v. 2, p. 505) ...

Original Text
29. Erysimum asperum.
E. foliis lineari-oblongis, inferioribus dentato-runcinatis, cauleque pubescentibus scabris, siliquis patentibus, stylo brevissimo crasso.

Erysimum lanceolatum. Pursh. fl. bor. am. 2. p. 436.
Cheiranthus asper. Nutt. gen. am. 2. p. 69. *
Hab. in campis Missouriensibus (Nutt.). (2). fl. jun. (v. s. sp. in h. Banks et Lamb.)
...

 

Other articles:
• Golden Checklist Flora:  Erysimum capitatum;  

Erysimum capitatum (Hook.) Greene “Sanddune Wallflower”

 

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.  

Hooker (1829, v. 1(1), p. 38) ...

Original Text
1. C. capitatus ; subasper, foliis lineari-lanceolatis magis minusve dentatis vel integris basi longe attenuatis cauleque pube bipartita arctissime appressa strigosis, floribus (majusculis) dense corymbosis, "siliquis pedicello triplo longioribus." — Douglas, MSS. C. asper. Schlecht. et Cham, in Linncaea, v. 1. p. 14.
[Latin diagnosis omitted.]
Hab. Common on rocky places of the Columbia, near the sea; and at Puget Sound, Douglas. Chamisso found it on the coast of California. — I regret that I do not possess the fruit of this plant ; but there can be no question of the accuracy of Chamisso and Schlechtendal, who assure us that the cotyledons are accumbent. It must therefore be separated from Erysimum, with the species of which, and especially with E. lanceolatum, as is well observed, it bears a very close affinity. It is quite a different plant from the Erysimum asperum, De Cand., the Cheiranthus asper of Nuttall, and which is also supposed to be the Erysimum lanceolatum of Pursh; but I know not upon what authority. I am indebted for specimens to M. de Chamisso, which agree in every respect with those from Mr. Douglas : both are remarkable for the great attenuation of the base of the leaves. It has been found by no other of our North American travellers : but is now cultivated in the gardens of the Horticultural Society, from seeds sent by Mr. Douglas from the Columbia.

Literature Cited:
- Greene, Edward Lee, 1891-97.  

Greene (1891, p. 269) ....

Original Text
2. E. capitatum, Greene. Dougl.; Hook. Fl. i. 38 (1829), under Cheiranthus. Stout and low, ½ — l½ ft. high, spariugly pubescent with appressed bifid or 2-parted hairs; leaves narrow, entire, or sinuately or angularly toothed or lobed: fl. large, cream-color or yellowish, in a depressed terminal corymb, scarcely fragrant: pods in a short raceme; valves nearly flat, with a strong midvein, 1½ lines wide, the whole 1½ — 2½ in. long, abruptly and stoutly short-pointed: seeds flattened. — Among the sandy or rocky hills of the seaboard only, from Monterey northward to Mendocino Co.; easily mistaken for E. aspertum, but we have not seen it with even yellow, much less orange-colored flowers. The petals are broader than in that species, but at San Francisco they are invariably almost white. Feb. — May.

 

Literature Cited:
- Watson, Sereno, 1888.

Other articles:
• Golden Checklist Flora:  Notes on Physaria montana, Watson, 1888;  

Lesquerella S.Watson

Watson (1888, v. 23, new ser. v. 15, p. 249) published a new genus Lesquerella saying:

Original Text
* The Old World genera of the Vesicaria and Alyssum group are variously understood by European botanists and are very troublesome. The species of Vesicaria upon which all are agreed (V. utriculata, V. Graeca and V. glabrescens, the first being one of the two original species) have stout erect leafy stems from a suffrutescent base, glabrous, or pubescent below with appressed 2-parted or somewhat stellate hairs, with large Erysimum-like flowers, very large globose coriaceous pods, nerveless septum, and wing-margined seeds. This is the genus as it is generally accepted on the continent, …
… [text omitted.] …
The American species differ from them all, more or less positively. In all our species there is a distinct nerve extending from the apex to the middle of the septum or beyond. The filaments are never toothed or appendaged; the petals are never narrowly unguiculate, and, except in one or two species, are yellow ; the ovules are never solitary in the cells, and the pubescence is always more or less stellate or lepidote.
… [text omitted.] …
To the species that have hitherto been placed in Vesicaria, I would, therefore, now give the generic name Lesquerella (in preference to reviving Lesquereuxia, the former name of a genus now merged in Siphonostegia), in honor of our venerable and in every way worthy veteran palaeontologist and bryologist, Leo Lesquereux. Our one flat-podded species that has been referred to Alyssum (A. Lescurii) appears to differ in no other respect than its less convex valves from a somewhat distinct group of species which can be separated, however, only as a section from the rest.

 

Other articles:
• Golden Checklist Flora:  Lunaria annua;  

Lunaria annua L. “Money Plant”

 

 

Other articles:
• Golden Checklist Flora:  Nasturtium officinale;  

Nasturtium officinale R. Br. “Watercress”

 

Literature Cited:
- Aiton, William T., 2nd. ed., 1810-13.  

W. T. Aiton (1812, v. 4, p. 110) noted that N. officinale was grown at Kew.

Original Text
officinale. 1. N. foliis pinnatis : foliolis ovatis subcordatis repandis.
Sisymbrium Nasturtium. Willden. sp. pl. 3. p. 489. Curtis lond. Engl. bot.855.
Water-cress
Nat. of Britain.
Fl. June–September. H. ♃
   

Literature Cited:
- Al-Shebaz, I. A., 2014.  

 

   

Noccaea fendleri (A. Gray) Holub “Fendler's Pennycress”

 

Literature Cited:
- Gray, Asa, 1849.  

Gray (1849, p. 10) referred Fendler's collection to Thlaspi cochleariforme

Original Text
44. Thlaspi cochleariforme, DC. Syst. 2. p. 381 ; Deless. Ic. Select. 2. t. 52. Santa Fe Creek, in the mountains ; March to June.

The image of the original publication, which came from George Engelmann's library, has several pencilled notations, such as “No!” and “Th. Fendleri Agr / Pl Wrightii 2. 14.”

Literature Cited:
- Gray, Asa, 1852-1853.  

Gray (1853, part 2, p. 14) published Thlaspi fendleri as a correction to his identification ...

Original Text
Thlaspi Fendleri: foliis carnosulis subintegerrimis, radicalibus petiolatis ovalibus, caulinis sagittato-oblongis amplexicaulibus ; racemo etiam fructifero brevi conferto ; floribus majusculis ; petalis calyce triplo longioribus; siliculis lato-obovatis subalatis basi acutis apice immaturis sinu aperto leviter obcordatis maturis truncato-subintegris, loculis 3-4-ovulatis ; stylo filiformi fructu dimidio breviore. — T. cochleariforme, Gray, Pl. Fendl p. 10, non DC, nec Hook. — On the Organ Mountains, northeast of El Paso ; May : mostly in fruit. (1322.) — Plant a span high, or lower ; the raceme an inch, or, even in full fruit, not over 2 inches, in length ; the pedicels closely approximate, spreading, in fruit 5 or 6 lines long. Petals (in Fendler's specimens) 4 lines long. Ovary obcordate by a broad and shallow sinus ; the cells 3 - 4-ovulate. Ripe silicles 4 lines long, with sharp but slightly winged edges, obscurely emarginate or even truncate at the broad summit ; the style a line and a half or two lines long. — The specimens of Fendler were mostly in flower only : mine with young fruit have the raceme more loose than in those gathered this spring by Mr. Wright ; which, being in fruit, plainly show that I was wrong in referring the New Mexican plant to T. cochleariforme. The latter has the fructiferous raceme long and loose, and has likewise smaller flowers, narrower pods, with a deep and narrow apical sinus, and a short style. Our plant is more like T. praecox ; which also has smaller flowers, and winged pods, with a very deep notch.

Literature Cited:
- Holub, Josef, 1998.

Other articles:
• Golden Checklist Flora:  Notes on Thlaspi, Mummemhoff, 1997;  

Holub (1998, v. 70, n. 2, p. 108) published some names in Noccaea for American taxa to complete the genus. This is explained in a note to the first name proposed.

Original Text
N o t e : F. K. Meyer ( 1973) undertook a generic reform (or rather a radical revolution) of the earlier standing concept of the genus Thlaspi . It was shown, that - especially on the basis of the anatomical structure of the seed testa - the small group Thlaspi s.s. with the type species T. arvense L., differs substantially from other groups of Thlaspi s.l., including many species. In Central Europe (s.l.) a few representatives of Microthlaspi F. K. Meyer and numerous species of Noccaea Moench occur. F. K. Meyer's classification has not been followed very much by other authors. The present author (Holub 1983: 205). accepted the genus Noccaea in circumscription proposed by F. K. Meyer. Most recently Kerguelen and Cerepanov also supported this classification by some new proposals of nomenclatural combinations with Noccaea. Justification of the generic exclusion of Thlaspi s.s. from the remainder of that genus (Thlaspi s.l.) and by this the acceptance of the genus Noccaea, are indicated by the most recent studies based on modern methods of molecular research (Mummenhoff et al. 1997; chloroplast DNA restriction-site variation - Mummenhoff & Koch 1994, Zunk et al. 1996; isoelectric focusing analysis of Rubisco - Mummenhoff & Zunk 1991). Nomenclatural combinations for taxa of the genus Noccaea were proposed especially for European and Oriental species of that genus. Here also some new nomenclatural combinations for taxa from the American continent are proposed.
Noccaea fendleri (A. Gray) Holub, comb. nova. — Bas.: Thlaspi fendleri Asa Gray, Smithsonian Contrib. Knowl. 5(6): 14, [= Plantae Wrightianae 2: 14], Washington 1853 .

 

Other articles:
• Golden Checklist Flora:  Noccaea fendleri glauca;  

Noccaea fendleri (A. Gray) Holub ssp. glauca (A. Nelson) Al-Shehbaz & M. Koch “Alpine Pennycress”

 

Literature Cited:
- Nelson, Aven, 1896.  

Thlaspi alpestre var. glaucum A.Nelson, Wyoming Agric. Exp. Sta. Bull. 28: 84 (1896).

Nelson (1896, p. 84) ...

Original Text
Thlaspi alpestre glaucum, n. var.
The perennial basal part of stem rather freely branched, herbaceous stems simple and erect, 6-10 inches high; radical leaves broadly to narrowly elliptical, entire or obscurely repand-denticulate; cauline deltoid-auticulate entire. It also differs from the species in the glaucus hue of the leaves, the laxer inflorescence and will marked notch at the apex of the capsule as well as in its habitat. The species flowers in early speint on open hillsides; the variety was collected in the forest almost at timber line, growing in the thick beds of Spruce needles. La Plata Mines, August 21, 1895 (No. 1777).

Literature Cited:
- Nelson, Aven, 1898a.  

Nelson (1898, v. 25, p. 275) ...

Original Text
Thlaspi glaucum (A. Nelson).
Thlaspi alpestre glaucum A. Nelson, First Rep. Fl. Wyo. 84. 1896.
The perennial, basal part of stem freely branched, the several or many herbaceous stems simple, erect or ascending, 1-3 dm. high ; root leaves numerous, petioled, from oval to oblong or obovate, entire or obscurely repand-denticulate ; cauline leaves deltoid-auriculate or oblong, entire, 10-20 mm. long, smooth or even glaucous ; raceme dense in flower, much elongated in fruit (5-15 cm.) ; sepals greenish, thin, ovate ; petals white, spatulate, about 6 mm. long ; pods obovate, obtuse or emarginate ; pedicels divaricate, 10-14 mm - long.
In 1897 an abundance of material of this form was secured, an examination of which leads me to think that it is worthy of specific rank. Its habitat is sub-alpine in open parks or among open spruce timber. It is separated from T. alpestre by its uni- formly larger size, numerous stems, numerous and larger stem leaves, longer pedicels and raceme (in fruit), longer, greenish sepals, etc.
Type specimen in Herb. University of Wyoming, no. 4176, Battle Lake, August 16, 1897. Prof. Henderson's no. 2893 from Kendrick, Idaho seems to be nearly the same.

Literature Cited:
- Holub, Josef, 1998.  

Holub (1998, v. 70, n. 2, p. 108) published some names in Noccaea for American taxa to complete the genus. This is explained in a note to the first name proposed.

Original Text
Noccaea glauca (A. Nelson) Holub, comb. nova. — Bas.: Thlaspi alpestre glaucum A. Nelson, First Report Fl. Wyoming, 84, 1896 (n.v.). – Syn.: Thlaspi glaucum (A. Nelson) A. Nelson, Bull. Torrey Bot. Club 25: 275, New York 1898.

 

Literature Cited:
- Ackerfield, Jennifer, 2015.
- Al-Shehbaz, Ihsan A., and Steve L. O'Kane, 2002.

Other articles:
• Golden Checklist Flora:  Notes on Physaria montana;
• Glossary:  cytology;  

Notes on Physaria

The taxonomic world knew for a long time that keeping Physaria and Lesquerella separate was very tenuous. More recently molecular data clearly showed that Physaria is nested within Lesquerella. Also, the data showed that Physaria evolved more than once from Lesquerella. Regardless of how many times Physaria evolved from Lesquerella, Physaria is polyphyletic and Lesquerella definitely paraphyletic. The two genera could no longer be maintained separately on either molecular, morphological, cytological, biogeographic, or ecological grounds. Therefore the two genera should be combined. Physaria is the older of the two names and thus has priority, but there are far more species in Lesquerella. It might make more sense to conserve Lesquerella and transfer the Physaria thereto. An proposal to conserve the name Lesquerella against Physaria was made to the Committee for Spermatophyta (2000). However, the proposal was denied. Therefore, names in Physaria were proposed for most of the former Lesquerella (Al-Shehbaz & O'Kane, 2004).

One of the taxa transferred to Physaria was P. montana a bladderpod found in Jefferson County, including Golden.

Seven taxa of Physaria have been reported for Jefferson County. Four of the taxa are unlikely to occur here, as the collections are misidentified or otherwise quesitonable data. The three confirmed Jefferson County taxa are:

  • P. bellii G. A. Mulligan
  • P. montana (A. Gray) Greene
  • P. vitulifera Rydb.
The four spurious reports of Physaria in Jefferson County are:
  • P. acutifolia Rydb. — there may be two collections of P. acutifolia from the Evergreen - Kittridge - Parmalee Gulch area. They are old records and a bit sketchy. Otherwise this is an western slope taxon.
  • P. didymocarpa (Hook.) A. Gray — misidentified, two vouchers at YU, duplicate at CS9266 determined P. vitulifera.
  • P. floribunda Rydb. — western slope taxon. Data point from NY showing P. floribunda in Golden is misidentified.
  • P. rollinsii G. A. Mulligan — generally known only from the Gunnison River basin, except for Rollins No. 5151, July 6, 1951, 2 miles west of Deckers, GH 01691383, which has been annotated P. rollinsii, date and name of annotator unknown.

The following key is abstracted from Ackerfield (2015).

(1b.) Fruit (and ovary) pubescent with stellate hairs … 4
(4b.) Plants not present in the alpine, found below 11,000 ft in elevation … 6
(6b.) Inflorescence usually obviously exceeding the leaves, or if included then the plants otherwise unlike the above; plants sometimes mound-forming, but the mounds usually not small and button-like … 8
(8b) Mature fruiting pedicels S-curved (sigmoid), ascending, or sometimes horizontal … 12
(12a) Plants of the eastern slope … 13

(13a) Fruit entire at the apex, not or only slightly inflated; ovules 4-20 per ovary … 14
(14a) Basal leaves suborbicular, obovate, or elliptic, mostly over 4 mm wide (rarely narrowly elliptic and 2-3 mm wide), usually with a well-defined petiole; stem leaves usually secund; ovules (8) 12-24 per ovary … P. montana

(13b) Fruit notched at the apex (didymous), usually inflated; ovules 4 per ovary … 16
(16b) Plants of the foothills, absent from the northeastern plains; fruit not conspicuously broader at the apex or appearing flared from the base … 17

(17a) At least some basal leaf margins deeply and broadly incised, rarely almost entire … P. vitulifera

(17b) Basal leaf margins shallowly dentate or entire … 18
(18a) Basal leaves gradually tapering to an ill-defined petiole; fruit 2-8 mm wide … P. bellii

   

Notes on Physaria bellii Mulligan

 

Literature Cited:
- Mulligan, Gerald A., 1966.  

10. Physaria bellii G. A. Mulligan, Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. 1966.

Other articles:
• S. & W. Deer Creek Canyon Road:  74500;
• Golden Checklist Flora:  Notes on Physaria vitulifera, Borland, 1988;
• Owens Street:  78000;  

Jim Borland collected Physaria bellii and P. vitulifera twice: 12 May 1988 and 22 June 1988.

Collections of P. bellii were:

  • Verbatim Date: 12 May 1988. Locality: United States, Colorado, Jefferson, On north road cut as Deer Creek Canyon road enters the canyon. Habitat: Road cut. Reproductive Condition: Flr & Frt. https://swbiodiversity.org/seinet/collections/individual/index.php?occid=13848645
  • Verbatim Date: 22 June 1988. Locality: United States, Colorado, Jefferson, 100 ft. S of junction of West Ute Ave. and Owens Street, west side of road. Habitat: On limestone/shale outcrop. Reproductive Condition: Frt. https://swbiodiversity.org/seinet/collections/individual/index.php?occid=13848646

Literature Cited:
- O'Kane, Steve L., 1988.  

O'Kane (1988, p. 463) ...

O'Kane, Steve L. (Colorado Natural Areas Program), Colorado's Rare Flora. Great Basin Naturalist. v. 48, n. 4, p. 433-484. 31 October 1988.

Original Text Comments
Physaria bellii Mulligan, Canadian J. Bot. 44: 1662-1663. 1966.  
Map 66  
Family. — Brassicaceae (Cruciferae).  
Federal status. — Category 2.  
Bells twinpod, endemic to the Niobrara Formation, grows on fine-textured soils derived from black shale high in calcium in the southern part of its range and from light- colored, limey shale in the north. The species is scattered along the foothills and hogbacks from Fourmile Canyon near Boulder to Box Elder Creek north of Owl Canyon. One old collection, however, exists from Cheyenne Mountain near Colorado Springs (Churchill s.n. MO), and a 1988 report (J. Borland, personal communication, specimen verified by W. A. Weber) places the species southwest of Denver in Jefferson County on a small outcrop of Niobrara shale at the mouth of Deer Creek Canyon. As Cheyenne Mountain is composed of Pikes Peak Granite, it is likely that the collection locality is in error unless a disjunct population occurred at the foot of the mountain on Pierre Shale. Bells twinpod is usually found in shrub communities dominated by Rhus trilobata and Cercocarpus montanus at elevations of 1,580 to 1,750 m. Oryzopsis and species of Stipa are common associates. Although not threatened everywhere in its range, threats to individual populations can be significant. Some populations are threatened by road construction and maintenance. The populations near LaPorte could be impacted by mining for cement products. The effects of grazing are not known. Two vouchers of J. Borland collections are at COLO.
    Boulder County: 
      TIN R70W: 
        Weber 3270, 4/10/47 (COLO, CS); 
      T2N R70W: 
        Naumann 2, 6/7/84 (CS); 
        Neely 2514, 5/5/85 (CS, UTC); 
      T2N R71W: 
        Denham & Denham 1315, 4/16/67 (COLO); 
        Gambill s.n., 5/7/84 (KHD);
        Mulligan & Crompton 2898, 8/24/64 (Isotype: COLO); 
        Ramaley 1016, 5/30/05 (COLO);
        Rolhns 7948, 5/24/79 (COLO, MO, RM);
        Weber 3372, 5/22/47 (COLO, CS); 
        Wiegand & Upton 3320, 6/12/22 (MO); 
        Wynn s.n. , 5/10/83 (KHD); 
      T3N R71W: 
        Wilken 12887, 5/18/77 (CS). 
    El Paso County: 
      T15S R67W:
        Churchill s.n., 6/22/12 (MO). 
    Larimer County: 
      Location obscure: 
        Osterhout 5616, 6/25/17 (COLO); 
      T4N R70W: 
        Popp s.n., 6/18/83 (CS); 
      T4S R69W: 
        Robinson s.n., 5/25/80 (COLO); 
      T5N R69W: 
        Naumann & Trout 4, 7/2/84 (CS); 
      T6N R70W: 
        Popp s.n., 6/29/83 (COLO); 
        Schromberg s.n., 5/8/83 (COLO); 
      T7N R69W: 
        Naumann & Trout 3, 5/6/84 (CS); 
        Popp s.n., 6/18/83 (CS); 
      T8N R69W: 
        Crandall 212, 5/14/1890 (CS); 
        Crandall 425, 5/2/1896 (CS, MO); 
        Crandall 427, 6/7/1898 (CS); 
        Crandall 426, 5/7/1898 (CS);
        Wilken 14302, 5/6/85 (CS); 
      T9N R69W: 
        Lanham s.n., 6/8/80 (COLO); 
        Neely 2734, 6/2/85 (CS, UTC); 
        Popp s.n., 6/17/83 (CS); 
      T10N R69W: 
        Ramaley 2743, 6/14/07 (COLO).
  
 

 

Other articles:
• Golden Checklist Flora:  Physaria montana;  Notes on Physaria;  

Physaria montana (A. Gray) Greene “Mountain Bladderpod”

 

Literature Cited:
- Gray, Asa, 1863.  

Gray (1863, v. 15, p. 58) article titled, “Enumeration of the Species of Plants collected by Dr. C. C. Parry, and Messrs. Elihu Hall and J. P. Harbour, during the Summer and Autumn of 1862, on and near the Rocky Mountains, in Colorado Territory,”

Original Text Comments
… 49. Vesicaria montana, n. sp.* from the middle mountains ; also collected last year at Eureka by Mr. Howard, but without fruit. … The asterisk (*) refers the reader to a footnote that is reproduced below.

The number 49 corresponds to a Hall & Harbour collection number 49 that is designated the holotype at GH (https://swbiodiversity.org/seinet/collections/individual/index.php?occid=23018520&clid=0)

I assume the Eureka place name is Eureka, San Juan County, Colorado, a former mining town 19 miles northeast of Silverton.


*Vesicaria Montana (sp. nov.): argenteo-incana : caulibus e radice perenni diffusus foliosis: foliis spathulatis, radicalibus subovatis petiolatis nunc 1-2-dentatis; racemo fructifero elongato; silicula ovali seu ellipsoidea cano-pubescente stylo gracili longiore pedicello patente sursum curvato paullo breviore. Habit of V. Ludoviciana, argyrea, and argentea; well-marked by the oval or oblong silicle (which is. in some specimens, 3 lines in length, but of scarcely half that breadth, while in others it is shorter and broader, barely oval in outline,) hoary, with a fine stellular pubescence, one-third longer than the style, commonly one-third or one-half longer than the pedicel, nearly terete; the valves of the same rather firm texture as those of V. Ludoviciana, more convex than those of V. alpina. Seeds four or six in each cell, wingless. Petals spatulate, light yellow. Filiments filiform.
This is a footnote at the bottom of page 58.

Literature Cited:
- Watson, Sereno, 1888.

Other articles:
• Golden Checklist Flora:  Notes on Lesquerella, Watson, 1888;  

Watson (1888, v. 23, new ser. v. 15, p. 249) placed V. montana A. Gray in Lesquerella saying:

Original Text
10. L. Montana. Pubescence often evidently stellate; caudex rarely branched: leaves oblanceolate, the radical often subovate on slender petioles and obscurely toothed: petals spatulate: pods 3 lines long, with long slender style; cells 4-8-ovuled. — V. montana, Gray. Northern Colorado and southern Wyoming, near and on the mountains.

Literature Cited:
- Greene, Edward Lee, 1891-97.  

Greene (1891, p. 249) in his Flora Franciscana ...

Original Text Comments
3. PHYSARIA, A. Gray. Low herbs (our species perennial) silvery- canescent with a dense close stellate pubescence, entire or pinnatifid leaves, and racemose yellow flowers of middle size; calyx more or less persistent as in Alyssum. Pods globose or ovoid, or the valves more inflated and distended, nerveless. Style long, persistent. Seeds few, in 2 rows, flattened, rarely somewhat margined: cotyledons accumbent.  
* Pods didymous. — Typical Physaria.  
1. P. didymocarpa, Gray, Gen. Ill. i. 162 (1849); Hook. Fl. i. 49. t. 16 (1829), under Vesicaria. Leaves rosulately crowded on a short crown or caudex, broadly spatulate, occasionally somewhat lyrate, those of the decumbent flowering branches oblanceolate, entire: racemes short: pods 2 — 6 lines broad, i. e., more or less widely didymously-inflated, the partition narrow or nearly obsolete. — East of the Sierra, thence common to Colorado. While there are collections from Colorado, including Jefferson County, that are determined P. didymocarpa, I think they are mis-identified specimens of P. vitulifera.

The species is known to occur in Wyoming.

* * Pod globose or ovate. — Genus Lesquerella, Wats.  
2. P. montana, Greene. Gray, Proc. Philad. Acad. 58 (1868), under Vesicaria. Habit of the preceding: rosulate radical leaves orbicular or obovate, long-petioled, those of the branches oblanceolate or spatulate, entire, or with few teeth : fl. 3 lines long: pods oblong-ovoid, 2½ lines long, on slender recurved pedicels; style a third shorter. — On Lassen's Peak, Lemmon, thence eastward and northward.  

 

Literature Cited:
- Mulligan, Gerald A., 1966.

Other articles:
• Golden Checklist Flora:  Physaria vitulifera;  

Physaria vitulifera Rydb. “Roundtip Twinpod”

It appears to me that, in addition to Rydberg's type with two vouchers, there are two “approximate topotypes:” Ripley & Barneby (1950) #10475 (GH 1691622, CAS 303257) and Mulligan (1963, GH 1691620).

Mulligan's collection was likely in connection of his study of the genus (Mulligan, 1966).

Other articles:
• Interstate 70:   in Clear Creek Canyon;  

Physaria vitulifera, Parry, 1861

The oldest known collection of P. vitulifera was made by C. C. Parry, in his first expedition to the Rocky Mountains. We know that he collected in upper Clear Creek and traveled in the direction of Pikes Peak, but he did not keep a diary or detailed itinerary.

Plants of Colorado
Brassicaceae

Physaria vitulifera Rydb. Bull. Torrey Bot. Club 28: 278. 1901.  Roundtip Twinpod.

Clear Creek Headwaters, Clear Creek County, Colorado. United States of America, Colorado, From the head-waters of Clear Creek, and the alpine ridges lying east of "Middle Park," Colorado Territory.

C. C. Parry 101. 1861

There are three existing vouchers of Parry's collection. They were all initially determined P. didymocarpa Gray. Several indicate that they may be an undescribed variety. One of them, GH1639631, was annotated P. vitulifera by Rollins in 1938. Mulligan agreed in 1966 and stamped the voucher “STET.”

The other two vouchers, PBRU39332 and YU137402, remain determined P. didymocarpa.

Other articles:
• Interstate 70:  Idaho Springs;  

Physaria vitulifera, Rydberg, 1895

Brassicaceae

Physaria vitulifera Rydb.  Roundtip Twinpod.

Idaho Springs, Clear Creek County, Colorado. United States of America, Colorado, Clear Creek Co., Idaho Springs 39.7425°N, 105.5136°W. WGS 1984

P. A. Rydberg 26-Aug-1895

Vouchers: NY3188335, RM29916.

There are 11 other collections at NY by Rydberg with the same date and location of Idaho Springs.

Literature Cited:
- Rydberg, Per Axel, 1901.  

Physaria vitulifera, Rydberg, 1901

Rydberg (1901) described a new Physaria that he called P. vitulifera from a collection he made at Idaho Springs. An image of this specimen at NY is available through SEINet. Rydberg (1895) also cited a collection by Parry made in 1861 in the headwaters of Clear Creek. An image of this voucher at GH is also available through SEINet.

Original Text Interpretation
Physaria vitulifera sp. nov.  
A densely tufted finely stellate-pubescent perennial with deep tap-root. Basal leaves numerous, fiddle-shaped, the larger 4-5 cm. long ; terminal lobe nearly orbicular to broadly obovate, subentire, obtuse ; upper sinuses rounded : lateral lobes 1-2 pairs, much smaller : stems ascending, or decumbent, 1-2 dm. high : stem-leaves 1-2 cm. long, obovate or oblanceolate, entire : pedicels short, usually curved in fruit : sepals about 4 mm. long, oblong, acute : petals clawed, 8-9 mm. long : fruit obtuse at the base, deeply divided above ; cells much inflated, round-obovate, divergent, about 5 mm. in diameter : style about 5 mm. long.  
This species, as well as the two following, differs from P. didymocarpa in the smaller fruit, which is divided only above, not at the base. In this respect they resemble P. Newberryi ; but the fruit is much smaller and not flattened on the sides. P. vitulifera differs from the following in the form of the leaves and the curved pedicels. It grows in dry places at an altitude of about 2500 m. The "following" are Physaria floribunda and Physaria acutifolia.
Colorado : Idaho Springs, 1895, P. A. Rydberg (type); Middle Park, 1861, C. C. Parry, 101.  

Rydberg's (1901) description of P. floribunda cites “Hills about Golden, 1892, Crandall, 53.” This collection can be found by searching SEINet for collections of Physaria by Crandall in 1892. Vouchers at CS and GH have been determined P. vitulifera. A voucher at YU has two specimens, determined as P. didymocarpa, one labelled as “Hills about Golden” and the other as “Hills about Dolores.” The Golden specimen looks more like Physaria montana than either P. vitulifera, P. floribunda, or P. didymocarpa.

 

Physaria vitulifera, Ripley & Barneby, 1950

On July 14, 1950, Ripley and Barneby made 9 collections from Breckenridge, Dillon, Loveland Pass, and Idaho Springs.

Plantae Occidentales Selectae
Brassicaceae

Physaria vitulifera Rydb. Bull. Torrey Bot. Club 28: 278. 1901.  Roundtip Twinpod.

Idaho Springs, Clear Creek County, Colorado. United States of America, Colorado, Clear Creek County, Idaho Springs Habitat: steep banks of granite gravel. (approx. topotype)

H. D. Ripley & R. C. Barneby 10475. 14-Jul-1950

The label notation of “(approx. topotype)” suggests they were looking for P. vitulifera at its type location of Idaho Springs. While there, they also collected two Astragalus, A. flexuosus and A. parryi. Voucher: GH16911622, CAS303257.

 

Physaria vitulifera, Mulligan, 1963

Brassicaceae

Physaria vitulifera Rydb.  Roundtip Twinpod.

Idaho Springs, Clear Creek County, Colorado. United States of America, Colorado, Clear Creek County, Northwest outskirts of Idaho Springs Elev. 2178 m.

Collected by permit: Permit Unspecified, issued: , to: .

G. A. Mulligan 2765. 18-Jun-1963

Voucher: GH1691620

Other articles:
• CO Highway 119:   near Russell Gulch;  

Physaria vitulifera, Higgins, 1968

The Higgins collection is probably not an “approximate topotype.” There is one voucher of P. vitulifera that incorrectly implies that Higgins collected the plant at or near Idaho Springs. The label of RM343593 states the collection was made in Clear Creek County, 4 miles south of Black Hawk. However, Higgins other collections on this day were made 7 miles west of Boulder and 4 miles south of Black Hawk. Of the collections 4 miles south of Black Hawk, only RM343593 gives the county as Clear Creek County, whereas the other seven vouchers of collections at that location give the county as Gilpin County. I think it is far more likely that Higgins was 4 miles southwest of Black Hawk on CO Highway 119. This would place his collection in Clear Creek Canyon approximately at the mouth of Russell Gulch. This location is 4 miles northeast of Idaho Springs.

Plants of Colorado
Brassicaceae

Physaria vitulifera Rydb. Bull. Torrey Bot. Club 28: 278. 1901.  Roundtip Twinpod.

Clear Creek Canyon, Gilpin County, Colorado. U.S.A., Colorado, Clear Creek, Ca 4 mi S of Black Hawk. Reproductive Condition: flowering and fruiting Habitat: Sandy soil; growing in the bottom of creek bed with Astragalus parryi.

Larry C. Higgins 1494. 4-Jun-1968

Voucher: RM343593, UTC176647

Other articles:
• U. S. Highway 6:  near CO Hwy 119;  

Physaria vitulifera, Mellichamp & Neson, 1968

This Mellichamp and Nesom collection is not an “approximate topotype” Being made 2.3 miles east of the junction of US Hwy 40 and US Hwy 6, it would be very close to the junction with CO Highway 119, and thus about 7 miles east of Idaho Springs.

Plants of Colorado
Brassicaceae

Physaria vitulifera Rydb. Bull. Torrey Bot. Club 28: 278. 1901.  Roundtip Twinpod.

Clear Creek Canyon, Clear Creek County, Colorado. Rocky roadbank, 2.3 mi. east of jct US 40 & US 6. Elev. 7600 ft.

T. L. Mellichamp & Guy Nesom 11-Jun-1968

Vouchers: UNCC19620

Other articles:
• Golden Checklist Flora:  Notes on Physaria bellii, Borland, 1988;  

Jim Borland collected Physaria bellii and P. vitulifera twice: 12 May 1988 and 22 June 1988.

Collections of P. vitulifera were:

  • Verbatim Date: 12 May 1988. Locality: United States, Colorado, Jefferson, Along west Alameda Ave. (rte. 26) above Rooney Road. Reproductive Condition: Flr & Frt. https://swbiodiversity.org/seinet/collections/individual/index.php?occid=13848718
  • Verbatim Date: 12 May 1988. Locality: United States, Colorado, Jefferson, Across street from coke oven along Rooney Road. Habitat: On mineralized soil. Reproductive Condition: Flr & Frt. https://swbiodiversity.org/seinet/collections/individual/index.php?occid=13848800
  • Verbatim Date: 12 May 1988. Locality: United States, Colorado, Jefferson, Becket Street, ¼ mi. E of junction Rte. 26. Habitat: On mineralized soil (not shale). Reproductive Condition: Flr & Frt. https://swbiodiversity.org/seinet/collections/individual/index.php?occid=13848801
  • Verbatim Date: 22 June 1988. Locality: United States, Colorado, Jefferson, Becket Street, 1/4 mi. E. of junction Rte. 26 (W. Alameda Parkway) and Becket Street. Habitat: On mineralized soil (not shale). Reproductive Condition: Frt. https://swbiodiversity.org/seinet/collections/individual/index.php?occid=13849334.

Literature Cited:
- Wichmann, Brenda L., 2023.

Other articles:
• Golden Checklist Flora:  Physaria vitulifera;  

Jennifer Ackerfield, with assistance from Sami Naibauer (population genetics lab coordinator and field botanist at the University of Northern Colorado’s (UNC’s) School of Biological Sciences), updated the group on a collaborative effort to elucidate the taxonomic status of a plant referred to as Physaria ×1, a suspected hybrid of Bell’s twinpod (Physaria bellii) and fiddleleaf twinpod (Physaria vitulifera). The suspected hybrid has now been determined to be within the species concept of Physaria vitulifera.

 

Other articles:
• Golden Checklist Flora:  Rorippa palustris;  

Rorippa palustris (L.) Besser “Bog Yellow-Cress”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 657) ...

Original Text
3. SISYMBRIUM filiquis, declinatis oblongo-ovatis, foliis pinnatifidis ferratis. amphibium.
α. Sifymbrium foliis pinnatifidis ferratis. Fl. fuec..
Sifymbrium foliis infomis capillaceis, fummis pinnatifidis. Hort. cliff. 337. Roy. lugdb. 341. Dalib. parif. 203.
Raphanus aquaticus, foliis in profundas lacinias divifis. Bauh. pin. 97. prodr. 38. t. 38. Fl. lapp. 262
paluftre.
β. Sifymbrium foliis fimplicibus dentatis ferratis. Hort. cliff. 336. Fl. fuec. 350. Roy. lugdb. 341. Hall. helv. 548.
Raphanus aquaticus, rapiftri folio. Bauh. pin. 97.
Raphanus aquaticus alter. Bauh. prodr. 38. t. 38.
aquaticum.
δ. Sifymbrium aquaticum, foliis variis. Vaill. parif. 185.
Sifymbrium foliis imis integris ovatis ferratis, fuperioribus pinnatis. Hall. helv. 548.
terreftre.
Habitat in Europae feptentrionalioris aquofis.  

Literature Cited:
- Leysser, Friedrich Wilhelm von, 1761.  

Leysser (1761, p. 126) in his first edition of Flora Halensis accepted Linnaeus' treatment of R. palustris as a variety of S. amphibium.

Original Text Interpretation and Comments
603. Sisymbrium (amphibium) filiquis declinatis oblongo-ouatis, foliis pinnatifidis ferratis. Linn. fpec. 657.  
α. Sifymbrium (paluftre) foliis pinnatifidis ferratis. Linn. fuec. 2. n. 593. β. Icon. orig. cent. 3.  
β. Sifymbrium (aquaticum) foliis simplicibus dentatis ferratis. Linn. fuec. 2. n. 593. α.  
H. ad foffas circa Paffenforf, & alibi paffin: fl. Maio & feqq.  

Literature Cited:
- Leysser, Friedrich Wilhelm von, 1783.  

In his second edition of Flora Halensis, Leysser (1783, p. 166) treated our plant at the rank of species without comment regarding why he elevated it to that rank.

Original Text Interpretation and Comments
679. Sisymbrivm paluftre filiquis declinatis oblongo-ouatis, foliis pinnatifidis ferratis, (petalis calyce breuioribus.) L. f. 3. p. 250. n. 3. α. In this edition, Leysser seems to have interchanged the letters ‘u’ and ‘v.’ For example, the description of the silique as “oblongo-ouatis,” would otherwise be written as “oblongo-ovatis.”
Ad foffas, aquas, pifinas paffim, rarior tamen praecedente; Maio et feqq.  

Literature Cited:
- Besser, Wilibald S., 1822.  

Besser (1822, p. 27) writing in an enumeration of plants in a section of eastern Europe, placed our plant in Rorripa.

Original Text
CCCXII. Roripa Scop. Nasturtium H. Kew. Sisymbrium L.
824. palustris.
825. amphibia.
826. sylvestris.

 

Other articles:
• Golden Checklist Flora:  Rorippa sinuata;  

Rorippa sinuata (Nutt.) Hitchc. “Spreading Yellowcress”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.

Other articles:
• Golden Checklist Flora:  Torrey & A. Gray, 1838-1843, publication details;  

Torrey & A. Gray (1838, v. 1, p. 73) proposed Nasturtium sinuatum from a manuscript by Thomas Nuttall describing plants he had seen in Oregon Territory and in the Arkansas.

Original Text Comments
5. N. sinuatum (Nutt.! mss.): “ decumbent ; leaves pinnatifid ; segments lanceolate, subserrtat or toothed on the lower margin ; pedicels spreading or recurved, longer than the oblong acute silique ; style nearly one-third the length of the silique.  
“Banks of the Oregon and its tributaries ; also in Arkansas. — Glabrous. Leaves all rqually pinnatifid ; the terminal segments more or less confluent. Flowers rather large, bright yellow. Sepals ovate. Petals oblong-ovate. Silique about one-tird of an inch long, slightly curved.” Nutt. Cotyledons o== “… the Oregon …” is a synonym for the Columbia River.

 

Other articles:
• Golden Checklist Flora:  Rorippa tenerrima;  

Rorippa tenerrima Greene “Modoc Yellowcress”

 

Literature Cited:
- Greene, Edward L., 1895.  

Greene (1895b, v. 3, p. 46) published R. tenerrima from collections in Modoc County, California by Mrs. R. M. Austin.

Original Text
Roripa tenerrima. Annual, weak and decumbent, very sparingly branching, 6 to 10 inches high, of delicate texture and glabrous: leaves few, lyrate-pinnatifid, the terminal lobe acutish: rachis of the few racemes almost capillary: pods rather distant, subconical, slightly curved, the tapering apex surmounted by a considerable beak-like style; valves and septum both very thin: seeds many, in 2 rows under each valve.
Collected sparingly, in Modoc Co., California, 1894, by Mrs. R. M. Austin.

   

Thlaspi L. “Pennycress”

Thlaspi L. s.s. is not known from Colorado, the only representative being Thlaspi arvense L. “Field Pennycress,” an introduced species.

Literature Cited:
- Mummenhoff, Klaus, Andreas Franzke, and Marcus Koch, 1997.

Other articles:
• Golden Checklist Flora:  Notes on Noccaea fendleri, Holub, 1998;  

Mummenhoff, et al. (1997) examined phylogenetics of Thlaspi s.l. using primarily European material. The main lineages recognized are Thlaspi s.s., Noccaea (Raparia) included, and Microthlaspi. Noccaea includes by far the highest number of species formerly classified in Thlaspi s.l. It will be up to Holub to propose names for the American Noccaea.

 

Other articles:
• Golden Checklist Flora:  Turritis glabra;  

Turritis glabra L. “Tower Rockcress”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 666) ...

Original Text
glabra. 1. TURRITIS foliis radicalibus dentatis hifpidis, caulinis integerrimus amplexicaulibus glabris. Hort. cliff. 339. Fl. fuec. 544. Roy. lugdb. 339. Hall. helv. 5600. Dalib. parif. 203
  Braffica fylveftris, foliis circa radicem cichoraceis. Bauh. pin. 112.
  Habitat in Sueciae, Germaniae, Angliae pafcuis fylvaticis.

 

Other articles:
• Golden Checklist Flora:  Cleomella serrulata;  

Cleomella serrulata (Pursh) Roalson & J. C. Hall “Rocky Mountain Beeplant”

[Syn: Cleome serrulata Pursh, Peritoma serrulata (Pursh) DC.]

Literature Cited:
- Moulton, Gary E., 1999.
- Pursh, Frederick, 1814.

Locations: Vermillion River.  

Pursh (1814, v. 2, p. 441) ...

Original Text Comments
2. C. glabra ; floribus hexandris, foliis ternatis, foliolis lanceolatis tenuissime serrulatis, racemo elongato, bracteis linearibus. serrulata.  
On the banks of the Missouri. ☉. Aug. v . s. in Herb. Lewis. Flowers pale purple or white.   Mouth of the Vermillion River, Clay County, South Dakota, or Cedar or Dixon County, Nebraska, August 25, 1804 (Moulton, 1999).

Literature Cited:
- Roalson, Eric C., Jocelyn C. Hall, James P. Riser II, Warren M. Cardinal-McTeague, Theodore S. Cochrane & Kenneth J. Sytsma, 2015.  

Cleomella serrulata (Pursh) Roalson & J.C.Hall, Phytotaxa 205(3): 140 (2015).

There is really nothing specific about Cleomella serrulata in this paper that focuses on the North American cleomids, except perhaps, that it fits into alternatives of Peritoma s.l., Peritoma s.s., and Cleomella s.l., along P. jonesii and P. lutea.

 

Other articles:
• Golden Checklist Flora:  Polanisia dodecandra;  

Polanisia dodecandra (L.) DC. “Sandyseed Clammyweed”

 

 

Other articles:
• Golden Checklist Flora:  Sedum lanceolatum;  

Sedum lanceolatum Torr. “Spearleaf Stonecrop”

 

Literature Cited:
- Torrey, John G., 1828.  

172. S. lanceolatum, foliis planis, subalternis, inferioribus confertis, oblongo-lanceolatis, acutiusculis, glabris, margine glanduloso-serrulatis ; caule ramoso, assurgente ; floribus cymoso-corymbosis ; petalis lanceolatis, patentibus.
Desc. Stem a little branched at the base, creeping. Leaves about half an inch long, rather crowded, oblong-lanceolate, obscurely 3-nerved, smooth, except on the margin, which is glandularly serrate under a lens. Cyme corymbose, the branches spreading, or reflexed. Flowers white ? decandrous ? Segments of the calyx lanceolate. Petals 7, lanceolate, acute. Stamens — .
Hab. Near the Rocky Mountains.

Literature Cited:
- Goodman, George J., and Cheryl A. Lawson, 1995.  

Original Text
James could have collected the type (NY) of S. lanceolatum in early July along his route from the chasm of the Platte, Jefferson County, south to Manitou Springs and Pikes Peak, El Paso County, Colorado.

 

Other articles:
• Golden Checklist Flora:  Jamesia americana;  

Jamesia americana Torr. & A. Gray “Fivepetal Cliffbush”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1840, v. 1(4), p. 594) published Jamesia americana with a tribute to the work of Dr. Edwin James.

J. Americana.
Dr. James ! — Shrub erect? with terete branches. Leaves, including the petioles, 1-2 inches long, ovate, simply serrate with broad mucronate teeth. Cymes shorter than the leaves : bracts subulate. Calyx persistent, cleft rather below the middle ; the segments mostly acute, two of them somewhat narrower ; one of the broader ones sometimes minutely 3-toothed at the apex. Petals twice or more the length of the calyx. Stamens deciduous. Styles more than twice the length of the ovary, much exserted beyond the calyx. Ovary free, except the base, the parieties rather thick and firm ; the dissepiments very short ; the placentae lunate, at first distinct, many-ovuled. — We much regret that we have not more adequate materials for describing this plant. Our specimens were collected by Dr. Edwin James (in Long's Expedition), but the particular locality is not recorded. It is probably rare or very local, as no other botanist seems to have met with it. It appears to be an entirely distinct genus, to which we have applied the present name in commemoration of the scientific services of its worthy discoverer, the botanist and historian of ‘Major Long's Expedition to the Rocky Mountains, in the year 1820,’ and who, during that journey, made an excellent collection of plants under the most unfavorable circumstances.

 

Other articles:
• Golden Checklist Flora:  Heuchera bracteata;  

Heuchera bracteata (Torr.) Ser. “Bracted Alumroot”

 

 

Other articles:
• Golden Checklist Flora:  Heuchera parvifolia;
Full Size ImageColl. No. 1655, Heuchera parvifolia  

Heuchera parvifolia Nutt. ex Torr. & A. Gray. “Littleleaf Alumroot”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & A. Gray (1838, v. 1, p. 581) published H. parvifolia in the section Heucherella.

§ 4. Filaments and styles subulate, very short : calyx obconic at the base, rotate, equal : petals small, fugacious : flowers small : panicles narrow, loose. — Heucherella.  
13. H. parvifolia (Nutt.! mss.): scabrous-puberulent ; scape naked ; leaves forming a small radical cluster, roundish-cordate, crenately 5-7-lobed, at length glabrous, ciliate ; the lobes short and rounded ; panicle racemose, rather loose ; bracts small, laciniate-ciliate ; flowers very small ; limb of the calys flat, dilated ; petals minute, caducous ; stamens shorter than the lobes of the calyx ; styles very short, conical. Typically when quoting a Nuttall manuscript, Torrey & A. Gray will place double quotes around the description. They did not do so in this case. I assume that was a printer's oversight in this case, and is not significant.
Rocky Mountains, Dr. James! Blue Mountains of Oregon, Nuttall! — Caudex somewhat ligneous, thick. Leaves an inch or more in diameter ; the lobes with one or two crenatures, not mucronate. Scape 8-12 inches : panicle branching below. Calyx-tube obconic, adherent nearly to the summit of the ovary. Filaments very short and thick, subulate : anthers large for the size of the flower. Seeds hispid. — Mr. Nuttall supposes this species to be nearly allied to H. Richardsonii. It seems to us, however, to form, with the succeeding species, a very well-marked section. The succeeding was “14. H. hirtiflora …”

 

Other articles:
• Golden Checklist Flora:  Micranthes rhomboidea;
• Field Notes:  Coll. No. 2262, 6 May 2020;
Full Size ImageColl. No. 2262, Micranthes rhomboidea
Full Size ImageColl. No. 2262, Micranthes rhomboidea  

Micranthes rhomboidea (Greene) Small. “Diamondleaf Saxifrage”

 

Literature Cited:
- Greene, Edward L., 1898.  

Greene (1898, v. 3, n. 19, p. 343) ...

Saxifraga rhomboidea. Stem leafless and scapiform, stoutish, 5 to 10 inches high, pubescent : leaves all in a radical tuft, usually depressed and rosulate, the earliest ones rhombic-ovate, the later more oval, 1 to 2½ inches long including the broad short petiole, variously toothed, often crenate, sometimes repand-dentate, glabrous or the margins short-hairy: flowers small, white, in a commonly dense ovoid capitate-terminal cluster, but this in fruit becoming somewhat obviously branched and thyrsiform : petals white, spatulate-obovate, usually emarginate.
Common species of dry open slopes, chiefly in the belt of Pinus ponderosa, in the Colorado Rocky Mountains, thence northward to Montana and southward to New Mexico; long referred to the Old World S. nivalis, a very different alpine or subarctic species. Mr. Holm, who has independently made a careful anatomical study of S. nivalis and its allies, informs me that this Rocky Mountain plant bears internal evidence of a closer relationship to S. integrifolia by far, than to S. nivalis. S. rhomboidea has an American but high-northern ally in S. radulina of page 308 preceding.

Literature Cited:
- New York Botanical Garden, 1905-1949.  

Small (1905, v. 22, pt. 2, p. 136) ...

9. Micranthes rhomboidea (Greene) Small.
Saxifraga nivalis Hook. Fl. Bor. Am. 1 : 248, in part. 1832. Not S. nivalis L.
Saxifraga rhomboidea Greene, Pittonia3: 343. 1898.
? Saxifraga rhomboidea austrina A. Nelson, Bot. Gaz. 31 : 394. 1901.
Leaves spreading or ascending, 2-6 cm. long, the blades mainly ovoid or rhombic- ovoid, sometimes oblong-ovoid, obtuse, crenate or dentate-serrate, nearly glabrous except the more or less ciliate margins, mainly green, sometimes purplish beneath, abruptly or gradually narrowed into petiole-like bases which are occasionally longer than the blade itself; scapes mainly erect, 8-31 cm. high, green, copiously glandular-pubescent, simple or Sparingly paniculate above ; cymules aggregated into a terminal head or in several heads terminating the short branches of the inflorescence; bracts subtending the peduncles narrow, green; sepals ovate to triangular, 1.5-3.5 mm. long, barely if at all ciliate, 3- veined; petals white, obovate to oblong-obovate, 2.5-4 mm. long, longer than the sepals, commonly notched at the apex, often contracted into broad claw-like bases ; filaments subulate ; follicles 3-3.5 mm. high, green or purple-tinged, the stout tips widely spreading.
Type locality : Mountains of Colorado.
Distribution: Montana, Wyoming and Colorado.

 

Other articles:
• Golden Checklist Flora:  Ribes aureum;  

Ribes aureum Pursh “Golden Currant”

 

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, p. 164) published Ribes aureum from collections by Lewis & Clark, and from garden-grown specimens he had seen.

Original Text
aureum. 8. R. inerme, glaberrimum ; foliis trilobis : lobis divaricatis inciso-pauci-dentatis petiolo basi ciliato brevioribus, racemis laxis dense-mujiifloris, calycibus tubulatis pedicellis longioribus : tubo gracili : laciniis oblongis obtusis, petalis linearibus laciniis calycis duplo brevioribus, bracteis linearibus longitudine pedicellorum, baccis glabris.
  On the banks of the rivers Missouri and Columbia M. Lewis. ♄ April, v. s. in Herb. Lewis. ; v. v. in Hort. Flowers in close racemes, beautiful golden-yellow ; berries red or brown, of an exquisitely fine taste, and considerably larger size than any of garden currants. The shrub before flowering has the appearance of a species of Crataegus.

 

Other articles:
• Golden Checklist Flora:  Ribes cereum;  

Ribes cereum Douglas “Wax Currant”

 

Literature Cited:
- Douglas, David, 1830.  

Original Text
I brought seeds of this species to England in October 1827. The plants flowered last April, in the Garden of the Society, but rather weakly, being only one year old.*
3. R. cereum : inerme, foliis subrotundis obtuse trilobis crenatis viscidis, racemis 3-5-floris pendulis pubescentibus longitudine foliorum, calycibus tubulatis laciniis ovatis reflexis : petala subreniformia duplo excedentibus, bracteis cuneiformibus apice dentatis, baccis rubris glabris.
This bush is of more humble stature and slender habit than the one last mentioned, growing erect, about five or six feet high, with white smooth bark on the old branches. The young shoots which are curved and flexible, are covered with a brown viscid scentless glutinous substance, which when exposed to the sun, acquires a rough, hardened, waxy, warty appearance. The leaves are nearly round, bluntly three-lobed, crenate, scarcely an inch long, of a leathery texture and almost veinless, clothed on the upper surface with white and (in dry weather) hardened waxy minute granulations, quite smooth below ; footstalks somewhat longer than the leaves. The clusters are dense, of the same length as the leaves, three or five-flowered, slightly pubescent, hanging in great profusion below the branches, with scarcely any partial footstalks ; bracteas wedge-shaped, glandular and toothed at the apex. The calyx is tubular, imperfectly four-sided, white, pink at the base, three-fourths of an inch long, with rounded, short, reflected segments double the length of the minute somewhat kidney shaped petals. Filaments, same length as the petals ; style slightly cloven. Berry spherical, small, red and glossy, thin-skinned, rarely containing more than three large seeds and a great quantity of insipid, viscid, red juice.
This species in point of beauty cannot be compared to the fragrant flowered R. aureum, and its varieties, nor can it vie with the gaudy inflorescence of R. sanguineum. It blossoms at the same season and with equal profusion, is equally hardy and as readily cultivated.
On dry exposed decayed granite rocks or schist, throughout the chain of the river Columbia from the Great Falls 45° 46' 17" N. Lat. to the source of that stream in the Rocky Mountains, 52° 07' 09" this is a common shrub, flowering in March and April, and ripening its fruit in June.

 

Other articles:
• Golden Checklist Flora:  Saskatoon Serviceberry;  

Amelanchier alnifolia (Nutt.) Nutt. ex M. Roem. “Saskatoon Serviceberry”

 

Literature Cited:
- Moulton, Gary E., 1999.  

Collected by Lewis & Clark on April 15, 1806 at The Dalles of the Columbia River. However, it was not published by Pursh.

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Nuttall, 1818, publication details;
Full Size ImageNuttall (1818) description of Aronia alnifolia.  

Order IV. — Pentagynia

340. Aronia. Persoon. Mespilus. L.

Calix 5-toothed. Petals 5. Berry inferior 5 to 10-celled; cells 1 or 2-seeded. Seeds cartilaginous.

Shrubs without spines, having alternate undivided leaves, and flowers which are corymbose or racemose, generally white; fruit a small black purple or scarlet pomois berry, containing seeds similar to those of apples.

Species. 1. … 6. * Alnifolia. Smooth: leaves roundish, upperpart toothed, pinnately nerved, under side somewhat glaucous; raceme simple, elongated; fruit black and sweet. Habitat. In ravines and on the elevated margins of small streams from Fort Mandan to the Northern Andes. Observation. A shrub 4 or 5 feet high; leaves roundish and retuse, somewhat attenuated at the base, toothed towards the summit; fruit dark purple, somewhat pruinose, very agreeable and saccharine; ripening about July and August.

Literature Cited:
- Nuttall, Thomas, 1834a.
Full Size ImageNuttall (1834) use of the name Amelanchier alnifolia  

In 1834, Nuttall referred to Amelanchier alnifolia in his review of plants collected by Nathaniel Wyeth:

37. Amelanchier alnifolia. Source of the Missouri.

Unfortunately this possible publication of a new name for the taxon is invalid because there is no reference (either direct or indirect) to the intended basionym Aronia alnifolia Nutt. (1818).

Literature Cited:
- Roemer, M. J., 1847.
Full Size ImageRoemer (1847) description of A. alnifolia  

Roemer validly published Amelanchier alnifolia in 1847, referring to Nuttall's 1834 use of the name.

3) prope apicem tantum serrata, subrotunda v. late elliptica, utrinque obtusissima v. retusa ; racemi densiflori ; petala lineari-oblonga, calyce 3–4plo longiora; stamina brevissima.

18. A. alnifolia Nutt. In Joun. Acad. Philad. VII. 22

A. ovalis β semiintegrifolia Hook. L. c. 201. – G.Don l. c.

A. canadensis δ alnifolia Torr. & A.Gr. l. c.

Aronia alnifolia Nutt. gen. I. 306

Pyrus alnifolia Ser. l. c. 637. 39.G.Don l. c. 649. 56.

Ad flum. Columbia pr. Fort Vancouver et ad “the grand Rapids,” in editoribus ad flumen Multnomah. 5 5.

 

There is one collection of A. utahensis made June 1, 1913 on Lookout Mountain. The name of the collector is unknown. This is one of 13 collections made that day on Lookout Mountain or at its base. The Lookout Mountain Road was under construction on that day, and opened to public use August 21, 1913, but not completed until December 1913 (Colorado Transcript, multiple dates).

 

J. H. Ehlers collected A. utahensis on a rocky slope of a hogback near Golden (COLO662346). There are three possible hog backs which may be this location. There is Dakota Ridge (North Hogback) at the very north of Golden. There is a small hogback informally named Eagle Ridge near the intersection of US Highway 6 and Heritage Road. Finally, at the very south end of Golden is Tin Cup Ridge, the northward extension of Dinosaur Ridge into Golden. There are three other vouchers of Ehlers collections made the same day, though none of them give any more details on the actual location.

Literature Cited:
- Jones, George Neville, 1946.  

 

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Some authors, such as Ackerfield (2015), treat A. utahensis as a variety of A. alnifolia, explaining that the two taxa overlap in morphology and distribution, and that it can be difficult, if not impossible, to assign one name or the other to some specimens. Then we would have:

  • Amelanchier alnifolia (Nutt.) Nutt. ex M. Roem. var. alnifolia “Saskatoon Serviceberry” and
  • Amelanchier alnifolia (Nutt.) Nutt. Ex M. Roem. Var. utahensis (Koehne) M. E. Jones “Utah Serviceberry”

 

Other articles:
• Golden Checklist Flora:  Amelanchier utahensis;  

Amelanchier utahensis Koehne “Utah Service-Berry”

 

 

Other articles:
• Golden Checklist Flora:  Cercocarpus montanus;
• Field Notes:  Coll. No. 2270.1, 9 May 2020;
Full Size ImageMountain Mahogany (Cercocarpus montanus) on South Table Mountain.
Full Size ImageColl. No. 2184, Cercocarpus montanus in fruit.  

Cercocarpus montanus Raf. “Alder-Leaf Mountain Mahogany”

Full Size Image
Coll. No. 2270.1, Cercocarpus montanus in bloom.
 

 

Other articles:
• Golden Checklist Flora:  Cotoneaster lucidus;  

Cotoneaster lucidus Schltdl. “Hairy Stem Cotoneaster”

 

 

Other articles:
• Golden Checklist Flora:  Crataegus erythropoda;

Locations: Magpie Gulch.
Full Size ImageColl. No. 2930, Crataegus erythropoda.  

Crataegus erythropoda Ashe “Cerro Hawthorn”

 

 

Other articles:
• Golden Checklist Flora:  Crataegus succulenta;  

Crataegus succulenta Schrad. ex Link “Fleshy Hawthorn”

(Syn: Crataegus chrysocarpa Ashe, Crataegus erythropoda Ashe, Crataegus macrantha Britton var. occidentalis (Britton) Eggl.)

 

Other articles:
• Golden Checklist Flora:  Fragaria virginiana;  

Fragaria virginiana Mill. “Virginia Strawberry”

 

Literature Cited:
- Miller, Philip, 1768.  

Miller (8th ed., 1768, p. FOEN-FRA) ...

2. Fragaria (Virginiana) foliis oblongo-ovatis ferratis, inferne incanis, calycibus longioribus, fructu fubrotundo. Strawberry with oblong, oval, fawed leaved, hoary ontheir under fide, longer empalements,and a roundifh fruit. Fragaria Virginia fructu Coccineo. Hift. Ox. 2. 186. Virginia Strawberry with a fcarlet fruit, commonly called the Scarlet Strawberry.

Hift. Ox. Rob. Morifoni Hiftoria Plantarum Univerfalis Oxonienfis. Pars ada, Oxoniae, 1680; Pars 3, abfoluta a Bobarto ; Oxon. 1699, fol.

 

Other articles:
• Golden Checklist Flora:  Holodiscus dumosus;  

Holodiscus dumosus (Nutt. ex Torr. & A.Gray) A.Heller. “Rock Spirea”

 

Literature Cited:
- Moulton, Gary E., 1999.  

There are two collections of Holodiscus discolor in the Lewis & Clark herbarium both collected May 29, 1806 on the Clearwater (Kooskooske) River, Camp Chopunnish, Kamiah, Idaho County, Idaho (Moulton, 1999).

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 1, p. 341-342) ...

Original Text

413. SPIRÆA. Gen. pl.862.

* Fruticosæ

...  
discolor 8. S. foliis ovatis lobatis dentatis subplicatis subtus niveotomentosis, paniculatis terminalibus pedunculatis ramosissimis.

On the banks of the Kooskoosky. ♄. June, July. v. s. in Herb. Lewis. A shrub about five feet high.

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.

Other articles:
• Golden Checklist Flora:  Torrey & A. Gray, 1838-1843, publication details;  

Torrey & A. Gray (1838, v. 1, p. 416) ...

Original Text
5. SPIRÆA. Linn. ; Gærth. fr. t. 69 ; DC. prodr. 2 p. 541.
...
§ 2. Flowers perfect : disk free at the margin, mostly crenate or with glandular teeth or lobes : carpels distinct, not inflated : ovules mostly several, pendulous : seeds mostly with a loose membranous testa, attenuate at each end : shrubs with entire or serrate exstipulate leaves. — Euspiraea.
...
* * * * Flowers in large and loose compound panicles : calyx deeply 5-cleft, spreading : disk sholly coherent, entire ; ovules 2, collateral !
8. S. ariæfolia (Smith): leaves broadly ovate, petioled, obtuse, pinnatifidly incised or somewhat lobed, dentate with mucronate teeth, almost glabrous above, canescently hairy or tomentose beneath ; panicle large and loose, much branched, and, with the calyx, tomentose-pubescent ; segments of the calyx acute, apreading ; carpels 5. broad, compressed, margined, very hirsute. — Smith ! in Rees, cycl. ; Seringe in DC. l.c. ; Lindl.! bot. reg. t. 1365 ; Hook.! fl. Bor-Am. 1 p. 173 ; & bot. Beechey, suppl. p. 338.
β discolor : leaves much smaller, cuneiform at the base, the serratures scarcely mucronate, silvery-tomentose beneath. — S. discolor, Pursh. ! fl. 1. p.342 ; Seringe, l. c. ; Torr. ! in ann. lyc. New York, 2. p. 195. S. dumosa, Nutt. ! mss.
N. W. Coast ! Oregon ! and California ! (Menzies ! Douglas ! Dr. Scouler ! Nuttall!) β. In the Rocky Mountains, Dr. James ! Nuttall ! and on the Kooskoosky River, Lewis ! June-July. — The ordinary forms is frequently somewhat arborescent, according to Nuttall, with a stout trunk, rising to the height of 12-14 feet. His S. dumosa is said to tbe a low shrub, confined to mountain regions. Douglas's Californian specimens are quite intermediate between the two, and probably came from the back country : the young leaves agree well with the description of Pursh.

Literature Cited:
- Geyer, Charles A., 1846.
- Hooker, Sir William J., 1847.  

Hooker (1847, v. 6., p. 217) published Nuttall's H. dumosa in his Catalogue of Geyer's collections of plants in the upper Missouri, Oregon Territory, and the intervening portion of the Rocky Mountains. IPNI judges this name as illegitimate.

Original Text
4. S. dumosa, Nutt. — S. discolor, Ph. (fide Torr. et Gr.) — S. ariæfolia, β. discolor, Torr. et Gr. Am. 1. p. 416.
Hab. Stony and sandy places of Platte River; a shrub, from 2-10 feet high : and at the mouth of Walla-Walla River, Upper Oregon. June. (n. 228.) — These are specimens of a dwarf shrub, with leaves smaller than those of a gooseberry, and the foliage and panicles very different from those of S. ariæfolia, with which Torrey and Gray unite it. Nuttall's, Geyer's and Gordon's specimens (the latter from the Upper Platte) are uniform.
Geyer (1846) published an account of his travels in Hooker's London Journal of Botany.

Literature Cited:
- Gray, Asa, 1849.

Other articles:
• Golden Checklist Flora:  Gray, 1849, publication details;  

A. Gray (1849, p. 40) in Planteæ Fendlerianæ ...

Original Text
ROSACEAE
...
188. S. dumosa, Nutt ! Mss. ; Hook. Lond. Jour. Bot. 6. p. 217. Steep mountain-sides, upper part of Santa Fé Creek ; July. Shrub about 4 feet high. — This is the S. discolor of Torrey in Ann. Lyc. New York (James's Collection), and a good species ; but the original S. discolor, Pursh, I believe to be only S. ariæfolia ; as certainly is a plant of Menzies so named by Pursh in Herb. Lambert.

Remarks in the International Plant Names Index

A. Gray used: S. Dumosa, Nutt! Mss.; Hook. Land. Jour. Bot. 6. p. 217. … This is the S. discolor of Torrey in Ann. Lye. New York (James's Collection), and a good species; but the original S. discolor, Pursh, I believe to be only S. ariaefolia; as certainly is a plant of Menzies so named by Pursh in Herb. Lambert. S. dumosa Nutt. ex Hook. (1847) is an illegitimate superfluous name for S. discolor Pursh (1813). Since A. Gray used S. dumosa Nutt. ex Hook., but excluded its illegitimacy causing synonymy (i.e., S. discolor Pursh), he created a later homonym (see Vienna Code Art. 48.1).

Literature Cited:
- Heller, A. A., 1898.

Other articles:
• Golden Checklist Flora:  Heller, 1898;  

Heller (1898, p. 4) published Holodiscus dumosa with Spiraea dumosa as the basionym.

Original Text
Holodiscus dumosa (Nutt.)
   Spiraea dumosa Nutt.: T. & G. Fl. N. A. 1: 416. 1840. as synonym.

Literature Cited:
- Harrington, H. D., 1964, 2nd ed..  

Harrington (1964, 2nd ed., p. 309) accepts Holodiscus microphyllus Rydb. and H. dumosus (Nutt.) Heller.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012, p. 338) accept only Holodiscus discolor (Pursh) Maximovicz and include H. dumosus and H. microphyllus therein.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015) accepts only Holodiscus dumosus (Nutt. ex Hook.) A. Heller.

Literature Cited:
- Lis, Richard, 2015.  

Lis (2015) accepts the following names in Holodiscus ...

  • Holodiscus discolor (Pursh) Maximowicz
    • Holodiscus discolor var. discolor
    • Holodiscus discolor var. dumosus (S. Watson) Maximowicz ex J. M. Coulter
  • Holodiscus microphyllus Rydberg
    • Holodiscus microphyllus var. glabrescens (Greenman) F. A. Ley
    • Holodiscus microphyllus var. microphyllus
    • Holodiscus microphyllus var. sericeus F. A. Ley

 

Other articles:
• Golden Checklist Flora:  Malus pumila;  

Malus pumila Mill. “Common Apple; Crabapple”

 

   

Physocarpus Maxim.

 

Literature Cited:
- Maximowicz, C. J., 1879.  

Maximowicz (1879, p. 219) is credited with publishing Physocarpus, though he credits "Camb." for the name, and notes Neilliae sectio Physocarpus Hook. F. Gen. pl. I 612. Names published were Ph. opulifolia L. Cod. 3724. (sub Spiraea), Ph. Torrey Wats., and Ph. Amurensis Maxim.

 

Other articles:
• Golden Checklist Flora:  Physocarpus monogynus;  

Physocarpus monogynus (Torr.) J.M. Coult. “Mountain Ninebark”

 

Literature Cited:
- Torrey, John G., 1828.  

Torrey (1828, p. 194) describes ...

Original Text
119. S. monogyna, foliis lato-ovatis, subtrilobis, inciso-serratis, glabris ; floribus corymboso-umbellatis, monogynis ; pedicellis glabris ; calycis lobis erecto-patentibus, ovariis villosis.
Desc. Fruticose. Branches alternate, covered with a loose bark as in S. opulifolia. Leaves roundish-ovate, broad and subcordate at the base, somewhat 3-lobed, incisely toothed, slightly pubescent beneath, smooth above ; petioles without stipules. Umbels corymbose, few-flowered, on short peduncles ; pedicels filiform, about half an inch long. Flowers perfect. Calyx campanulate, 5-lobed ; lobes ovate-obtuse, somewhat spreading, pubescent. Petals —. Stamens 20 ; filaments inserted on a torus, which is free at the margin. Germen solitary, stipulate, ovate, compressed, villous, 3-seeded, acuminate with the persistent filiform style ; stigma small, capitate.
Hab. On the Rocky Mountains.
Obs. This species, so far as I have examined it, is constantly monogynous. In many respects if resembles the genus Neillia of Don prod. fl. Nep.

Literature Cited:
- Maximowicz, C. J., 1879.  

Maximowicz (1879, p. 219) ... chooses to use Watson's name over Torrey's basionym.

2 Ph. Torreyi Wats. in Proceed. Amer. Acad. New ser. XI. 136. (sub Neillia). Foliis parvulis obtuse trilobis; corymbis plurifloris; staminibus 30 petala glabra, carpellis 1-2 stellatotomentosis calycem vix superantibus. Spiraea monogyna Torr. in Ann. Lyc. N. Y. II. 194. S. opulifolia var. γ. pauciflora Torr. et Gray l. c.
Hab in America boreali occidentali : montibus Colorado et occidentem versus ad Sierram Nevadam, ex Watson. Vidi specc. Fendler pl. Neo-Mexic. n. 187. et Parry pl. Colorado .207
Staminum circiter 30 series 3 : 10 per paria sepalis, 10 paulo breviora petalis et sepalis opposita, cum praecedentibus igitur alterna, 40 intima omnium breviora cum serie secunda alternantia. Ovula suturae ventrali prope apicem inserta, summa horizontalis, infima pendula.

Literature Cited:
- Coulter, John M., 1891.  

John M. Coulter (1891, p. 104) published P. monogyna from the Guadalupe Mountains of west Texas.

Original Text
4. PHYSOCARPUS Maxim.
Diffuse shrubs, with palmately lobed leaves, corymbose flowers, and 1 to 5 divergent inflated membranaceous dehiscent 2 to several-seeded carpels.
1. P. monogyna. A small shrub : leaves ovate or often cordate, 3-lobed and toothed, sometimes densely white-tomentose beneath : flowers on short pedicels in simple umbel-like corymbs : ovaries densely tomentose and but 1 or 2. (Spiraea monogyna Torr. Neillia Torreyi Watson. Physocarpus Torreyi Maxim.) — In the Guadalupe Mountains.

Literature Cited:
- Oh, Sang-Hun, and Daniel Potter, 2005.  

“ … in Physocarpus one dispersal event from western North America to eastern Asia occurred.”

 

Other articles:
• Golden Checklist Flora:  Potentilla fissa;  

Potentilla fissa Nutt. ex Torr. & A.Gray. “Bigflower Cinquefoil”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & A. Gray (1838, v. 1, 446) published Potentilla fissa from Nuttall and Wyeth collections on the plains of the Rocky Mountains towards Oregon.

Original Text
34. P. fissa (Nutt.! mss.): “viscidly pubescent ; stem erect, branching, leafy ; leaves pinnately 9-11-foliolate, on shory petioles ; leaflets unequal, roundish or oval, deeply incised or incisely toothed, the teeth entire ; stipules entire or toothed ; flowers rather crowded ; segments of the calys ovate, acute, shorter than the roundish (sulphur-yellow) petals.”
β. major : larger in all its parts ; flowers more crowded. — P. arguta, Nutt.! in jour. acad. Philad. 7. p. 21, not of Pursh. P. glutinosa, Nutt.! July.
Plains of the Rocky Mountains towards the Oregon, Nuttall! July. β Headwaters of the Oregon, Capt. Wyeth! — “Stem about a span high. Leaflets small, the lower ones roundish ; those of the upper cauline leaves ovate. External sepals much smaller, often toothed. Flowers rather large.” Nutt. — The plant which we have joined as a variety of this species seems to be a larger plant, and bears more resemblance to P. arguta.

 

Other articles:
• Golden Checklist Flora:  Potentilla gracilis;  

Potentilla gracilis Hook. “Slender Cinquefoil”

 

Literature Cited:
- Hooker, William J., 1830a.
Full Size ImageHooker's Plate 2984 of Potentilla gracilis  

Hooker (1830, v. 57, t. 2984) ...

( 2984 )
POTENTILLA GRACILIS. TALL UPRIGHT ClNQUEFOIL.
*********************
Class and Order.
Icosandria Polygynia.
( Nat. Ord. — Rosaceae. )
Generic Character.
Cal. 10-fidus, laciniis alternis minoribus. Pet. 5. Caryopses nudae, plurimae, in receptaculo sicco, saepe piloso. Spr.
Specific Character and Synonym.
Potentilla gracilis ; caule erecto elato superne corymboso-paniculato molliter hirsuto, foliis longe petiolatis 5-natis superioribus solummodo sessilibus, foliolis lanceolatis profunde pinnatifido-serratis subtus albo-tomentosis, stipulis magnis lanceolatis, calyce senceo petalis obcordatis longiore.
Potentilla gracilis. Douglas MSS.

Descr. Root perennial, with many brown scales at its summit, from which arises a stem, a foot to a foot and a half high, rounded and slender in the wild specimens, more robust when cultivated, every where clothed with rather long, more or less patent, soft and silky hairs, pamculato- corymbose upwards. Leaves few upon the stem, the upper ones sessile and ternate, the uppermost ones laciniated and bracteiform, the rest, and especially the radical ones, upon long hairy stalks, quinate : the leaflets three to tour inches long, cut into very deep pinnatifid segments or serratures, pointing upwards, varying somewhat in breadth, dark green above with a few scattered hairs, beneath clothed with a dense, white, somewhat downy substance, mixed with silky hairs. The stipules are particularly large, lanceolate, acuminate, hairy, especially at the margin and beneath, where they are whitish. Flowers in the wild state almost corymbose, in the cultivated plant paniculated, but still nearly level-topped, peduncles forked, with generally a single flower between the forks. Cal. of five broadly-lanceolate, and five alternate, smaller, almost subulate segments, all of them clothed with rather long and silky hairs. Petals broadly obcordate, wavy, longer than the calyx.
This is a handsome, and, in our gardens, a strong-growing species of the Genus, but much better deserving the name of gracilis, as seen on the banks of the Columbia and the plains of the Multnomah rivers, where it was discovered by Mr. Douglas, than as it appears after cultivation. I cannot refer it to any described species, nor does it appear to have been gathered by any other Naturalist, except by Dr. Scouler, who accompanied Mr. Douglas as far as Fort Vancouver ; unless it be the same as a Rocky Mountain species, collected by Mr. Drummond, as I suspect it will prove to be.
The seeds were introduced to the Horticultural Society, and the plants, from which the annexed figure was taken, flowered there in the month of July, 1829.
Fig. 1. A Panicle, with the upper part of the Stem. 2. An upper Stem-leaf. 3. A radical Leaf, nat. size. 4. Petal, and 5. Calyx. — Slightly magnified.

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1838-1840, v. 1, p. 440) ...

13. P. fastigiata (Nutt. ! mss.) : “canescently silky-tomentose ; stem erect, leafy ; leaves palmately 5-7-foliolate ; leaflets cuneate-oblong, incisely or pinnatifid-serrate ; stipules mostly entire ; flowers crowded, fastigiate ; segments of the calyx lanceolate, the outer ones much smaller ; petals obovate, a little longer than the calyx ; achenia smooth.
&beta. “larger, more densely clothed with soft silky hairs.
“Plains of the Rocky Mountains. — Plant 7-8 inches high. Flowers much smaller than in P. rigida.” Nuttall.

Literature Cited:
- Watson, Sereno, 1873.  

Watson (1873, v. 8, p. 557) ...

15. P. gracilis, Dougl. (P. Blaschkeana, Turcz.) Villous and more or less tomentose ; stems 2-3° high ; stipules ovate or lanceolate, entire or subincised ; leaflets mostly 7, sometimes 5, very rarely but 3, cuneate-oblong, obtuse, incisely serrate or pinnatifid, tomentose beneath, green above and subvillous or appressed-silky, 1 - 2½' long ; flowers in a loose subfastigiate cyme, the pedicels at length elongated and slender ; calyx with the narrow bractlets shorter than the broad acute or lanceolate sepals ; petals broadly obcordate, 3 - 4" long, a little exceeding the calyx; carpels very numerous (40 or more). — From the Saskatchewan to Southern Alaska, and southward to New Mexico, Utah, and California. The leaflets occasionally show a tendency to a pinnate arrangement, and the species is distinguishable from the digitate form of P. Hippiana, var. pulcherrima, only by the more numerous carpels and the usually fewer and more deeply incised leaflets. Specimens like 159 Hall & Harbour tend to unite the two species.
...
Var. fastigiata (P. fastigiata, Nutt., P. olopetala, Lehm.) is a form, often low, with a shorter and more crowded cyme, the pubescence more dense and silky, especially upon the calyx and short pedicels.

 

Other articles:
• Golden Checklist Flora:  Potentilla norvegica;  

Potentilla norvegica L. “Norwegian Cinquefoil”

 

 

Other articles:
• Golden Checklist Flora:  Potentilla pensylvanica;  

Potentilla pensylvanica L. “Prairie Cinquefoil”

 

 

Other articles:
• Golden Checklist Flora:  Potentilla recta;  

Potentilla recta L. “Sulphur Cinquefoil”

 

 

Other articles:
• Golden Checklist Flora:  Potentilla rivalis;  

Potentilla rivalis Nutt. “Brook Cinquefoil”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.
Full Size ImageColl. No. 1682.1, Potentilla rivalis
Full Size ImageTernate leaf of Coll. No. 1682.1, Potentilla rivalis  

Torrey & A. Gray (1838, v. 1, p. 437) published P. rivalis from a Nuttall manuscript.

Original Text
2. P. rivalis (Nutt.! mss.) : “clothed with soft somewhat viscous pubescence ; stem erect, much branched ; radical leaves pinnately 5-foliate ; the leaflets crowded, and the 3 upper ones confluent ; those of the cauline leaves 3, often confluent, oblong, cuneiform at the base, coarsely serrate ; stipules ovate, nearly entire ; flowers numerous, small, on rather short pedicels ; calyx-segments acute ; petals inconspicuous; achenia smooth and even.
“In alluvial soil along the Lewis River.&rdquo July. — A very distinct species, allied to P. Norvegica. Cauline leaves small. Flowers inconspicuous.

 

Other articles:
• Golden Checklist Flora:  Prunus americana;
• Field Notes:  Coll. No. 2443, 4 Sep 2020;
Full Size ImageColl. No. 2443, Prunus americana
Full Size ImageColl. No. 2443, Prunus americana  

Prunus americana Marshall. “American Plum”

Full Size Image
Coll. No. 2443, Prunus americana
 

Literature Cited:
- Marshall, Humphry, 1785.  

Marshall (1785, p. 111) ...

1. Prunus americana. Large Yellow Sweet Plumb.
This generally rifes to the height of twelve or fifteen feet, fpreading into many ftiff branches. The leaves are oblong, oval, acute pointed, fharpiy fawed on their edges and much veined. The flowers generally come out very thick round the branches, often upon thick fhort fpurs; and are fucceeded by large oval fruit, with a fweet fucculent pulp. We have a great variety of thefe, growing naturally in a good, moift foil, with reddifh and yellowifhi fruit, but differing much in fize, tafte, and confiftence.

 

Other articles:
• Golden Checklist Flora:  Prunus domestica;  

Prunus domestica L. “European Plum”

 

 

Other articles:
• Golden Checklist Flora:  Prunus pensylvanica;  

Prunus pensylvanica L. f. “Pin Cherry”

(Syn: Cerasus pensylvanica (L. f.) Loiseleur)

Literature Cited:
- Linne´, Carl von, 1782.  

Linnaeus (1782, p.252-253) ...

Original Text
Penfylva-
nica.
PRUNUS umbellis feffilibus, foliis ovato-lanceolatis, ramulis pallide punctatis.
  Habitat in America feptentrionali. ♄. H. U.
  Facies P. Cerali, fed ramuli punctis albidis adfperli, up in P. Virginiana.
  Folia P. Cerafi, lanceolato-ovata, ferrulata, laevia, bafi glandulus duabus, faepe rubris.
  Umbellae feffiles.
  Flores P. Cerafi minores.

 

Other articles:
• Golden Checklist Flora:  Prunus virginiana;  

Prunus virginiana L. “Chokecherry”

 

 

Other articles:
• Golden Checklist Flora:  Rosa acicularis ssp. sayi;  

Rosa acicularis Lindl. ssp. sayi (Schwein.) W. H. Lewis “Say's Acicular Rose”

(Syn: Rosa acicularis Lindl., Rosa sayi Schweinitz )

Literature Cited:
- Schweinitz, Lewis D., 1824.
- Schweinitz, Lewis D. de, 1824.  

Schweinitz (1824, pp. 388-389) ...

50. Rosa *Sayi, L. v. Schw.
This appears to me a Rose quite distinct from any American one, although it is past flowering; the germen being manifestly not globose, (which is the case with all the rest except laevigata,) nor do I find any European one sufficiently agreeing. I describe it thus:
Germen oblong ovate, perfectly smooth, and proportionably large, crowned by erect calyx leaves, exceeding it in length, which are villous, and expand at summit. Peduncle smooth, or somewhat glandularly hispid, rigid. Common petiole villous and aculeate on the back, with three pairs of ovate, sessile, deeply serrate, small leaflets, and a single one on the lengthened petiole, furnished at base with clasping, glandulosely villous stipules. Upper side of the leaflets smooth, the underside glaucously villous. The young branches thickly set with thin, unequal, hispid spines.

Literature Cited:
- Lewis, Walter H., 1959.  

Rosa acicularis subsp. sayi (Schwein.) W.H.Lewis, Brittonia 11(1): 19 (1959).

 

Other articles:
• Golden Checklist Flora:  Rosa arkansana;  

Rosa arkansana Porter “Prairie Rose”

 

Literature Cited:
- Porter, Thomas C., and John M. Coulter, 1874.  

Porter & Coulter (1874, p. 38) ...

Rosa Arkansana, Porter (n. sp.) Stem stout, erect, leafy, 1° high, glabrous and glaucous, armed with weak, deciduous, bristly prickles ; leaflets 9-11, ovate and oblong-ovate, 1' or more in length, acute or obtuse, glabrous, sharply serrate; midrib and long stiipules somewhat prickly and minutely glandular; flowers numerous, termmal, corymbed, on peduncles about 1' long; fruit globose, smooth, glaucous; calyx-segments ovate, reflexed in fruit, with terminal and sometimes lateral appendages, more or less glandular and tomentose pubescent on the margins; petals broadly obcordate or emarginate, longer than the calyx-segments, rose-color; flowers 2' in diameter. — This rose may possibly be an extreme form of R. blanda, but it differs in so many points that I have ventured to describe it as new. — Banks of the Arkansas near Cañon City, Brandegee. Raton Mountains, Dr. Bell. Texas, Wright.

 

Other articles:
• Golden Checklist Flora:  Rosa woodsii;  

Rosa woodsii Lindl. “Mountain Rose”

 

Literature Cited:
- Lindley, John, 1820.  

Lindley (1820, pp. 21-22) ...

14. ROSA Woodsii.
R. stipulis sepalisque conniventibus, foliolis oblongis obtusis glabris.

R. lutea nigra Promv. nomencl. 24.

Hab. juxta flumen Missouri Americae septentrionalis (v. v. c hort. Sabine.)

In honorem cel. Josephi Woods qui primus veris Rosarum characteribus ad species distinguendas usus est.

A low shrub with upright, dull, dark branches, having very numerous, straight, slender, scattered prickles, with a few setae at their base, the former becoming stipulary towards the extremities ; branchlets often unarmed. Leaves without pubescence ; stipules very narrow and acute, convolute and fringed with glands ; stalks armed with straight unequal prickles ; leaflets 7-9, shaped like those of R. rubella, shining, flat, simply serrated, paler beneath. Flowers pink, appearing in the spring. Fruit naked, ovate, with short, connivent, entire sepals which are free from glands as is the peduncle.
As it is scarcely probable that any new British rose will be detected, worthy of bearing the name of Mr. Woods, of whose high merit I have already had occasion to speak, the present species has been selected by Mr. Sabine and myself for that purpose. That it is essentially distinct from every other is very evident even from the incomplete account I have been able to give of it. I first saw it growing in Mr. Sabine's garden at North Mimms late in the month of November ; most of the leaves had fallen, but a few heps still remained on the bush. Its habit without foliage bears more resemblance to that of a stunted cinnamomea than to any thing else. In character it approaches R. carolina, particularly in the remarkable convolution of stipulae. From this its numerous ramifications, weak prickles and short shining leaves sufficiently distinguish it. It moreover flowers in the spring and has naked fruit with conniving sepals.
I am assured by Mr. Sabine that this is the plant which was sent to France from a nursery here as a new American Rose with black and yellow flowers, and noticed as such in Promvillie's book.
Said to be a native of the country near the Missouri.

 

Other articles:
• Golden Checklist Flora:  Rubus deliciosus;  

Rubus deliciosus Torr. “Delicious Raspberry”

(Syn: Oreobatus deliciosus Torr. (E. James ex Torr.) Rydb.)

Literature Cited:
- Torrey, John G., 1828.  

Torrey (1828, v. 2, p. 196) ...

125. R. deliciosus, caule fruticoso, ramosissimo, inermi; ramis petiolisque pubescentibus; foliis simplicibus, rotundato-cordatis, breve 3-5-lobis, villoso-pubescentibus, rugosis; bracteis lanceolatis, unidentatis ; floribus subcorymbosis, terminalibus ; laciniis calycinis ovato-oblongis, acuminatis, apice foliaceis, petalis brevioribus.
Desc. Shrub much branched ; branches flexuous, terete, pubescent. Leaves suborbicular-cordate, 2-2½ inches in diameter, villously pubescent, rugous beneath, 3-5-lobed and serrate ; texture firm; petioles half an inch in length, terete, pubescent ; stipules lanceolate, acuminate, shorter than the petioles, with a single tooth near the base. Flowers 4-6 in a terminal corymbose panicle, rather smaller than those of R. odoratus, purple. Pedicels 6-8 lines long, not glandular. Calyx about two-thirds as long as the corolla, pubescent ; sepals ovate-oblong, acuminate, somewhat foliaceous at the extremity. Petals ovate, obtuse.
Hab. On the Rocky Mountains.
Obs. This species resembles R. odoratus, but is more nearly allied to R. nutkanus. It differs from the latter, particularly in not being glutinous, and in the smooth calyx, which is longer than the petals. Dr. James states, that the fruit is large and delicious.

 

Other articles:
• Golden Checklist Flora:  Rubus idaeus var. strigosus;  

Rubus idaeus L. var. strigosus (Michx.) Maxim. “American Red Raspberry”

 
 

Rubus strigosus Michx., Fl. Bor.-Amer. (Michaux) 1: 297 (1803).

 

Rubus idaeus var. strigosus (Michx.) Maxim., Bull. Acad. Imp. Sci. Saint-Pétersbourg 17: 161 (1872).

 

Other articles:
• Golden Checklist Flora:   Sanguisorba minor;  

Sanguisorba minor Scop. “Burnet”

 

Literature Cited:
- Scopoli, Giovanni Antonio, 1772.  

Scopoli (1772, t. 1, p. 110-111) ...

168. Sanguisorba Minor.
Sanguiforba fpicis fubrotundis ; floribus polygamis. Fl. Carniol. p. 279. n. 1.
Ptterium inerme caulibus fubangulofis. LINN. Syft. Nat. p. 629. Spec. Plant. p. 994. n. 1.
Pimpinella polyftemon. HALL. Hift. n. 706.
Sanguiforba polyanthera, foliis ovatis ; fpica brevi. BOEHM> Lipf. 17.
Pimpinella fanguiforba, minor. MATTHIOL. Diofc. p. 657.
Diagn. Spicae fubrotundae. Flores polygami, polyandri, digyni.
Habitat in aridis et apricis.
Pinnarum paria (ad 12). Flores feminei fuperiores digyni. Hermaphrodidi plures, ftaminibus longis (ad 40), pariter digyni. His addit HAMMERUS Mafculos, etiam a me vifos.

 

Other articles:
• Golden Checklist Flora:  Sorbus aucuparia;  

Sorbus aucuparia L. “European Mountain Ash”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 477) ...

Original Text
1. SORBUS foliis pinnatis utrinque glabris. Hall. helv. 250. aucuparia.
Sorbus foliis pinnatis. Hort. cliff. 188. Fl. fuec. 400. Mat. med. 235. Roy. lugdb. 274.  
Sorbus fylvefttis (sic), foliis domefticae fimilis. Bauh. pin. 415.  
Sorbus fylveftris. Cam. epit. 161.  
Habitat in Europae frigidioribus.  

 

Other articles:
• Golden Checklist Flora:  Astragalus agrestis;  

Astragalus agrestis Douglas ex G. Don. “Purple Milkvetch”

 

Literature Cited:
- Don, George, 1831-1838.  

Don, George [1798-1856] A general history of the dichlamydeous plants :comprising complete descriptions of the different orders...the whole arranged according to the natural system. London : J.G. and F. Rivington, 1831-1838. p. 258

64. A. agrestis (Dougl. mss.) stem erect, smooth ; leaflets ovate-lanceolate, obtuse, beset with adpressed hairs ; peduncles elongated, longer than the leaves ; spikes of flowers capitate ; calyx villous ; breacteas lnceolate, not half so long as the calyx. ♃ H. Native of North America, near the Columbia river in fields. Flowers purple.
Field Milk-Vetch. Pl. 1 foot.

H. = Hardy
♃. = Perennial herbaceous.

 

Other articles:
• Golden Checklist Flora:  Astragalus crassicarpus;  

Astragalus crassicarpus Nutt. “Groundplum Milkvetch”

 

Literature Cited:
- Nuttall, Thomas, 1813.

Other articles:
• Golden Checklist Flora:  Fraser's Catalogue, publication details;  

Nuttall (1813, No. 6) published his Astragalus crassicarpus ...

Original Text
6. * Astragalus crassicarpus. ‡ Fruit about the size and form of A. physodes, but thick and succulent. Collected above the River Platte.

 

Other articles:
• Golden Checklist Flora:  Astragalus drummondii;  

Astragalus drummondii Dougl. ex Hook. “Drummond's Milkvetch”

 

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.  

Hooker (1831) 1(3): 153-4 ...

Original Text
16. A. Drummondii ; elatus, erectus, caulibus sulcatis foliisque subtus pedunculisque valde molliter hirsutis, stipulis ovatis valde acuminatis, foliolis 13-15-jugis liueari-oblongis oblongisve obtusis basi in petiolulum perbrevem attenuatis, pedunculis folio longioribus, racemis elongatis laxis, bracteis subulatis pedicello longioribus, floribus pendulis, calycibus nigro-hirsutis, legumiuibus stipltatis subsecundis cylindraceis glabris paululum falcatis semibilocularibus, sutura superiore obtusa, inferiore introflexa profunde canaliculata. (Tab. LVII.) — Douglas, MSS. in Herb. Hert. Soc.
Longer Latin diagnosis omitted.
Hab. First, I believe, gathered by Mr. Wright, very many years ago, in Hudson's Bay. (Herb. nostr.) Eagle and Red-Deer Hills of the Saskatchawan. Douglas. Carlton-House. Dr. Richardson. — The present very fine species, which Mr. Douglas wishes should bear the name of the indefatigable Drummond, has been long known to me from a specimen gathered more than thirty years ago, by Mr. Wright, son of the late eminent Dr. Wright of Edinburgh, in Hudson's Bay, but which, from its habit, (and being destitute of fruit,) I was led to refer to the genus Phaca. Its nearest affinity seems to be with the A. galegoides, Nutt., (A. racemosus, Pursh and De Cand.) : but that, besides being an inhabitant of more southern latitudes, is described as only subpubescent, whereas our plant is so remarkable for its hairiness, that Mr. Douglas in his MSS. has aptly compared it to the Oxytropis pilosa of the Old Worid.*
Tab. LVII Astragalus Drummondii. Fig. 1, Flower; fig. 2, Alae and carina, including the stamens and pistil; fig. 3, Pistil ; fig. 4, Raceme, with fruit: —natural size. Fig, 5, Section of a legume; fig. 6, Seed : — all but fig. 4 more or less magnified.

 

Other articles:
• Golden Checklist Flora:  Astragalus flexuosus;  

Astragalus flexuosus G. Don “Flexible Milkvetch”

 

Literature Cited:
- Don, George, 1831-1838.  

George Don (1832, v. 2, p. 256) ...

Original Text
33. A. flexuosus (Doug. mss.) plant erect, flexuous, downy in a young state ; stipulas distinct ; leaflets distant, linear, obtuse, beset with close-pressed hairs beneath ; flowers distant, racemose; calyx smoothish ; beateas not much longer than the pedicels. ♃ H. Native of North America, near the Columbia river. Flowers purple. Plant with the habit of Vicia cracca.
Var. β, alba (Dougl. mss.) flowers white.
Flexuous-stemmed Milk-Vetch. Pl. 1 foot.

 

Other articles:
• Golden Checklist Flora:  Astragalus laxmannii robustior;  

Astragalus laxmannii Jacq. var. robustior (Hook.) Barneby & S. L. Welsh “Prairie Milkvetch”

 

Literature Cited:
- Jacquin, Nicolaus Joseph van, 1776.  

Jacquin, Nicolaus(Nicolaas) Joseph von (1727-1817), Hort. Bot. Vindob. 3: 22 (t. 37) (1776). [Botanical Garden of Vienna.]

Original Text Comments
TABULA TRIGESIMA SEPTIMA. Plate Thurty Seven
ASTRAGALUS LAXMANNI Astragalus Laxmanni
Numerofas inter planta Imperii Ruffici, quas hortus acceptas referre debet liberalitati illuftris Erici Laxmanni, etiam hicce Altragalus eft, ex feminibus Sibiricis enatus. Ex radice perenni ... [Latin diagnosis omitted] ... nitida. Floret Junio & Julio. Ad latus feorfim vexillum, ala, carina, legumen & femen fifuntur. Among the numerous plants of the Russian Empire, these gardens must be appreciated for your generosity the illustrious Eric Laxmannus, who is also here Astragalus, sprung from the seeds of the Siberians. Perennial root ... [Latin diagnosis omitted] ... sharp. Flowers June & July. The flag, wing, keel, pod and seed are presented separately.

Other articles:
• Glossary:  dolabriform;  

(Syn: Astragalus adsurgens Pall. ) Pallas, Peter Simon von, 1800-1803, Species Astragalorum Descriptae et Iconibus Coloratis Illustratae a P. S. Pallas, Eq... cum Appendice. Lipsiae [Leipzig]. p. 40. and Tab XXXI.

Original Text Translation
XLIV. ASTRAGALUS adfurgens. Tab. XXXI. XLIV. ASTRAGALUS adsurgens. Tab. XXXI.
Latin diagnosis omitted. I don't see any reference to attachment of the hairs, i.e., dolabriform or medifixed.
Crefcit hic Aftragalus tantun in regionibus Trans-Baicalenfibus, cim A. Laxmanni promifcue, frequens ad Selengam, Ononem, circa Tarei-noor et ufque in Mongoliae defertum. Planta tota, etiam foliis, duriufcula, ut Glycirrhizae, minunque tenera cum generibus plerisque. Astragalus grows here only in the regions of the Trans-Baicalans, and promiscuously to[with] A. Laxmannus, frequent to the Selengam, Ononem, around the Tarei-noor, and it was carried all the way into Mongolia. The whole plant, including its leaves, is quite hard, as in most species of Glycirrhiza and less tender.
  Transbaikal, Trans-Baikal, Transbaikalia, or Dauria is a mountainous region to the east of or "beyond" (trans-) Lake Baikal in Far Eastern Russia.

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.  

Hooker (1831, v. 1, -. 149-150) ... publication of vaeirty robustior

Original Text
Leguminibus rectis.
4. A, adsurgens; elongatus, adscendens vel prostratus, glabriusculus, foliolis 8-12-jugis oblongis, stipulis ovatis acuminatis membranaceis, pedunculis folio longioribus, spicis oblongis ovatis rotundatisve, floribus densis erectis, vexillo alis vix 1-3 longiore, leguminibus erectis compactis oblongis subtriquetris hinc sulcatis appresso-pubescentibus vix calyce nigro-hirsuto longioribus. — “Pall. Astr. n. 44. t 31.De Cand, Prodr, v. 2. p. 287. — A. Laxmanni. Pall. Astr. t. 30? (fide DC.) Jacq. Hort. Vind. v. 3. t 37? (sed spicis elongatis et foliolis angustioribus.) De Cand. Prodr. v. 2. p. 287. — β. robustior ; calycibus albido-pilosis, pilis nigris perpaucis. A. nitidus. Douglas, MSS. in Herb. Hort. Soc.
Hab. α. Plains of the Assinaboin and Saskatchawan Rivers, as for as the mountains. Dr. Richardson; Drummond; Douglas. β. Common in the mountain-vallies, from the Kettle Falls to the sources of the Columbia, on the West side of the Rocky Mountains. Douglas. — l possess the Asiatic A. adsurgens and A. Laxmanni from Dr. Fischer and Mr. Prescott ; but I am unable to distinguish them specifically. Our specimens from America seem to be identical with them. Flowers purple-blue.

Literature Cited:
- Barneby, R. C. and S. L. Welsh, 1996.  

Astragalus laxmannii Jacq.(1776) has priority over A. adsurgens Pall. (1800), Barneby and Welsh (1996) published the required name to recognize this:

Original Text
Astragalus laxmannii var. robustior (Hooker) Barneby & Welsh, comb. nov., based on A. adsurgens var. robustior Hooker, Fl. Bor.-Amer. 1:149. 1831.
Astragalus nitidus var. robustior (Hooker) M. E. Jones, Contr. W. Bot. 10:64. 1902.
Astragalus adsurgens ssp. robustior (Hooker) Welsh, Iowa State J. Sci. 37: 357. 1963.

 

Other articles:
• Golden Checklist Flora:  Astragalus parryi;  

Astragalus parryi A. Gray “Parry's Milkvetch”

 

 

Other articles:
• Golden Checklist Flora:  Astragalus shortianus;  

Astragalus shortianus Torr. & A. Gray “Short's Milkvetch”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & A. Gray (1840, v. 1, p. 331) ...

Original Text
12. A. Shortianus (Nutt. m.s.): “ " stemless, canescent with appressed shining hairs; leaflets 5-7 pairs, roundish-elliptical or ovate, very obtuse ; stipules ovate, obtuse; peduncles shorter than the leaves ; raceme oblong ; calyx clothed with white hairs, with rather long subulate teeth ; legume large and turgid, cymbiform, with a short curved point, black and tranversely wrinkled.
“Rocky Mountains, towards the plains of the Oregon. — Almost entirely silvery white. Leaflets nearly as broad as long, twice as large as in the preceding species, which it nearly resembles. Flowers ochroleucous ?” Nuttall.

Named for Charles Wilkins Short, friend of Nuttall and Gray (Weber & Wittmann, 2012).

 

Other articles:
• Golden Checklist Flora:  Colutea arborescens;  

Colutea arborescens L. “Bladder Senna”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 723) ...

Original Text
1. COLUTEA arborea, foliolis obcordatis. Hort. cliff. 365. Roy. lugdb. 374. Sauv. monfp. 238. arborefcens.
Colutea veficaria. Bauh. pin. 396.  
Colutea. Dod. pempt. 784.  
β. Colutea africana, fenne foliis, flore fangvineo. Comm. rar. 11. t. 11.  
Habitat in Auftria, G. Narbonenfi, Italia praecipue ad Vefuzium.  

 

Other articles:
• Golden Checklist Flora:  Dalea candida;  

Dalea candida Willd. “White Prairie Clover”

 

 

Other articles:
• Golden Checklist Flora:  Dalea purpurea;  

Dalea purpurea Vent. “Purple Prairie Clover”

 

Literature Cited:
- Ventenat, Etienne P., Jacques M. Cels, and Henri J. Redoute, 1801.  

Described by Ventenat (1800, Pl. 40) ...

Original Text
(Short excerpt.)
Interpretation
Plante herbacee, vivace, remarquable par l'elegance de son port et par l'eclat de ses fleurs, decouverte par Michaux dans le pays des Illinois. Elle a ete introduite chez Cels en l'an 5, et elle fleurit en fructidor. Herbaceous plant, perennial, remarkable for the elegance of its habit and the brilliance of its flowers, discovered by Michaux in the Illinois country. It was introduced to the garden of J. M. Cels in year 5 [1794?], and it flowers in Fructidor [late August to early September].

 

Other articles:
• Golden Checklist Flora:  Glycyrrhiza lepidota;  

Glycyrrhiza lepidota Pursh “American Licorice”

 

Literature Cited:
- Nuttall, Thomas, 1813.

Other articles:
• Golden Checklist Flora:  Fraser's Catalogue, publication details;  

Nuttall (1813) writing in Fraser's Catalogue

Original Text
45 *Liquiritia lepidota. ‡ Met from the source to the confluence of the Missourie ; and is probably the liquorice mentioned by Sir Alexander M'Kenzie, as found on the coasts of the North Pacific Ocean.

IPNI (2021) annotates this name as nom. inval.

Literature Cited:
- Pursh, Frederick, 1814.

Other articles:
• Golden Checklist Flora:  Pursh, 1814, publication details;  

Pursh (1814, v. 2., p. 480) validly published Glycyrrhiza lepidota indicating Fraser's Catalogue as a source of the name and that he had seen it dried, alive, and in flower.

Original Text Comments
  579. GLYCYRRHIZA. Gen. pl. 1197.  
lepidota. 1. G. foliolis oblongis acutis sericeo-villosis, leguminibus racemosis oblongis hispidis. &mdash. Fraser. catal.

On the banks of the Missouri. ♄ July, Aug. v. s. v. v. s. fl.

I have seen it dry; I have seen it alive or in flower.

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Nuttall, 1818, publication details;
• Glossary:  vexillum;  

Nuttall (1818, v. 2, p. 106) also published Glycyrrhiza lepidota. He does not indicate it is a new name. Nevertheless, it is treated as an isonym. Nuttall does give credit to Bradbury for first detecting the species around St. Louis.

Original Text
506. GLYCYRRHIZA. L.
Calix mostly bilabiate, gibbous at the base. Vexillum including the wings and carina. Legume subovate or oblong, compressed, and mostly hispid, 2 to 6-seeded.
Herbaceous; leaves pinnate, stipules cauline; flowers capitate, spiked or racemose.
Species. 1. G. lepidota. T. N. in Fras. Catal. Ph. 2, p. 480. Leaflets oblong-lanceolate, acute, everywhere squamulose, under surface covered with glandulose atoms; spikes axillary, acute, flowers crowded; legume oblong, many-seeded, echinate, setæ uncinate.
Hab. Abundant around St. Louis, where it was first detected by Mr. John Bradbury, F. L. S.; it is also common on the alluvial banks of the Missouri to the Mountains, and is in all probablity the Liquorice mentioned by Sir A. Mackenzie as indigenous to the coasts of the North Pacific Ocean.
Obs. Roots flagelliform, creeping, and very long, possessing in no inconsiderable degree the taste of liquorice. Stem erect, 3 to 5 feet high; spikes pedunculate; flowers whitish, dense, sessile; calix almost equally 5-parted, segments subulate. Vexillum ovate-oblong, nearly straight. Legume oblong, compressed, hispid, 5 or 6-seeded, much resembling the fruit of Xanthium spinulosum, not spontaneously opening. Nearly allied to G. foetida, and like that species emitting a somewhat disagreeable resinous odor. This plant appears to destroy the artificial distinction by which Glycyrrhiza and Liquiritia have been separated; as it can be equally referred to either one or the other.
The South of Europe, Tartary and the Levant furnish the other 6 species of this genus.

 

Other articles:
• Golden Checklist Flora:  Hedysarum boreale;  

Hedysarum boreale Nutt. “Utah Sweetvetch”

 

Literature Cited:
- Nuttall, Thomas, 1818.  

508. HEDYSARUM. L.
Calix 5-cleft. Carina transversely obtuse. Loment of several articulations: joints i-seeded, compressed, and mostly hispid.
Herbaceous or suffruticose; leaves simple, ternate, or pinnate; stipules cauline, often bath general and partial; flowers mosth racemose, rarely solitary, racemes axillary or terminal, often paniculate; flowers (in all the North American species) by pairs, or by 3's, each pair or aggregate subtendtd by 3 unequal and deciduous bractes, the 2 lateral ones minute, calix naked at the base, the lowest segment of the border often elongated; flowers violaceous, rarely, white, the carina often expanding and ejecting the stamina; the leaves in a few species exhibit a spontaneous motion.
...
18. * boreale. Caulescent, subdecumbent, leaves pinnate (7 or 8 pair), leaflets oblong-obovate, partly villous; racemes long pedunculate, axillary, stipules sheathing, subulate; articulations of the loment nearly round, and rugose. H. alpinum? Mich. Fl. Am. 2. p. 74. Hab. In arid and denudated soils around Fort Mandan, on the banks of the Missouri. Flowering in June and July. Flowers of a fine red and numerous; common petiole very short; calix subulate, wings of the corolla short.

 

Other articles:
• Golden Checklist Flora:  Ladeania lanceolata;  

Ladeania lanceolata (Pursh) A. N. Egan & Reveal “Dune Scurfpea”

 

Literature Cited:
- Pursh, Frederick, 1814.  

Syn: Psoralea lanceolata Pursh (1814, v. 2, p. 475) ...

Original Text
4. P. pubescens ; foliis ternatis : foliolis elongato-lanceolatis, petiolis crassis, spicis axillaribus folio vix longioribus densifloris, floribus pedicellatis, bracteis pedicello vix longioribus, dentibus calycis coloratis. lanceolata.
On the banks of the Missouri. ♃. July, Aug. v. s. Flowers bright blue, small.  

“Collected at an unknown place and date, probably on the Missouri River (Moulton, 1999).”

Literature Cited:
- Rydberg, Per Axel, 1915.  

Syn: Psoralidium lanceolatum (Pursh) Rydberg (1919, v. 24, p. 13)

Original Text
1. Psoralidium lanceolatum (Pursh) Rydberg.

Psoralea lanccolata Pursh. Fl. Am. Sept. 475. 1814
Psoralea elliptica Pursh. Fl. Am. Sept. 741. 1814.
Psoralea arenaria Nutt. Gen. 2: 103. 1818.
Psoralea laxiflora Nutt.; T. & G. Fl. N. Am. 1: 299. 1838.
? Psoralea scabra Nutt.; T. & G. Fl. N. Am. 1: 300. 1838.
Lotodes ellipticum Kuntze, Rev. Gen. 193. 1891.
Lotodes ellipticum angustissimum Kuntze, Rev. Gen. 193. 1891.
A perennial, with a creeping branched rootstock; stem 1.5-4 dm. high, aromatic, glandular- punctate throughout, sparingly strigose; leaves palmately 3-foliolate; stipules linear-lanceolate to subulate, 3-10 mm. long; petioles 1-2 cm. long; leaflets 1-4 cm. long, oblanceolate to linear or those of the earlier leaves obovate-oblanceolate, acute to rounded and mucronate at the apex, sparingly strigose, especially along the veins beneath, and conspicuously punctate; peduncles 2-5 cm. long; racemes short, 1-2.5 cm. long; bracts minute; calyx campanulate, sparingly strigose, 2 mm. long; lobes nearly equal, obtuse, glandular-punctate; corolla white or slightly purple-tinged, 5-6 mm. long; banner almost orbicular; blades of the wings obliquely oblong- oblanceolate; keel-petals scarcely lobed at the base and usually purple-tipped; stigma capitate; pod globose, 5 mm. long, conspicuously glandular-warty, sparingly strigose or glabrate, the beak short, erect.

Type locality: Banks of the Missouri [probably Montana]
Distribution: Missouri to Saskatchewan, Alberta, and Arizona
Illustrations: Hook. Fl. Bor. Am. pl. 51; Britt. & Brown, Ill. Fl. f. 2086; ed. 2. f. 2491.

Literature Cited:
- Egan, Ashley N., and James L. Reveal, 2009.  

Egan & Reveal (2009, v. 19, p. 310-314) ...

1. Ladeania lanceolata (Pursh) A. N. Egan & Reveal, comb. nov. Basionym: Psoralea lanceolata Pursh, Fl. Amer. Sept. 475. 1814 [actual date, Dec. 1813]. Psoralidium lanceolatum (Pursh) Rydbergm N. Amer. Fl. 24: 13. 1919. TYPE. U. S. A. Along the Missouri River, 1811, T. Nuttall s.n. (lectotype, designated by Grimes, 1989: 22, PH-LC 182; duplicate, NY [PH-LC fragm.]). Figure 1.
Although traditionally the type was considered to have been collected by Meriwether Lewis in 1804, Reveal et al. (1999:42) showed that this sheet could only have been gathered by Thomas Nuttall in 1811. If indeed Pursh had Lewis 42 prior to 1813, that specimen is now lost and Grimes' typification is an effective lectotypification (Barbeby, 1989). ...

 

Other articles:
• Golden Checklist Flora:  Lathyrus eucosmus;  

Lathyrus eucosmus Butters & H. St. John “Bush Vetchling”

 

Literature Cited:
- Butters, Freferick K., and Harold St. John, 1917.  

Butters, Frederick K., and Harold St. John, Studies in certain North American Species of Lathyrus Rhodora. 19(224): 156-163. August, 1917.

Lathyrus eucosmus, n. sp. — L. polymorphus of Torrey and Gray in part, and of later American authors, not of Nuttall; L. decaphyllus of Britton, Mem. Torr. Bot. Club. v. 207 (1894), not of Pursh or Hooker — humilis, ramosus saepe decumbens pilosus vel saepius basim versus glabratus, 1.5-3.5 dm. altus; caule 1-3 mm. crasso striato 4-angulato haud alato; stipulis lineari-lanceolatis attenuatis semi- sagittatis 5-22 mm. longis, 1-4 mm. latis, nervosis; foliolis 4-12 subalternis elliptico-lanceolatis mucronatis 1.5-6 cm. longis, 3-13 mm. latis, nervosis; cirrhis simplicibus vel 2-3-fidis; ramis cum pedunculis 1-3 folia superantibus 2-5-floriferis instructis; floribus magnis pulchris purpureis 1.8-3 cm. longis; calyce campanulato oblique 5-dentato, dentibus superioribus brevibus adscendentibus, inferioribus attenuatis patentibus.
Plant low and branching, often decumbent, pilose, or more frequently glabrate towards the base, 1.5-3.5 dm. tall: stem striate and 4-angled, not at all winged, 1-3 mm. in thickness: stipules linear- lanceolate, attenuate semisagittate, 5-22 mm. in length, 1-4 mm. in breadth, with prominent raised nerves; leaflets 4-12 in number, sub-alternate, elliptic-lanceolate and mucronate, the longitudinal nerves raised and prominent, leaflets 1.5-6 cm. long, 3-13 mm. wide; tendrils unbranched, or 2-3-parted: branches of the stem bearing 1-3 peduncles, which are 2-5-flowered, and exceed the leaves : flowers beautiful, large and purple, 1.8-3 cm. long; calyx campanulate obliquely 5-toothed, the upper teeth short and ascending, the lower elongate and divergent.
Specimens examined: Colorado: Rocky Mountains, 1862, E. Hall & J. P. Harbour, no. 111; low lands by streams on the plains, Golden City etc., May 25, 1870, E. L. Greene, no. 94; Gunnison, altitude 7680 ft., July 7, 1901, C. F. Baker, no. 355; Sapinero, altitude 7250 ft., June 19, 1901, C. F. Baker, no. 181; clay hillside, altitude 5400 ft. Naturita, May 16, 1914, Edwin Payson, no. 312; Gato, June 18, 1899, C. F. Baker, no. 432; roadsides below Mancos, July 8, 1898, C. F. Baker, F. S. Earle and S. M. Tracy, no. 413; collected on Long's 1st expedition, Dr. James. New Mexico: altitude 7044 ft., Santa Fe, June, 1874, J. T. Rothrock, no. 3; altitude 7200 ft., Santa Fe, May 4, 1897, A. A. & E. Gertrude Heller, no. 3658 (type in Gray Herb.); 1847, A. Fendler, no. 115; Gray, altitude 6000 ft., June 13, 1898, Josephine Skehan, no. 24; Fort Wingate, 1882, W. Matthews, no. 18. Arizona: Little Colorado, Dr. Newberry; 1880, Mr. & Mrs. J. G. Lemmon.

 

Other articles:
• Golden Checklist Flora:  Lathyrus lanszwertii;  

Lathyrus lanszwertii Kellogg “Lanszwert's Pea”

Lathyrus lanszwertii Kellogg, Proceedings of the California Academy of Natural Sciences, v. 2, p. 150 (1863). https://www.biodiversitylibrary.org/item/24102#page/164 Note the title of this journal includes “Natural” which was dropped in 1869.

 

Other articles:
• Golden Checklist Flora:  Lupinus argenteus;  

Lupinus argenteus Pursh “Loosely Flowered Silver Lupine”

 
 

Harrington (1954) and Ackerfield (2015) neither use keel decoration as a key character nor do they describe the decoration of the keel. California floras, e.g., Munz (1965) and Baldwin (2012), use a ciliate keel, as a key character to identify L. argenteus. Welch, et al. (1993) note that the keel can be glabrous or variously ciliate.

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 2, p. 468) ...

Original Text
argenteus. 4. L. perennis ; foliis digitatis : foliolis (5-7) lineari-lanceolatis acutis supra glabris subtus argenteo-sericeis, calycibus alternis inappendiculatis : lapio superiore ontusa ; inferiore integro.
  On the banks of the Kooskoosky. M. Lewis. ♃. June, July. v. s. in Herb. Lambert. Flowers small, cream-coloured.

 

Other articles:
• Golden Checklist Flora:  Lupinus caudatus;  

Lupinus caudatus Kellogg “Kellogg's Spurred Lupine”

Of those collections of Lupinus caudatus in Jefferson County determined to an infraspecific name, nearly all are determined var. argophyllus.

Literature Cited:
- Gray, Asa, 1849.  

The first use of the name argophyllus was by Gray (1848) in Plantae Fendlerianae Novi-Mexicanae.

Original Text
166. Lupinus decumbens, Torr. in Ann. Lyc. New York, 2. p. 191. Var. argophyllus; caule foliisque argyreo-sericeis, supra viridiusculis. &mdash: A foot high, a silvery white species; more so than in the original specimens of L. decumbens, which imperfect as they are, I am confident belong to the same species with ours. It is a very handsome plant, with flowers as large as in L. perennis, and apparently light blue. A characteristic of the species is the saccate, almost spurred base of the calyx, which is quite a conspicuous as in L. laxiflorus, next to which it should be placed. The same plant was gathered near the sources of the Platte in the first expedition of Col. Fremont, and forms part of what was called L. ornatus in the Botanical catalogue of that expedition.

Literature Cited:
- Kellogg, Albert, 1863.  

Kellogg, Albert. 1863. Lupinus caudatus. Proceedings of the California Academy of Natural Sciences. v. 2 (1858-1862), p. 197. https://www.biodiversitylibrary.org/item/24102#page/211/

Original Text
Lupinus caudatus (Kellogg). Fig. 61.
Stem persistent, somewhat decumbent ; leafy and branching, silvery or satiny, appressed pubescent throughout.
Leaflets five to seven, linear-lanceolate, acute, mucronate, narrowed towards the base, about as long as the petiole.
Stipules persistent, small lance-acumuinte.
Flowers blue, scattered and sub-verticillate, floral portion about twice the length of the peduncle, or two or three times the length of the petioles.
Bracts caducous, twice the length of the pedicels ; calyx tubular campanulate, upper lip straight, two-toothed, (not colored) short and somewhat subulately spurred at the base, (spur erect nearly half the length of the pedicel) ; lower lip entire, elongated, carinate ; linear bracts conspicuous ; banner satiny pubescent on the back, chiefly along the middle portion ; wings with an erect claw.
Keel, silky-ciliate.
Stigma, naked.
Legumes (embryo) linear, silky ; seeded.
Closely allied to L. calcaratus (Kellogg). But the general appearance is quite distinct, being very silvery sericeous, with blue flowers. The calyx is not colored ; the upper lip straight ; the spur short, sharp and erect ; the leaflets straight and radiating, only five or six in number ; the flowers also fewer, and the spike less crowded. Found in the same localities, but more rare.

Literature Cited:
- Cockerell, T. D. A., 1902.  

Cockerell, T. D. A., 1902. Notes on Southwestern Plants. Torreya. v. 2, n. 3 (March 1902)

Original Text
Lupinus Helleri Greene, Pittonia, 4 : 1 34. Santa Fe, N. M. This name may be objected to on account of the prior L. Hellerae Heller, though I do not consider the names identical. However, it appears to me that L. Helleri is L. decumbens argophyllus, A. Gray (Pl. Fendl. 37. 1849), so its proper name will be Lupinus argophyllus. The type locality of argophyllus is “around Santa Fe,” where, in fact, it is abundant.

Literature Cited:
- Welsh, Stanley L., 1978.  

Lupinus caudatus var. argophyllus (A.Gray) S.L.Welsh, Great Basin Naturalist 38(3): 327 (1978).

Original Text
...
Three rather weak varieties are known from Utah. They are separable only arbitrarily, but seem to represent at least trends within variation.
...
Var. argophyllus (A. Gray) Welsh stat. nov. based on Lupinus decumbens var. argophyllus A. Gray Mem. Amer. Acad. 4:37. 1849. [L. caudatus ssp. argophyllus (A. Gray) Phillips; L. argophyllus (A. Gray) Cockerell; L. laxiflorus var. argophyllus (A. Gray) M. E. Jones; L. helleri Greene; L. aduncus Greene]. Pinyon-juniper, mountain brush, ponderosa pine, and grassland communities at 1570 to 2430 feet in Beaver, Garfield, Kane, and San Juan cos.; Wyoming south to New Mexico; 7 (iii).

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012) do not accept infraspecific names for Lupinus caudatus Kellogg.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015) accepts two varieties of Lupinus caudatus in Colorado: var. argophyllus (A. Gray) L. Philips (sic), and var. utahensis (S. Watson) S. L. Welsh.

 

The Jepson Manual of California treats Lupinus caudatus as a synonym of L. argenteus var. heteranthus.

 

Other articles:
• Golden Checklist Flora:  Medicago;  

Medicago L.

Please summarize the phylogeny of medicago and the development of alfalfa as an agricultural crop.

ChatGPT:

Medicago is a genus of flowering plants in the legume family, Fabaceae, which includes more than 80 species, some of which are used as forage crops. Alfalfa, also known as lucerne, is the most widely cultivated species in the genus and is a major forage crop in many parts of the world.

Alfalfa is believed to have originated in central Asia, and its cultivation dates back at least 2,000 years. It was introduced to Europe by the Arabs in the eighth century and was later brought to the Americas by Spanish colonizers in the sixteenth century. The plant was valued for its high protein content, drought tolerance, and ability to fix atmospheric nitrogen, making it an important crop in areas with poor soils.

In terms of phylogeny, Medicago is a member of the tribe Trifolieae, which also includes clovers and other forage legumes. Molecular phylogenetic studies suggest that the genus Medicago is monophyletic, meaning that all species in the genus share a common ancestor, and that it is most closely related to the genus Melilotus.

Alfalfa is a tetraploid species, meaning that it has four sets of chromosomes, and its genome is quite large, with an estimated size of around 800 million base pairs. This complexity has made genetic improvement of alfalfa a challenge, but recent advances in genomics and molecular breeding techniques are helping to overcome some of these difficulties.

Overall, the development of alfalfa as an agricultural crop has been a long and complex process, but the plant's many useful characteristics have made it an important component of many farming systems around the world.

 

Other articles:
• Golden Checklist Flora:  Medicago lupulina;  

Medicago lupulina L. “Black Medick”

Medicago lupulina L. “Black Medick” is a common little weed found on North and South Table Mountains and Tin Cup Ridge. It is also adventive in gardens.

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 779) ...

Original Text
7. MEDICAGO fpicis ovalibus, leguminibus reniformibus monofpermis, caule procumbente. Fl. fuec. 621. Dalib. parif. 281. lupulina.
Trifolium leguminibus fpicatis reniformibus nudis monofpermis, caule procumbente. Hort. cliff. 375. Roy. lugdb. 380.  
Trifolium pratenfe luteum, capitulo breviore. Bauh. pin. 328.  
Trifolium pratenfe luteum. Fuchf. hift. 819.  
Habitat in Europae pratis. ♂  

 

Other articles:
• Golden Checklist Flora:  Medicago sativa;  

Medicago sativa L. “Alfalfa”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 778) ...

Original Text
fativa. 5. MEDICAGO pedunculis racemofis, leguminibus contortis, caule erecto glabro. Hort. cliff. 377. Roy. lugdb. 281.
  Medica legitima. Cluf. hift. 2. p. 242.
  Medica fativa. Morif. hift. 2. p. 150. f. 2. t. 16. f. 2.
  Foenum burgundicum. Lob. ic. 2. p. 36.
  Habitat in Hifpaniae, Galliae apricis.

Literature Cited:
- Prosperi, Jean-Marie, Eric Jenczewski, Marie-Helene Muller, Stephane Fourtier, Jean-Paul Sampoux, and Joelle Ronfort, 2014.  

Prosperi, et al., 2014 summarize current thought on the domestication of alfalfa.

It is difficult to identify the first traces of alfalfa (Medicago sativa L.) domestication. Different authors suggested that alfalfa was cultivated 9,000 years ago in some rare locations in its centre of origin (Near East to Central Asia). Sinskaya in 1950 suggested that its distribution spread throughout the Middle East by 1,000 BC, and from there, to China and India. But actually, we have no element to validate these assumptions. The first written references mentioned the introduction of alfalfa to Greece by the Medes armies, its spread into Italy and then all over the Roman Empire in Europe. During the Middle Ages, the interest in alfalfa regressed considerably in Europe.
Alfalfa was reintroduced into Spain through North Africa with the Moors. It then crossed the Pyrenees around the fifteenth century. Thereafter, it was introduced to South America (Mexico, Peru, Chile) by the Spanish in the sixteenth century, and then to the United States through California at the beginning of the nineteenth century as “Chilean clover.”

 

Other articles:
• Golden Checklist Flora:  Melilotus albus;  

Melilotus albus Medik. “White Sweet Clover”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 765) ...

Original Text Interpretation
TRIFOLIUM  
* Meliloti leguminibus nudis polyfpermis.  
...  
4. TRIFOLIUM leguminibus racemofis nudis difpermis, caule erecto. Hort. cliff. 376. Hort. upf. 223. Fl. fuec. 619. Mat. med. 355. Roy. lugdb. 223. Dalib. parif. 225. M. officinalis.  
Trifolium odoratum f. Melilotus. Dod. pempt. 567.    
Melilotus officinarum germaniae. Bauh. pin. 331.    
γ. Melilotus officinarum germaniae, flore alba. Tournef. inft. 407.   The Greek letter “γ” corresponds to the letter “c” in the excerpt from Medikis (1786), below.
Lotus fylveftris, flore albo. Tabern. hift. 893.    
Habitat in Europae campeftribus. ☉ ♂    

Literature Cited:
- Medikus, Friedrich Kasimir, 1786.
Full Size ImagePage from Medikus (1786) publishing Melilotus albus.  

Medikus, Friedrich Kasimir (1736-1808), [May] 1786. Versuch einer neuen Lehrart die Pflanzen nach zwei Methoden zugleich, nehmlich nach der künstlichen und natürlichen, zu ordnen, durch ein Beispiel einer natürlichen Familie erörtert. [Attempt at a new method of classifying plants according to two methods at the same time, namely according to the artificial and the natural, explained by an example of a natural family.] Vorlesungen der Churpfälzischen physicalisch-öconomischen Gesellschaft. [Lectures of the Churpfälzische Physical-Economic Society.] Mannheim. 2: 382 (1787). http://ds.ub.uni-bielefeld.de/viewer/fullscreen/2241757_002/392/

The following excerpt is taken from the image at left.

4. Melilotus albus.  
Trifolium M. officinalis c L. The German letter “c” in the text at left corresponds with the Greek letter “γ” in the Linnean description above.

 

Other articles:
• Golden Checklist Flora:  Melilotus officinalis;  

Melilotus officinalis (L.) Lam. “Yellow Sweet Clover”

 

Literature Cited:
- Lamarck, Jean-Baptiste Pierre Antoine de Monet de, 1778.  

Melilotus officinalis (L.) Lam., Flore Françoise, ou Descriptions Succinctes de Toutes les Plantes qui Croissent Naturellement en France...Paris (Lamarck) 2: 595 (1779).

Original Text
III. Epi de fleurs pointu , lache , & ayant un ponce ou plus de longueur.
Melilot officinal. Melilotus officinalis.
Melilotus officinarum germaniae. Tournef. 407.
β Melilotus vulgaris altiffima frutefcens flore albo. Ibid.
Sa tige eft haute de deux pieds , dure & rameufe ; fes feuilles font petiolees , compofees de trois folioles glabres, ovales-oblongues , quelquefois un peu etroites & dentees dans leur partie fuperieure. Les fleurs font petites, de couleur jaune ou blanche, pendantes & difpofees fur des epis greles , laches & affez longs. Il leur fuccede des legumes courts , pendans , un peu rides, & qui renferment une on deux femences. La variete β s'eleve jufqu'a fix pieds , & porte des epis greles & fort longs. Cette plante croit dans les champs & fur le bord des hales , © ou a* ; fes feuilles & fes fleurs font refolutives , emollientes & anodines.

 

Other articles:
• Golden Checklist Flora:  Oxytropis lambertii;  

Oxytropis lambertii Pursh “Purple Locoweed”

Oxytropis DC., Astragologia 24, 66 (ed. qto.); 19, 53 (ed. fol.) (1802), nom. cons., includes 606 accepted species.

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 2, addendum, p. 740) ...

Original Text Comments
Oxytropis Lambertii. — O. acaulis, sericeo pilosa ; foliolis (19) lanceolato-ellipticis utrinque acutis, scapis folia aequantibus, spicis capitatis, bracteis linearibus longitudine calycis.  
On the Missouri. Bradbury. ♃". Aug. Sept. v. v. in Hort. Lambert. Flowers a beautiful purple. It approaches near to O. argentata and setosa.  

Literature Cited:
- Welsh, Stanley L., 1991.

Other articles:
• Glossary:   porrect;  

Original Text
The porrect beak of the keel is diagnostic for the genus, even though some species of Astragalus have extended keel apices; none are truly porrect.

 

Other articles:
• Golden Checklist Flora:  Oxytropis sericea;  

Oxytropis sericea Nutt. ex Torr. & A. Gray “White Locoweed”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1838, v. 1, p. 339) ...

Original Text
6. O. sericea (Nutt. ! mss.): “stemless, somewhat caespitose, shining and whitish with appressed silky hairs; leaflets linear-oblong or lanceolate (those of the primary leaves elliptical and obtuse) ; scapes longer than the leaves; spikes short, elongated in fruit ; bracts lanceolate, acuminate, shorter than the calyx; teeth of the calyx short and subulate ; legumes somewhat cylindrical, acuminate, 2-celled, canescently pubescent.”
Rocky Mountains. Nuttall! — Leaflets about ¾ of an inch long. Wings emarginate. Scapes stout and rigid in fruit. Legumes (including the beak) nearly an inch long, coriaceous, compressed contrary to the sutures. — Nearly related to O. Lamberti.

 

Other articles:
• Golden Checklist Flora:  Pediomelum tenuiflorum;  

Pediomelum tenuiflorum (Pursh) A. N. Egan “Slimflower Scurfpea”

 

Literature Cited:
- Pursh, Frederick, 1814.  

(Syn: Psoralea tenuiflora Pursh) Pursh (1814, v. 2, p. 475) ....

Original Text
3. P. pubescens, ramosissima; foliis ternatis; foliolis ellipticis utrinque rugoso-punctatis, pedunculis axillaribus folio longioribus sub-3-floris. tenuiflora.
On the banks of the Missouri. M. Lewis. ♃ Sept. v. s. in Herb. Lewis. Flowers very small, pale blue.  

Literature Cited:
- Egan, Ashley N., and James L. Reveal, 2009.  

Egan, Ashley N., and James L. Reveal (2009, v. 19, 310-314) ...

Original Text
Psoralidium tenuiflorum (Pursh) Rydberg was first collected on the historical expedition of Meriwether Lewis and William Clark on 21 September 1804 at the Big Bend of the Missouri River, which would later become Lyman County, South Dakota, U.S.A. (Moulton, 1999).
...
Frederick Pursh (1813) later described it as Psoralea tenuiflora Pursh. Since then, the species has experienced a murky taxonomic history. Rydberg (1919) designated Psoralea tenuiflora as the type of a new genus, Psoralidium Rydberg, wherein he recognized 14 species. … the variation across this species is great, but with no distinct segregation as to definitely allow specific designation, an ascentainment that the authors of this paper share.
Pediomelum tenuiflorum (Pursh) A. N. Egan, comb. nov. Basionym: Psoralea tenuiflora Pursh, Fl. Amer. Sept. 2: 475. Dec. 1813. Lotodes tenuiflorum (Pursh) Kuntze, Revis. Gen. Pl. 1: 194. 1891, nom illeg. Psoralidium tenuiflorum (Pursh) Rydberg. N. Amer. Fl. 24: 15. 1919. TYPE: U.S.A. South Dakota: Lyman Co., Big Bend of the Missouri River, 21 Sep. 1804, M. Lewis s.n. (lectotype, designated by Grimes, 1990: 36, PH-LC 184; duplicated, NY [fragm.], PH-LC 183).

 

Other articles:
• Golden Checklist Flora:  Robinia pseudoacacia;  

Robinia pseudoacacia L. “Black Locust”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 722) ...

Original Text
PfeudoAcacia. 1. ROBINIA pedunculis racemofis, foliis impari-pinnatis. Hort. upf. 212.
  Robinia aculeis geminatis. Hort. Cliff. 354. Gron. virg. 82. Roy. lugdb. 372.
  Acacia americana, filiquis glabris. Raj. hift. 1719.
  Acaciae affinis virginiana fpinofa, filiqua membranacea plana. Pluk. alm. 6. p. 73. 4. 4.
  Habitat in Virginia. ♄
  Conf. Pfeudo-Acacia hifpida, floribus rofeis. Catesb. car. 3. p. 20. t. 20.
POWO (2022) states the tree is native to Alabama, Georgia, Illinois, Kentucky, Pennsylvania, Tennessee, and Vermont; not Virginia. Of course at Linnaeus' time, Virginia was a colony, not a state. USDA Plants maps its nativity as US.

Literature Cited:
- Burns, Russell M., and Barbara H. Honkala, 1990.  

Black locust has a disjunct original range, the extent of which is not accurately known. The eastern section is centered in the Appalachian Mountains and ranges from central Pennsylvania and southern Ohio, south to northeastern Alabama, northern Georgia, and northwestern South Carolina. The western section includes the Ozark Plateau of southern Missouri, northern Arkansas, and northeastern Oklahoma, and the Ouachita Mountains of central Arkansas and southeastern Oklahoma (https://www.srs.fs.usda.gov/pubs/misc/ag_654/volume_2/robinia/pseudoacacia.htm, 16 March 2022).

 

Other articles:
• Golden Checklist Flora:  Securigera varia;  

Securigera varia (L.) Lassen “Purple Crownvetch”

 

 

Other articles:
• Golden Checklist Flora:  Thermopsis rhombifolia var. divaricarpa;  

Thermopsis rhombifolia var. divaricarpa (A. Nelson) Isely. “Spreadfruit Goldenbanner”

 

Literature Cited:
- Fraser, John, 1813.  

Fraser's (1813) Catalogue, most likely written by Nuttall, is the first reference to T. rhombifolia.

Original Text Comments
26. *Cytisus rhombifolius. ‡ Fl. not seen. I am a little surprised this is not marked by an “M” from the Missouri because in Nuttall (1818), two paragraphs down, Nuttall will tell us that he collected this plant at Fort Mandan.
 

Symbols used by Fraser (Nuttall).
* – New Species.
‡ – Perennial.
M. – from the Missourie. ;

Literature Cited:
- Pursh, Frederick, 1816.  

Pursh (1814) published Cytisus rhombifolia in the supplement to Vol. 2 of his Flora Americae Septentrionalis. Many of the entries in the supplement were Bradbury collections or from Nuttall descriptions in Fraser's Catalogue.

Original Text
Cytisus rhombifolius — C. pubescens ; racemis terminalibus erectis, leguminibus falcatis subarticulatis, foliolis oblongo-rhomboideis obtusis, stipulus rotundato-ovatis obliquis.

C. rhombifolius. Fraser. Catal. 1813.
In Upper Louisiana. Bradbury. v. s. in Herb. Bradbury. The stipulae are large and foliaceous ; the specimen I have seen was in fruit, and Mr. Bradbury has seen only one plant in flower.

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Glossary:  carina;  vexillum;  

Nuttall (1818) published it as Thermia rhombifolia eschewing the Thermopsis of R. Brown.

Original Text
401.THERMIA. Thermopsis. R. Brown. Hort. Kew. 3. p. 3.
Calix subcampanulate, half 4-cleft, the supper segment truncate and emarginate. Corolla papilionaceous, petals nearly equal in length; vexillum reflected at the sides; carina obtuse. Legume compressed and falcate, attenuated at the base, many-seeded.
Herbaceous; leaves ternate petiolate, stipules large and foliaceous; spikes terminal, interrupted, subverticillate, erect; flowers yellow. Very distinct in habit from Cytisus but requires further comparison with that genus?
Species. 1. *rhombifolia. Leaflets rhombi-ovate … … raceme interrupted. Cytisus rhombifolius T. N. in Fras Catal. 1813. Pursh, Flor. Am. Sept. 2. p. 741. Suppl. Obs. Roots perennial … … — On denudated argillaceous hills near Fort Mandan. This plant is very closely allied to Sophora lupinoides of Pallas, Thermopsis lanceolata of Brown, and they appear inseparable in genus, that species when in perfection produces a long verticillated spike of flowers; some of Pallas's specimens, however, in the herbarium of A. B. Lambert, Esq., have a single verticill of flowers only as in the starved specimen figured in the Botanical Magazine, in this species the leaves are on both sides closely covered with a silky villous; the primary leaves it appears occur sometimes simple but always accompanied by the stipules after the manner of Baptisia.

Literature Cited:
- Richardson, John, 1823.  

Richardson (1823) wrote the botanical appendix for Franklin's (1823) report of the voyage to the polar sea.

Original Text Comments

No. VII.

BOTANICAL APPENDIX,

BY

JOHN RICHARDSON

 
 
The collections of Pallas and Pursh, now belonging to Mr. Lambert, rendered the power of referring to his valuable Herbarium an object of the utmost importance to me ; and the desire of promoting the science, whoch so eminently distinguishes his character, induced him cheerfully to accord it. I include this note because it confirms that Lambert had collections made by Pallas. Pallas and Lambert must have been frequent correspondents. This is possibly relevant to explanation of how a Pallas' manuscript describing Chrysocoma nauseosa might have been seen by Pursh.
 
137. Thermopsis rhombifolia: Nuttall. Am. 1. p. 282. (C.)  

(C.) Denotes the sandy plains in the neighborhood of Carlton, strongly resembling the plains of the Missouri, upon which the American botanists have lately made extensive collections.

 
 

Literature Cited:
- Nelson A., 1898.  

Nelson (1898) ...

Original Text Comments
The species of this region, as they now appear to be, are T. montana, T. rhombifolia, and the two proposed species. The fruits of these are very characteristic, a fact shown in the accompanying plate. The four fruiting racemes are all from the Wyoming material, are mature and typical each of its species They were photographed on one plate, hence the size is relatively correct (about one-third natural size).  
Thermopsis rhombifolia n. sp. — … …  
It is found near streams, preferring the moist, rich soil among the open underbrush. Type specimens in Herb. Univ. of Wyo., no. 3424 by Elias Nelson, Pole creek, July 22, 1897 ; and no. 3903 by the writer, Johnson's ranch, Big Laramie river, August 8, 1897.  

Literature Cited:
- Harrington, H. D., 1964, 2nd ed..  

Harrington (1964, 2nd ed.) treated the Colorado taxa at the rank of species, saying, “… The mature or nearly mature fruit is necessary to be sure of the species. Even then some intergradations occur in Colorado plants.”

Literature Cited:
- Isely, Duane, 1978.  

Isely (1978) proposed var. divaricarpa despite recognizing that infraspecific divisions and those with T. macrocarpa are not that well-defined.

Original Text Comments
Thermopsis rhombifolia var. divaricarpa (A. Nels.) Isely, Comb. Nov.  .
Thermopsis divaricarpa A. Nels. Bot. Gaz. 25:275. 1898. Type: United States. Wyoming, Albany Co.: Johnson Ranch, 8 Aug 1897, A. Nelson 3902 (Holotype: RM!; Isotypes: GH!, NY!).  
I view Thermopsis of the western states as one vast complex, but have divided it into two “convenience” species, T. macrophylla of the Pacific states, and T. rhombifolia, which extends east to the high plains. These combinations provide the needed names. Isely also proposed var. montana based on a Nuttall type, and var. ovata based on T. ovata Robinson ex Piper.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittman (2012) treat Thermopsis rhombifolia, T. montana, and T. divaricarpa at the rank of species.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015) ...

Original Text Comments
This is a polymorphic species with considerable variation and is often divided into three species (T, divaricarpa, T. montana, and T. rhombifolia). However, when one examines all three species together, considerable overlap in morphology is evident. … Intermediates between all three taxa can be seen where their ranges overlap.  

 

Other articles:
• Golden Checklist Flora:  Trifolium hybridum;  

Trifolium hybridum L. “Alsike Clover”

 

 

Other articles:
• Golden Checklist Flora:  Vicia americana;  

Vicia americana Willd. “American Vetch”

Vicia americana Muhl. ex Willd., Sp. Pl., ed. 4 [Willdenow] 3(2): 1096 (1802).

 

Other articles:
• Golden Checklist Flora:  Vicia ludoviciana;  

Vicia ludoviciana Nutt. “Louisiana Vetch”

There is one collection of Vicia ludoviciana in Golden s.l. that was made under the south side of Castle Rock.

In Jefferson County, there is just one other collection, from Chatfield Farms. Otherwise, in Colorado there are collections along the northern Front Range, along the Arkansas River between Canon City and Pueblo, and then in scattered locations in other parts of Colorado.

The flowers of Vicia ludoviciana are quite small, (4.5)6-8(9) mm., and mostly white, compared to those of V. americana that are (12)15-22(25) mm. and mostly colored.

Literature Cited:
- Graustein, Jeannette E., 1967.  

Nuttall left Philadelphia on 2 October 1818, arriving at Fort Smith 24 April 1819. In mid-May he set off with a small Army detachment and several Cherokees for the Red River, traveling up the Poteau River and down the Kiamichi River. Nuttall collected on the Red River for three full days before time for the detachment to return to Fort Smith. He lost his companions and stayed in the area until mid-June. During this time he made many collections of new species, including Vicia americana (Graustein, 1967, p. 144).

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.

Other articles:
• Golden Checklist Flora:  Torrey & A. Gray, 1838-1843, publication details;  

Torrey & A. Gray (1838, v. 1, p. 271) ...

Original Text
8. V. Ludoviciana (Nutt. mss.): glabrous (except the young shoots) ; leaflets 10-12, elliptical or obovate, obtuse or emarginate ; stipules subulate, simple or semisagittate ; peduncle 2-6-flowered, at length longer than the leaves ; flowers (minute) closely approximated ; teeth of the calyx broad, acuminate, shorter than the tube ; legume broadly sabre-shaped, glabrous, 5-6-seeded ; seeds compressed, dark brown.
Grassy places on the Red River, and in Texas, Dr. Leavenworth ! “In Louisiana, Mr. Tainturuer,Nuttall. — ♃ Stem 2-3 feet long, rather stout, strongly angled, climbing. Leaflets 6-8 lines long, 2 lines wide, commonly emarginate. Stipules very small. Flowers blue, smaller than in V. Cracca, rarely solitary, often 2-6 on a peduncle. Legume ¾ of an inch long and 3 lines wide. — Mr. Nuttall in his manuscript describes the peduncles as 1-2-flowered, which is the case in some of our specimens ; but the peduncles are more commonly at least 4-flowered.

 

Literature Cited:
- Roth, Albrecht Wilhelm, 1788-1800.

Other articles:
• Golden Checklist Flora:  Vicia villosa;  

Vicia villosa Roth “Hairy Vetch”

Roth, Albrecht Wilhelm, 1757-1834 Tentamen florae Germanicae; continens enumerationem plantarum in Germania sponte nascentium. [The German flora containing a listing of plants that grow naturally in Germany.] Lipsiae, In Bibliopolio I.G. Mlleriano, 1788-1800. Volume 2, part 2, page 186.

“In locis limosis prope Vegesack ante aliquot annos inveni plantam, quae quotannis in horto sterilliori culta non mutavit habitum. [A few years ago I found a plant in muddy areas near Vegesack, which has not changed its habit of being cultivated every year in a more sterile garden.]&rduo;

Vegesack is located about 20 km (12 mi) north from the centre of Bremen-city at the mouth of the river Lesum.

 

Other articles:
• Golden Checklist Flora:  Erodium cicutarium;  

Erodium cicutarium (L.) L'Her. ex Aiton “Redstem Stork's Bill”

 

Literature Cited:
- Aiton, William, 1789.  

Aiton (1789, v. 2, p. 414) ...

Original Text
cicutarium. 3. E. pedunculis multifloris, foliis pinnatis : foliolis feffilibus pinnatifidis. L' Herit. n. 12.
  Geranium cicutarium. Sp. pl. 951. Curtis lond.
  Hemlock-leav'd Crane's-bill.
  Nat. of Britain.
  Fl. April-September. H. ☉.

 

Other articles:
• Golden Checklist Flora:  Geranium caespitosum;  

Geranium caespitosum James “Pineywoods Geranium”

 

Literature Cited:
- James, Edwin, ed., 1823.  

James (1823, v. 2, p. 3) ...

Original Text Interpretation and Comments
About the sandstone ledges we collected a geranium † intermediate between the crane's bill and herb robert, the beautiful calochortus, [C. elegans, Ph.] and a few other valuable plants. The party was at the mouth of Platte Canyon.

 
† G. caespitose, sub-erect, pubescent, sparingly branched above. Radical leaves reniform deeply 5-7 cleft. The flower is a little larger than that of G. robertianum, and similarly coloured, having whitish lines towards the base of the corrolla. We also saw here the Campanula decipiens. Pers. Lysimachia ciliata, Ph. Troximon glaucum, N. with two or three belinging to genera with which we were unacquainted.
Campanula decipiens = ?, maybe Campanula rotundifolia L
Lysimachia ciliata Ph. !, except author was L.
Troximon glaucum N[utt.] = Agoseris glauca (Pursh) Raf.

Literature Cited:
- Aedo, Carlos, 2001.
- Goodman, George J., and Cheryl A. Lawson, 1995.  

Goodman and Lawson (1995, p. 199-201) have a long account about this specimen and its proper name, reaching the conclusion that it should be G. intermedium James. Aedo (2001) acknowledges the name but accepts our G. caespitosum James.

 

Other articles:
• Golden Checklist Flora:  Geranium richardsonii;  

Geranium richardsonii Fisch. & Trautv. “Richardson's Geranium”

 

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.  

Hooker (1840, v. 1, p. 116) … Parts 2 and 3, pages 49-144 of Hooker's Flora of North America were issued in 1830, so even though the flora was not sonsidered complete until 1840, the pages including G. albiflorum Hook. were published in 1830. Meanwhile G. albiflorum Ledeb., of the Russian flora, was published in 1829, giving it priority.

Original Text
2. G. albiflorum ; caule subangulato erecto dichotomo inferne glabro superne piloso- glanduloso, foliis profunde 5-partitis ovato-acuminatis inciso-subpinnatifidis subpilosis, radicalibus longe petiolatis, superioribus oppositis breve petiolatis 3-partitis magis acurainatis, calycibus glanduloso-pilosis, petalis integris (albis) intus filamentis que basi hirsutis, (Tab. XL.)
Long Latin description omitted.
Hab. Vallies in the Rocky Mountains. Drummond. — This fine species of Geranium seems to hold a middle rank between the American G. maculatum and the Europaean G. pratense: yet it is, I feel assured, truly distinct from both. In the former, G. maculatum, the leaves are much less divided, the segments broader, and the peduncles are quite destitute of glandular hairs : in the latter, the leaves are much more deeply divided, the segments greatly narrower, and more truly pinnatifid. In both, the blossoms are purple ; in ours, the flowers are constantly white, even when cultivated, as the plant is in our gardens, and of a firmer texture. The G. longipes of De Candolle, according to his description, also approaches our plant.
Tab. XL. Fig. 1, Root-leaf: — natural size. Fig. 2, Petal; fig. 3, Stamen; fig. 4, Calyx and pistil, with the filaments of the stamens : — slightly magnified.
Parts 2 and 3, pages 49-144 of Hooker's Flora of North America were issued in 1830.

Literature Cited:
- Fischer, F. E. T., and Ernst Rudolph von Trautvetter, 1838.  

Fischer & Trautvetter (1838, p. 37) …

Original Text Translation and Comments
948. Geranium Richardsonii Fisch., Trautv. G. albiflorum Hook. fl. bor. amer. tab. 40, Bot. mag. tab. 3124. (non Ledeb.). Nomen erat mutandum ob G. albiflorum Ledeb. “The name was to be changed because of G. albiflorum Ledeb.”

 

Other articles:
• Golden Checklist Flora:  Oxalis stricta;  

Oxalis stricta L. “Common Yellow Oxalis”

 

 

Other articles:
• Golden Checklist Flora:  Linum lewisii;  

Linum lewisii Pursh “Prairie Blue Flax”

 

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 1, p. 210) ...

Original Text
Lewisii 2. L. foliolis calycinis ovatis acuminatis, petalis cuneatis apice rotundatis, foliis sparsis lanceolato-linearibus mucronatis, caulibus altis numerosis.
  In the valleys of the Rocky-mountains and on the banks of the Missouri. M. Lewis. ♃. July. v. v. Flowers large, blue; a very good perennial, and it probably might become an useful plant if cultivated.

 

Other articles:
• Golden Checklist Flora:  Linum perenne;  

Linum perenne L. “Blue Flax”

(Syn: Adenolinum perenne (L.) Rchb. )

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 277) ...

Original Text
2. LINUM calycibus capfulisque obtufis, foliis alternis lanceolatis integerrimis. perenne.
Linum foliis alternis lanceolatis integerrimis, calycibus apice obtifus, capfulis muticis. Hort. upf. 72.  
Linum perenne, ramis foliisque alternis lineari-lanceolatis. Hort. cliff. 114. Roy. lugdb. 434. Sauv. monfp. 53.  
Linum perenne majus caeruleum, calitulo majore. Morif. hift. 2. p. 573.  
Linum fylvestre caeruleum perenne erectus, flore & capitula majore. Raj. angl. 3. p. 362.  
Habitat in Sibiria et Cantabrigiae. ♃  

Other articles:
• Golden Checklist Flora:  Tribulus terrestris;  

 

 

Other articles:
• Golden Checklist Flora:  Chamaesyce fendleri;  

Chamaesyce fendleri (Torr. & A.Gray) Small “Fendler's Sandmat”

 

Literature Cited:
- Torrey, John, and Asa Gray, 1857b.

Locations: Big Spring.  

Torrey & Gray (1857) described Euphorbia fendleri in their report on the botany of the Pacific Railroad Survey. The collector was Dr. J. M. Bigelow. Dr. Gray, though, had already seen this plant among Fendler's collections.

Original Text Comments
Euphorbia Fendleri, (n. sp.): branching and diffuse from a somewhat woody caudex. smooth ; leaves stipulate, opposite, broadly ovate or orbicular-ovate, on very short petioles, subcordate and oblique at the base; involucres solitary, on short peduncles; gland transversely oval, with a narrow entire somewhat 2-lobed border; capsule smooth; seeds obovate, a little rugose transversely, gelatinous when moistened. Big Springs of the Colorado; April. This species is No. 800 of Fendler's New Mexican collection. It is a small plant, throwing off many branches that spread on the ground, forming a little patch from three to six inches in diameter. The leaves are 3-4 lines long, and are often of a purplish tinge, especially underneath. Big Springs of the Colorado is now Big Spring, Howard County, Texas. The creek is now called Beals Creek, and it joins the Colorado River some 35 miles to the southeast.

Literature Cited:
- Small, John Kunkel, and Per Axel Rydberg, 1903.  

Small (1903) made some nomenclatural changes without comment.

 

Other articles:
• Golden Checklist Flora:  Chamaesyce glyptosperma;  

Chamaesyce glyptosperma (Engelm.) Small “Ribseed Sand Mat”

 

Literature Cited:
- Torrey John, 1859.  

In Torrey, John (1859), the section on Euphorbiaceae was written by George Engelmann, M.D. That would explain why Dr. Engelmann is considered the author of a name published by Dr. Torrey. The botanist on the Mexican Boundary Commission was Dr. Charles Parry.

Original Text
Euphorbia glyptosperma (nov. spec.) : erecto patula seu demum decumbens ; foliis e basi valde obliqua (latere inferiore producta) aequilatis oblongis s. oblongo-linearibns obtusis versus apicem subserratis s. integriusculus ; stipulis setaceis laciniatis, anthodiis alaribus demum in glomerulos laxos laterales confertig ; appendiculis brevibus integris seu crenatis ; stylis brevibus apice bilobis, stigmatibus subglobosis ; seminibus ovatis argute rugosis ad angulos acutos crenatis. E. polygonifolia, Hook. FL Bar, Am. fide spec. auctoris non Linn.
β. tenerrima : foliis parvulis angustis apice vix crenulatis ; involucri minuti glandulis vix seu non appendiculatis. On the Rio Grande ; also on the Arkansas, and extending to the upper Missouri. (No. 1853, 1855, and 1856, Wright.) From a few inches to a foot high. The larger northern forms have leaves 3 to 6 lines long and 1 to 2 lines wide. In β the leaves are 1 to 3 lines long and ½ to 1 line wide ; involucrum in the latter only 0.3 line long. Seed very sharply cross-ribbed, similar to that of E. prostrata, and notched at the angles.

Wright is Charles Wright, a friend of Asa Gray, who collected in Texas and New Mexico in 1849 and 1851-53.

 

Other articles:
• Golden Checklist Flora:  Euphorbia myrsinites;  Euphorbia esula;  

Non-Native Euphorbia L. — Spurge

Several of our non-native Euphorbias as noxious weeds, e.g., E. myrsinites and E. esula, and are discussed among the Noxious Weeds, above.

 

Other articles:
• Golden Checklist Flora:  Euphorbia brachycera;  

Euphorbia brachycera Engelm. “Horned Spurge”

 

Literature Cited:
- Torrey John, 1859.  

George Engelmann described E. brachycera in Torrey (1859) citing a collection by Charles Wright in western New Mexico.

Original Text
Euphorbia brachycera (nov. spec.): annua (?), multicaulis, ramosa, erecta, glaberrima; foliis in petiolum brevissimum angustatis lanceolatis seu lineari-lanceolatis acutis mucronatis ; umbella 3-fida seu rare 4-5-fida; ramis pluries bifidis; bracteis inferioribus ovato-Ianceolatis superioribus rhombeo-orbiculatis mucronatis; glandulis brevissime obtuseque cornutis; seminibus majusculis ovatis maculis irregularibus saepe confluentibus leviter impressis. Western New Mexico; Wright, (No. 1821.) Many stems a foot high from a stout but apparently annual or biennial root; leaves 6-9 lines long, about 2 lines wide, patulous (not erect as in E. esulaeformis) regularly lanceolate ; upper part of the stem quite ramose ; horns shorter than in any of our species, and sometimes almost rudimentary. Seed 1 line in length, similar to those of E. esulaeformis, montana and Roemeriana.

 

Other articles:
• Golden Checklist Flora:  Euphorbia esula;  

Euphorbia esula L. “Leafy Spurge”

This plant has low severity poison characteristics. The sap contains diterpine esters in milky latex which is toxic on ingestion and highly irritant externally, causing photosensitive skin reactions and severe inflammation, especially on contact with eyes or open cuts. The toxicity can remain high even in dried plant material. Prolonged and regular contact with the sap is inadvisable because of its carcinogenic nature.

Linnaeus (v. 1, p. 461) notes that the plant is native to Germany, Belgium, and France.

 

Other articles:
• Golden Checklist Flora:  Euphorbia marginata;  

Euphorbia marginata Pursh “Snow on the Mountain”

 

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 2, p. 607) ...

Original Text
14. E. umbella 3-fida: bis dichotoma, involucro foliiforme, involucellis oblongis cordatis margine membranaceis coloratis, foliis lanceolato-oblongis subcordato-amplexicaulibus acutis glaucescentibus glabris, appendicibus calycinis petaloideis subrotundis, capsulis pilosis. marginata.
On the Yellow-stone river. M. Lewis. ☉. July. v. s. in Herb. Lewis. A very handsome species ; the white margin of the involucre and white petal-like appendices have a fine contrast with the elegant soft green leaves.  

   

Euphorbia myrsinites L. “Myrtle Spurge”

 

Other articles:
• Golden Checklist Flora:  Euphorbia myrsinites;  

Linnaeus (1753, v. 1, p. 461-462) ...

Original Text
50. EUPHORBIA umbella fuboctifida : bifida, involucellis fubovatis, foliis fpathulatis patentibus carnofis mucronatis margine fcabris. Diff. euph. 52. Myrfinites.
Euphorbia inermis, foliis fuperioribus reflexis latioribus lanceolatis, umbella univerfali trifida , partialibus bifidis. Hort. cliff. 199. Hort. upf. 141.  
Euphorbia inermis, foliis ligulatis fpinula terminatis, ad umbellam duodenis. bracteis trigonis fpinula terminatis. Sauv. monfp. 51.  
Tithymalus myrfinites latifolius. Bauh. pin. 296.  
Tithymalus myrfinites legittimus. Cluf. hift. 2. p. 189.  
Habitat in Calabria, Monfpelli. ♃  
Caules multi, pedales, diflexi, virides, inferme a cafu foliorum cicatrifati. Folia alterna, fpathulata, coriacea, concava, glauco-viridia, patentia, mucronata, margine fubscabra, fuperiora reflexa. Umbella 7-9 fida radiis femel bifidis. Involucrum univerfale foliolis 7-9, ovatis, tenuioribus, acutis ; Involucella diphylla, fubcordata, lariora, concava, acuta, margine fubfcabra. Flores intrainvolucella primaria & fecundaria mafculi, reliqui hermaphroditi. Calyces ore ferrati. Petala 4, flava, bicornia apicibus teretibus nitentia. Capufulae glabrae.  

Linnaeus does not mention toxicity of the species.

Locations given by Linnaeus are Calabria (Italy) and Montpellier (France). POWO shows the native distribution to be Italy south and east to Iran. Nativity or distribution in France is not mentioned.

 

Other articles:
• Golden Checklist Flora:  Euphorbia peplus;  

Euphorbia peplus L. “Petty Spurge”

 

 

Other articles:
• Golden Checklist Flora:  Euphorbia spathulata;  

Euphorbia spathulata Lam. “Warty Spurge” or “Spoonleaf Spurge”

Online (SEINet and CCH2) records contain determinations of the following synonyms:

  • Euphorbia arkansana
  • Euphorbia arkansana var. atrosemina
  • Euphorbia dictyosperma
  • Euphorbia obtusata
  • Tithymalus mexicanus
  • Tithymalus missouriensis
  • Tithymalus spathulatus

Literature Cited:
- Lamarck, Jean-Baptiste, 1786.  

Lamarck, Jean-Baptiste Pierre Antoine de Monet de [1744-1829] and Jean-Louis-Marie Poiret, Jean-Louis-Marie [1755-1834], 1786. Encyclopédie méthodique. Botanique . Paris,Liège : Panckoucke;Plomteux, 1783-1808. https://www.biodiversitylibrary.org/item/15260

Original Text Translation
56. Euphorbe fpatule, Euphorbia fpathulata. Euphorbia umbella trifida , dichotoma : foliis alternis oblongo-fpathulatis fubferratis. N. 56. Euphorbe spatule, Euphorbia spatulata. Euphorbia umbellum trifid, dichotomous: leaves alternate oblong-spathulate with subserrate. N
Euphorbia caule dichotomo , foliis ovato-oblongis , floribus fingularibus in fingulo dichotomiae finu feffilibus. Commerf. Herb. Euphorbia dichotomous stem , ovate-oblong leaves, flowers singular in each dichotomous branch, sessile.
Plante glabre , dont les tiges hautes d'un pied ou un peu plus , font munies de rameaux petits & alternes. Ses feuilles font aiternes , feffiles , oblongues-fpatulees , legerement dentelees , glabres , & a peine longues d'un pouce. L'ombelle eft compofee de trois rayons deux ou trois fois bifides. Les trois folioles de la collerette font ovales-oblongues ; les bractees leur reffemblent , mais elles font un peu plus petites & asuminees. Les capfules font glabres. M. Commerfon a trouve cette plante pres de Monte-Video. Elle paroit herbacee. ( v. f. ) Hairless plant, whose stems a foot or a little more high, are provided with small & alternate branches. Its leaves are alternate, thin, oblong-spatulate, slightly serrated, glabrous, and barely an inch long. The umbel is composed of three rays two or three times bifid. The three leaflets of the collar are oval-oblong; the bracts resemble them, but they are a little smaller & asuminees. The capsules are glabrous. Mr. Commerfon found this plant near Monte-Video. Herbaceous. (Seen in the dried state.)

 

Other articles:
• Golden Checklist Flora:  Edwin James collections;  Tragia ramosa;  

Tragia ramosa Torr. “Branched Noseburn”

 

Literature Cited:
- Torrey, John G., 1828.  

Torrey (1828, v. 2, p. 245) ...

Original Text
407. Tragia ramosa, caule herbaceo, piloso, ramosissimo ; foliis petiolatis, ovato-lanceolatis, argute serratis, subtus hirsutis, basi subcordatis ; racemis ♂ filiformibus, paucifloris, basi ♀
Desc. Stem about a span high, much branched, slender, hairy. Leaves alternate, ovate-lanceolate, coarsely serrate, with acuminate serratures, base acute or subcordate ; under surface hirsute, upper with a few appressed hairs; petioles one-third of an inch long, with subulate stipules at the base. Racemes axillary towards the upper part of the stem. Sterile flowers superior, 6-8, on short, filiform, bracteate pedicels. Calyx 4-parted, segments lanceolate, recurved, apex inflexed. Stamens 4. Pistil abortive. Fertile flowers. Calyx 5-parted. Stigmas 3, large, recurved. Capsule tricoccous, hispid. Seeds perfectly spherical, smooth.
Hab. Sources of the Canadian?
Obs. Resembles T. urticifolia, but differs in its narrow leaves, much branched, and smoother stem.

Literature Cited:
- Torrey John, 1859.  

Torrey (1859, v. 2, pt. 1, p. 200-201) ...

TRAGA RAMOSA, Torr. in Ann. Lyc. New York, 2, p. 245. T. angustifolia, Nutt. l. c. T. scutellariaefolia, Scheele^ l. c. Gravelly hiills of the Limpio; Bigelow. Ravines of the Organ mountains, April ; Parry. On the upper Rio Grande, New Mexico; Fendler, No. 776. Perhaps only a var, of the last. It differs chiefly in being much smaller and erect, with the leaves scarcely at all cordate. Var.? leptophylla : foliis linearibus integris vel remote denticulatis. — Near Howard's Springs; Bigelow; No. 1796, Wright. Plant woody at the base, about a span high, and branched from the base. Stem and branches sparingly hirsute. Leaves 1-1¼ inch long and 1-2 lines wide. Spikes few-flowered ; the lowest flowers fertile. Flowers as in T. ramosa, etc.

 

Other articles:
• Golden Checklist Flora:  Rhus glabra;  

Rhus glabra L. “Smooth Sumac”

 

 

Other articles:
• Golden Checklist Flora:   Rhus trilobata;
Full Size ImageRhus trilobata at the southern end of the Survey Field.  

Rhus trilobata Nutt. Squawbush.

Rhus L. is a Linnean name published in his Species plantarum in 1753, although the name had been previously used in six other works that Linnaeus cites. The genus is native to temperate and tropical Asia, Europe, Africa, the Pacific region, and North and South America. There are ten other names that are synonyms for Rhus including Lobadium Raf. that is now treated as a group within Rhus and contains our R. trilobata.

Literature Cited:
- Aiton, William, 1789.

Other articles:
• Glossary:  ♄;
Full Size ImageAiton (1789) description of Rhus aromatica  

Since Rhus trilobata is sometimes reduced to the rank of variety or subspecies under Rhus aromatica Ait. we need to understand the source of that name and its distribution. R. aromatica was described from a plant growing in the Kew Gardens, in 1789. The plant was provided to the garden in 1772 by John Bartram. The source of the plant was “Carolina,” which we now separate into North and South Carolina. Aiton's description is:

Mixed Latin and English My Interpretation of the Mixed Latin and English
11. R. foliis ternatis : foliolis sessilibus ovato-rhombeis inciso-serratis pilosiusculis.
Aromatic Sumach.
Nat. of Carolina. Mr. John Bartram.
Introd. 1772.
Fl. May.     H. ♄.
aromati-
cum.
11. R[hus] leaves in three parts : leaflets sessile, ovate-rhomboid, serrate, puberulent(?).
Aromatic Sumac.
Native to Carolina. [Collected by] Mr. John Bartram.
Introduced (to Kew Garden) in 1772.
Flowers in May.       Hardy. Shrubby.
aromatica
As was common at the time, Aiton used the astrological sign for Saturn to indicate shrubby plants.
Full Size Image
Abbreviations used by Aiton (1789)

Literature Cited:
- Graustein, Jeannette E., 1967.
- Torrey, John, and Asa Gray, 1838-1843.
Full Size ImageNuttall's description of Rhus trilobata in Torrey & Gray (1838).  

Rhus trilobata was published by Torrey & Gray (1838-1843) from a manuscript written by Nuttall. From the location “Rocky Mountains” and the date of publication — 1838 — it is likely Nuttall made his collection on the Nathaniel Wyeth expedition of 1834-1836 (Graustein, 1967, pp. 277-304).

It seems as though the author should be “Nutt. ex Torrey & A. Gray,” but no one seems to spell it out that way.

Rhus. ANACARDIACEAE 219
§ 3. Flowers dioecious or polygamous : disk glandular, deeply 5-lobed (lobes opposite the petals) : drupe globose, villous : nut smooth, compressed: flowers in short aments, preceding the leaves. Leaves 3-foliolate. — Lobadium, Raf.
8. R. aromatica (Ait.) …
9. R. trilobata (Nutt ! Mss.) : “leaves glabrous, small ; lateral leaflets obovate, obtuse, 3-lobed at the apex or nearly entire ; terminal leaflet cuneiform, 3-lobed at the summit, the middle lobe sometimes 3-toothed.
“In the central chain of the Rocky Mountains. — A low leafy shrub ; the leaves much smaller than in R. aromatica : terminal leaflet broad, ½–1  inch in length ; lateral ones smaller. Drupes scarlet, acid ; the nut flat, scarcely striate.” Nutt.

Graustein (1967), Nuttall's biographer, does not mention Rhus trilobata or Nuttall's collection of it.

Literature Cited:
- Watson, Sereno, 1871.

Other articles:
• Golden Checklist Flora:  Rhus trilobata, Barkley, 1937;

Locations: Raft River Mountains. Salt Lake Valley.  

The Geological Exploration of the Fortieth Parallel was a geological survey made by order of the Secretary of War under the direction of Brig. and Bvt. Major General A. A. Humphreys, Chief of Engineers, by Clarence King, U. S. geologist. More commonly known as the Fortieth Parallel Survey, the survey conducted field work from 1867 to 1872, exploring the area along the fortieth parallel north from northeastern California, through Nevada, to eastern Wyoming.

Volume 5, Botany, was written by Sereno Watson (1871) with the assistance of Drs. Gray and Torrey and others. Asa Gray reduced R. trilobata to the rank of variety, which was published in this volume by Watson.

Rhus aromatica, Ait., Var. trilobata, Gray. (R. trilobata, Nutt.) Leaves small, rarely exceeding 1' in length, usually glabrous ; leaflets lobed, the divisions entire or sparingly crenate. — Growing in dense leafy clumps, 3-6° high, and having a heavy disagreeable odor, It is the prevalent western form, extending from Western Texas to Southern California, and throughout the Rocky Mountains to the Upper Missouri. Found on the foot-hills around Salt Lake Valley and at the north base of the Raft River Mountains, Utah. (219.)

The Raft River Mountains are just northwest of the Great Salt Lake. The number (219) is the collection number that Watson applied when he distributed the specimens.

Full Size Image
Gray's description of Rhus aromatica var. trilobata in Watson (1871).

Literature Cited:
- Barkley, Fred Alexander, 1937.

Other articles:
• Golden Checklist Flora:  Rhus aromatica var. trilobata, Watson, 1871;  

Barkley (1937) treats both Rhus aromatica and R. trilobata at the species rank. R. aromatica var. trilobata Gray (in Watson, 1871) is placed in synonomy with R. trilobata. No comment is made regarding the treatment of either species, except that R. trilobata is an

... extremely variable assemblage of plants probably best treated taxonomically as a single polymorphic species running into several more of less consistent geographic variations…
Barkley also recognizes eight varieties of R. trilobata.

Literature Cited:
- Harrington, H. D., 1964, 2nd ed..  

Harrington (1964, 2nd ed.) accepts Rhus trilobata Nutt. Ex T. & G. and one variety, var. simplicifolia (Greene) Barkley.

Literature Cited:
- Weber, W.A., B.C. Johnston, and R. Whittmann., 1981.

Other articles:
• Golden Checklist Flora:  Rhus aromatica ssp. trilobata, Weber, 1989;  

Weber, et al. (1981) published Rhus aromatica Ait. subsp. pilosissima (Engelm.) W. A. Weber, without the similar new combination for R. trilobata. This will be corrected in Weber (1989).

Literature Cited:
- Weber, William A., 1989.

Other articles:
• Golden Checklist Flora:  Rhus aromatica, Weber, et al., 1981;  

Rhus aromatica Ait. Subsp. trilobata (Nutt.) W.A. Weber was published in Weber (1989) with the following explanation:

This is a combination inadvertantly omitted from an earlier treatment of Rhus (Weber, et al. 1981).
Neither publication gives any hint of why Weber reduces R. trilobata to a variety of Rhus aromatica

Literature Cited:
- Welsh, Stanley L., N. Duane Atwood, Sherel Goodrich, and Larry C. Higgins, 1993.  

Welsh et al. (1993) accept R. aromatica var. trilobata, and one variety, var. simplicifolia (Greene) Conq.

Literature Cited:
- Miller, Allison J., David A. Young, and Jun Wen, 2001.  

Miller, et al. (2001) paper about the phylogeny and biogeography of Rhus based on ITS sequence data is hidden behind a paywall.

Abstract. Rhus L. (sensu lato) has been considered the largest and most widespread genus in the Anacardiaceae. Controversy has surrounded the delimitation of the genus. Historically, seven segregate genera have been recognized: Actinocheita, Cotinus, Malosma, Melanococca, Metopium, Searsia, and Toxicodendron. These genera, together with Rhus s.str., are commonly referred to as the Rhus complex. Rhus s.str. Includes two subgenera, Lobadium (ca. 25 spp.) and Rhus (ca. 10 spp.). Sequences of the internal transcribed spacer (ITS) region of the nuclear ribosomal DNA were employed to examine the monophyly of Rhus s.str. And to provide insight into the phylogenetic and biogeographic history of the genus. The ITS data set indicates that Rhus s.str. Is monophyletic. Actinocheita, Cotinus, Malosma, Searsia, and Toxicodendron are distinct from Rhus s.str., although the relationships among these genera of the Rhus complex are not well resolved. Rhus subgenus Rhus is paraphyletic; the monophyletic subgenus Lobadium is nested within it. The ITS data set indicates that, for Rhus, the Madro-Tertiary floristic element (subgenus Lobadium) had a single origin within the Arcto-Tertiary floristic element (subgenus Rhus).

Literature Cited:
- Yi Tingshuang, Allison J. Miller, and Jun Wen, 2004.
- Yi, Tingshuang, Allison J. Miller, and Jun Wen, 2004.  

Phylogenetic and biogeographic diversification of Rhus (Anacardiaceae) in the Northern Hemisphere

Literature Cited:
- Yi, Tingshuang, Allison J. Miller, and Jun Wen, 2007.  

Yi, et al. (2007) published a similar paper on phylogeny of Rhus based on sequences of nuclear and chloroplast genetics found that

(1)Species of Rhus form a monophyletic group … (9) R. aromatica and R. trilobata are sister taxa.

With two notable exceptions … species of subgen. Lobadium were resolved into two clades: … and (2) R. aromatica–R. trilobata and R. integrifolia–R. ovata.

The present distributions of R. microphylla, R. aromatica–R. trilobata, R. lanceolata, and R. copallina indicate that the opportunity for hybridization between some or all of these species exist (and likely existed) in the southwestern U. S. and northern Mexico.

This is one of several phylogeny of Rhus papers that treat both R. aromatica and R. trilobata at the rank of species, although the relative rank of those two entities is not central to the purpose of those papers.

Literature Cited:
- Andres-Hernandez, A. R., and T. Terrazas, 2009.  

Andres-Hernandez (2009) published their analysis of leaf architecture of Rhus. This was part of their PhD thesis, and will be integral to a later paper (2014) on phylogeny of Rhus using structural and molecular data.

Literature Cited:
- Schalau, Jeff, and Gene Rwaronite, 2010.  

In their publication on wildfire risk reduction in Arizona's interior chaparral, Schalau & Twaronite (2010) list Rhus trilobata as suitable for use in eildfire survivable space.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012) accept R. aromatica Aiton ssp. trilobata (Nuttall) W. A. Weber, which Weber published in 1989, and one subspecies, ssp. pilosissima (Engelmann) W. A. Weber, which Weber published in 1981.

Literature Cited:
- Andres-Hernandez, Agustina Rosa, Teresa Terrazas, Gerardo Salazar, and Helga Ochoterena, 2014.  

For R. aromatica, the leaf texture is described as membraneous and the aureole is described as imperfect. For R. trilobata, the leaf texture is described as chartaceous and the aureole is described as incomplete.

This is another of several phylogeny of Rhus papers that treat both R. aromatica and R. trilobata at the rank of species, although the relative rank of those two entities is not central to the purpose of those papers.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015) accepts Rhus trilobata Nutt., placing R. aromatica Aiton in synonomy, and accepts three varieties: var. simplicifolia (Greene) Barkl., var. pilosissima Engelm., and var. trilobata.

Literature Cited:
- Idaho Firewise, 2017.  

Idaho Firewise (2017) lists Rhus trilobata as one of the best shrubs for Zones 1 and 2.

 

Kuhns & Daniels (2018) list Rhus trilobata as a recommended firewise shrub for Utah.

   

Toxicodendron Mill.

 

Literature Cited:
- Miller, Philip, 1754.  

Original Text
TOXICODENDRON, Poifon-tree, vulgo.
The Characters are ;

The Flower confifts of five Leaves, which are placed orbicularly, and expand in form of a Rofe ; out of whofe Flower-cup rifes the Pointal, which afterward becomes a roundifh, dry, and, for the moft part, furrowed Fruit, in which is contained one comprefs'd Seed.

The Species are ;

1. Toxicodendron triphyllum glabrum. Tourn. Three - leav'd fmooth Poifon-tree.

2. Toxicodendron triphyllum, folio finuato pubefcente. Tourn. Three- leav'd Poifon-tree, with a finuated hairy Leaf.

3. Toxicodendron rectum, foliis minoribus glabris. Hort. Elth. Upright Poifon-tree, with fmaller fmooth Leaves.

4. Toxicodendron rectum pentaphyllum glabrum, foliis latioribus. Smooth five-leav'd upright Poifon-tree, with broader Leaves.

5. Toxicodendron amplexicaule, foliis minoribus glabris. Hort. Elth. Climbing Poifon-tree, with fmaller fmooth Leaves.

6 Toxicodendron foliis alatis, fructu rhomboide. Hort. Elth. Poifon-tree with wing'd Leaves, and a Fruit fhap'd like a Rhombus.

The two firft Species were brought from Virginia, many Years fince, where they grow in great Plenty, as it is probable they do in moft other Northern Parts of America. The firft Sort feldom advances in Height ; but the Branches trail upon the Ground, and fend forth Roots, by which they propagate in great Plenty.
The fecond Sort will grow upright, and make a Shrub about four or five Feet high, but rarely exceeds that in this Country. This may be propagated by Layers, and is equally as hardy as the former.
The third Sort here mention'd grows erect to the Height of five or fix Feet : the Leaves of this Kind are much fmaller, than thofe of the common Poifon-oak ; but the Branches of this are flexible, fo that it will never make a Shrub of any great Height or Strength.
The fourth Sort was found in Maryland, from whence the Seeds were fent to England. This grows more upright than the former, and by the Appearance of the young Plants, feems to be a Shrub of much larger Growth.
The fifth Sort is a Native of Virginia, from whence I received the Seeds : this puts out Roots from the Branches, which faften themfelves to the Stems of Trees, or the Joints of Walls, by which the Branches are fupported.
The fixth Sort is a low Shrub, feldom rifing more than five Feet high. All the Sorts of Toxicodendron differ in Sex, the Male never producing any Fruit, having fmall herbaceous Flowers, without any Embryoes.
Thefe Plants are preferv'd by the Curious in Botany, for the fake of Variety ; but as there is little Beauty in them, they are not much cultivated in England. The Wood of thefe Trees, when burnt, emits a noxious Fume, which will fuffocate Animals when they are fhut up in a Room where it is burnt : an Inftance of this is mention'd in the Philofophical Tranfactions by Dr. William Sherard, which was communicated to him in a Letter from New-England by Mr. Moore, in which he mentions fome People who had cut fome of this Wood for Fuel, which they were burning, and in a fhort time they loft the Ufe of their Limbs, and became ftupid ; fo that if a Neighbour had not accidentally open'd the Door, and feen them in that Condition, it is generally believ'd they would foon have perifh'd. This fhould caution People from making ufe of this Wood.
All thefe Sorts are hardy Plants, which will thrive in the open Air in this Country ; but they love a moift Soil, and fhuld be planted under Trees in Wilderneffes, where they will thrive very well, and endure the Cold better than where they have a more open Expofure. They may be propagated by Seeds, or from Suckers, which fome of the Sorts fend forth in plenty, or by laying down the Branches of thofe Sorts which do not put forth Suckers ; which in one Seafon will be fufficiently rooted to tranfplant ; when they mould be planted where they are defign'd to remain. The beft Time to remove thefe Shrubs is in March, becaufe then there will be no Danger of their fuffering by Froft.
When a Perfon is poifoned by handling this Wood, in a few Hours he feels an itching Pain, which provokes a Scratching, which is followed by an Inflammation and Swelling. Sometimes a Perfon has had his Legs poifon'd, which have run with Water. Some of the Inhabitants of America affirm, they can diftinguifh this Wood by the Touch in the Dark, from its extreme Coldnefs, which is like Ice : but what is mention'd of this poifonous Quality, is applicable to the fixth Sort here mention'd ; which, by the Defcription, agrees with this Species.
This Sort of Poifon-tree is not only a Native of America, but grows plentifully on the Mountains in Japan, where it is called Fafi No Ki ; and from this Tree they extract one Sort of their Varnifh or Lacca, which they ufe for japanning their Utenfils : but this is not their beft Kind of Varnifh, that being made of the Juice of another Tree, which is alfo very poifonous, and nearly allied to this ; and from which fome Writers think it differs only by Culture.
The Juice of this Tree is milky, when it lffues oat of the wounded Part ; but foon after it is expofed to the Air, it turns black, and has a very ftrong fetid Scent, and is corrodeing : for I have obferved, on cutting off a fmall Branch from one of thefe Shrubs, that the Blade of the Knife has been changed black in a Moment's time, fo far as the Juice had fpread over it ; which I could not get off without grinding the Knife.
As this Tree is very common in Virginia, Carolina, and New-England, it would be well worth the Inhabitants Trial, to make this Varnifh.

Literature Cited:
- Greene, Edward L., 1905.  

Greene (1905, p. 114) segregated Toxicodendron from Rhus, again.

Segregates of the Genus Rhus
No taxonomic problem is easier, no fact more thoroughly established, than the identity of the original species, i.e., the type species of the genus Rhus ; because during more than a dozen centuries before even Tournefort, the species was but one, and that familiar to all writers about plants as the variously useful shrub of the whole Mediterranean region commonly called Rhus, but also long before Linnaeus written of under the binary name of Rhus coriaria, which name he also adopted. The genus was all this while supposed to be monotypical ; Rhus coriaria, the only Rhus. This fact is so easily apparent in bibliography, that there is no room for any controversy as to what is the type of the genus; and neither Tournefort nor Linnaeus, with the genus in view, could well have done otherwise than they did in placing it first in the list of species ; placing it as the type.
In the seventeenth century the genus received two indubitable accessions from North America in the shrubs now known as Rhus hirta and R glabra. Nobody questioned or doubted that these were of that genus. But along with these importations from our shores came the Poison Ivy ; a type which no authority did at first, or for a long time after, think of as possibly to be associated with Rhus congenerically.
Tournefort, before the end of the seventeenth century, proposed for the two forms known to him the rank of a genus, which he very fitly named Toxicodendron. Linnaeus suppressed the genus; but Philip Miller promptly restored it; and several more since Miller's time have insisted on its validity as a proper genus, so that now it bids fair for permanent recognition in the taxonomy of coming years.
A recension of the species of Toxicodendron is no easy task; so far from easy, I find it one of the most difficult I have hitherto undertaken. The best treatment of the genus extant, as to the early and typical species, is that of Dillenius in 1732. Linnaeus twenty years later, as his custom was, reduced the genus to Rhus and confused the species. Philip Miller sixteen years after this restored the genus, and also the Dillenian species of it, adding excellent descriptions of two or three new ones. In these two classic revisions of Tournefort's Toxicodendron, and not at all in Linnaeus, lie the means of identifying all the species early recognized.
The following represents my present understanding of the names and principal syonymy of the known species.

Greene goes on to nominate T. vulgare Mill. as the type of Toxicodendron.

 

Other articles:
• Golden Checklist Flora:  Toxicodendron rydbergii;  

Toxicodendron rydbergii (Small ex Rydb.) Greene “Western Poison Ivy”

 

Literature Cited:
- Rydberg, Per Axel, 1900a.  

Rydberg (1900, pp. 268-269) first recognized Western Poison Ivy as a distinct species in his Catalogue of the Flora of Montana and Yellowstone National Park.

* Rhus Rydbergii Small.
A single-stemmed shrub, less than a meter high, with grayish, somewhat striate bark ; leaves pinnately 3-foliolate with petioles 6-12 dm. long; leaflets 3-10 cm. long, broadly ovate, often somewhat rhomboid, rather thick, bright green, strongly veined beneath, glabrous except the veins on the lower surface, with wavy or sinuately toothed margins ; flowers in small conical axillary panicles, which are much shorter than the petioles, their branches short ; flowers yellow; petals about 3 mm. long, ovate, whitish yellow with greenish streaks; fruit when ripe white and shining,' a little depressed-globular, 5-6 mm. in diameter.
It has been invariably mistaken for R. Toxicodendron, which has pubescent sinuatelv lobed leaflets, and is confined to the southeastern United States. From R. radicans (R. Toxicodendron var. radicans), it differs in never being a climber, but always an erect shrub, in the thicker, somewhat glaucous leaves, the smaller and denser panicles, and the larger flowers and fruit. Both grow together in Nebraska, and they always remain distinct. The present species occurs on hillsides and in open woods, from Kansas to Arizona and British Columbia.
Montana: Great Falls, 1885, R. S. Williams, 291; northern Montana, F. W. Anderson.

Literature Cited:
- Greene, Edward L., 1905.  

Greene (1905, p. 117) provided a name for Rhus Rydbergii in Toxicodendron.

Original Text
T. Rydbergii. Rhus Rydbergii, Small, in Rydb. Fl. Mont. 268, in part. Well distinguished by Mr. Small, for the plant of Montana, occurring in Wyoming, mountain districts of Colorado, southward even to New Mexico, apparently, but hardly including that of Washington and Oregon.

 

Other articles:
• Golden Checklist Flora:  Acer glabrum;  

Acer glabrum Torr. Rocky Mountain Maple.

 

Literature Cited:
- Torrey, John G., 1828.  

Torrey (1828, p. 172) ...

Original Text
48. Acer glabrum, foliis subrotundis, 5-7-lobis, basi truncatis, lobis acute dentatis, utrinque glabris ; corymbis pedunculatis ; fructibus glabris, alis divergentibus, lato-ovatus.
Hab. On the Rocky Mountains.
Obs. Leaves in long petioles, green on both sides, very smooth on every part, slightly cordate at the base, 5-7-lobed ; the sinuses acute. Umbels on peduncles about a half an inch long. Fruit with very broad diverging wings.

There was but one specimen of this plant in the collection, which was in fruit. The flowers remain to be examined.

 

Other articles:
• Golden Checklist Flora:  Acer negundo L. “Box Elder Maple”;  

Acer negundo L. “Box Elder Maple”

(Syn: Acer negundo L. var. interius (Britton) Sarg., Negundo aceroides (L.) Moench )

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, vol. 2, p. 1056) ...

Negundo. 9. ACER foliis compofitis, floribus racemofis. Hort. cliff. 144. Gron. virg. 154. Roy. lugdb.460.
  Acer maximum, foliis trifidis & quinquefidis, virginianum Pluk. alm. 7. t. 123. f. 45.
  Habitat in Virginia. ♄

 

Other articles:
• Golden Checklist Flora:  Xanthoceras sorbifolium;  

Xanthoceras sorbifolium Bunge

First published as Xanthoceras sorbifolia Bunge, although the current preferred orthography is X. sorbifolium Bunge, probably something to do with proper Latin.

Literature Cited:
- Bunge, Alexander von, 1835.  

Bunge (1835, p. 85) ...

XXI. SAPINDACEAE
65 Xanthoceras Bge.
Char. gen.
Flores abortu polygamo-monoici.
Sepala quinque subaequalia obtusa.
Petala quinque hypogyna basi villosa.
Glandulae quinque petalis alternae lingulatae reflexae.
Stamina octo hypogyna.
Antherae apice glandula auctae biloculares intus dehiscentes.
Ovarium globosum triloculare, loculis octo-ovulatis.
Stylus simplex crassus.
Stigma capitatum trilobum.
Capsula corticata trilocularis trivalvis, valvis medio septiferis, dehiscens, polysperma.
Semina magna obovato-subglobosa, compressa.
Affinitas. Ab omnibus Sapindaceis loculis fructus multiovulatis differt.
Nomen a glandulis inter petala sitis cornuformibus flavis desumptum.
Xanthoceras sorbifolia Bge.
Hab. In montosis Chinae borealis rarior. Floret Aprili; Junio Julioque fructus maturat.
Arbor 10-15 pedes alta, foliis impari-pinnatis, foliolis serratis ; floribus terminalibus gemmaceis, racemosis, albis. Capsula magnitudine mali, trigono-subglobosa. Semina magna, nitida, nigra.
 

ChatGPT rewrote my prose above as follows:

“In a quiet alley of Golden, Colorado, nestled between a fence post and a telephone pole, lies a wondrous collection of Xanthoceras sorbifolium Bunge, more commonly known as the "Yellowhorn" tree. The remarkable discovery of this tree was made in 2023, despite it being several years old. The homeowner was surprised to find it there, as they had not planted it themselves nor were they even aware of its existence.

“Interestingly, this particular Yellowhorn tree is not the only one of its kind in Colorado. Two other collections have been reported in Logan County, one dating back to 1960 and the other all the way back to 1919. Furthermore, the Denver Botanic Garden is home to several specimens of the tree, including a hardy cultivar known as "'Psgan' Clear Creek," which was developed at Green Acres Nursery in Golden, Colorado.

“The name "Yellowhorn" is derived from the large orange nectaries found inside the base of the flower, and the leaves bear a striking resemblance to the "Mountain ash" of the Sorbus family. However, this beautiful tree has been given many other common names, including "Chinese Flowering Chestnut," "Flowering Yellowhorn," "Tree Goldenhorn," "Popcorn Shrub," and "Shiny Leaf Yellowhorn."

“Initially classified in the Sapinadaceae, or Soapberry Family, the Yellowhorn tree was later discovered to be a sister group to Sapindaceae s.l. after a recent phylogenetic analysis. This led to the suggestion that it should be placed in its own monotypic family, Xanthoceraceae. However, the Angiosperm Phylogeny Group currently places the tree in its own subfamily, Xanthoceratoideae Thorne & Reveal, within the Sapindaceae family. Despite some disagreements over its classification, one thing is certain: the Yellowhorn tree is a remarkable and fascinating species that continues to captivate botanists and nature lovers alike.”

 

Other articles:
• Golden Checklist Flora:  Ceanothus fendleri;  

Ceanothus fendleri A. Gray “Fendler's Ceanothus”

 

Literature Cited:
- Gray, Asa, 1849.  

Ceanothus fendleri A.Gray, Mem. Amer. Acad. Arts 29 (1849).

106. Ceanothus Fendleri (sp. nov.) : intricato-ramosissimus ; ramis ramulisque teretibus gracilibus saepe spinescentibus cinereo-puberulis demum glabratis laevibus ; foliis parentis (⅓-½ unc. longis) ovalibus scu ellipticis obtusis integerrimis eglandulosis trinerviis subtus sericeo-canescentibus supra glabriusculis viridibus ; glomerulis densis sessilibus ; floribus glabris albis. — Mountains east of Santa Fe, in sunny places ; June (in flower), and July, in fruit. Shrub about a foot and a half high and two feet in diameter. — Allied to C. depressus, Benth. Pl. Hartw. no. 29; but much more slender, the Thyme-shaped leaves smaller and not glandular, &c. Fruit about as large as in C. Americanus.

 

Other articles:
• Golden Checklist Flora:  Ceanothus herbaceus;  

Ceanothus herbaceus Raf. “Jersey Tea”

 

Literature Cited:
- Rafinesque, C. S., 1808.  

Ceanothus herbaceus Raf., Med. Repos. 5: 360 (1808).

19. Canothus herbaceus ; stem annual ; leaves ovate semi-serrated smooth, flowers in a terminal thyrsus ; grows near the falls of the Potowmack, between the rocks.

 

Other articles:
• Golden Checklist Flora:  Rhamnus cathartica;  

Rhamnus cathartica L. “European Buckthorn”

 

Literature Cited:
- Torrey, John, 1824.  

Torrey (1824) knew of the plant, and thought it to be introduced to America.

2. R. catharticus L. : spines terminal ; flowers generally 4-cleft, polygamo-dioecious ; leaves ovate ; stem erect ; berries 4-seeded. Willd. Spec. I. p. 1093. Smith Fl. Brit. I. p. 263. Eng. Bot. t. 1629. Roem. & Schult. V. p. 276.
A small tree or large shrub, with spreading branches and grayish bark. Leaves fasciculate and opposite, on petioles one-third their length, nervose, an inch and a half long ; margin erosely denticulate. Flowers clustered, on short axillary pedicels. Segments of the calyx lanceolate, recurved. Petals very small, yellowish-green. Stamens 4, a little longer than the flowers ; anthers didymous, oblong. Stigma 4-cleft. Berries black, nauseous and cathartic.
Hab. On mountains and in woods. Abundant in Williamstown, and other parts of New-England, apparently indigenous. Dewey. In the most retired parts of the Highlands of New-York. Barratt. Buck-thorn.
I suspect this plant has been introduced into this country at an early period, and that it is not really indigenous. Dr. Barratt informs me, that the old inhabitants of the Highlands remember to have seen it in gardens before it appeared in the woods.

Literature Cited:
- Knight, Kathleen S., Jessica S. Kurylo, Anton G. Endress, J. Ryan Stewart, and Peter B. Reich, 2007.  

Knight, et al. (2007) reviewed the characteristics of R. cathartica.

Physiological studies have uncovered traits including shade tolerance, rapid growth, high photosynthetic rates, a wide tolerance of moisture and drought, and an unusual phenology that may give R. cathartica an advantage in the environments it invades. Its high fecundity, bird-dispersed fruit, high germination rates, seedling success in disturbed conditions, and secondary metabolite production may also contribute to its ability to rapidly increase in abundance and impact ecosystems. R. cathartica impacts ecosystems through changes in soil N, elimination of the leaf litter layer, possible facilitation of earthworm invasions, unsubstantiated effects on native plants through allelopathy or competition, and effects on animals that may or may not be able to use it for food or habitat.

 

Other articles:
• Golden Checklist Flora:  Parthenocissus quinquefolia;  

Parthenocissus quinquefolia (L.) Planch. “Virginia Creeper”

 

 

Other articles:
• Golden Checklist Flora:  Parthenocissus vitacea;  

Parthenocissus vitacea (Knerr) Hitchcock “Woodbine, Thicket Creeper”

 

Literature Cited:
- Knerr, E. B., 1893.  

Knerr (1893, v. 18, 1893) ...

Notes on a variety of Ampelopsis quinquefolia— For the past ten years I have been observing a variety of the Virginia creeper that is quite marked by characteristic points of difference from the type species. Other students of botany must have noticed this variation and it is strange if nothing has been published upon the subject, but thus far I have failed to find anything1 . My attention was first called to the variation by neighbors who had transplanted the Virginia creeper from the woods but complained that it failed to cling to the side of the house. That was in the Miami valley in Ohio. I have since observed it in southern Iowa and in eastern Kansas. It does seem to me that the peculiarities are sufficient to distinguish it as permanent variety and it should be so recognized in our manuals.
In the first place the habit of growth is quite different from the type species. As is well known this latter climbs by clinging very closely to its support whether that be a tree or a wall. The variety does not cling so closely to its support. In fact it is'impossible for it to cling to a wall or even a tree unless the bark be very rough, owing to the structure of its tendrils. It climbs more like the grape and the clematis by trailing over low shrubbery to that which is higher, until it may reach the lower branches of a tree when it may rise to a considerable able height by reaching from branch to branch rather than by clinging close to the body of the tree and larger branches.
Sometimes in transplanting the Virginia creeper this variety is hit upon and the unobserving wonder why it fails to cling to the side of the house. On examination the tendrils will be found to be more like grape tendrils, long curling and grasping by recurved tips, rather than short, digitate and clinging by disk-like expansions as in the case of the typical species. The leaves also differ quite perceptibly, being much larger for the same age in the variation, and having longer petioles both for the leaf proper and for the leaflets. The margins are more distinctly serrate with larger teeth. The internodes of the stem are much longer in the variety, causing the leaves to be fewer and more scattered. The nodes are more swollen as are the leaf petioles at the base, making a much larger leaf scar, but the axillary buds are smaller.
The stem of the type species is quite rough, furrowed and warty especially as it grows older, while the variety is much smoother. The fruit ot the variety is more abundant, berries larger and in more open corymbs.
In short the whole aspect of the variety is more grape-like and for this reason I suggest the name A. quinquefolia, var. vitacea. — E. B. Knerr, Midland College, Atchison, Kansas.


1[See W. R. Lazenby in this journal xiii 233 and xv 233. —Eds.]

 

Other articles:
• Golden Checklist Flora:  Vitis riparia;  

Vitis riparia Michx. “Riverbank Grape”

 

Literature Cited:
- Hitchcock, A. S., 1894.  

Hitchcock (1894, p. 26) ...

PARTHENOCISSUS, Planch.
Calyx shortly -5 toothed. Petals expanding. Leaves palmately compound.
P. quinquefolia, (L.) Planch. Leaflets 5, clinging to rough supports by disk-like terminations. Woods, common. The older parts provided with aerial rootlets.
P. vitacea, (Knerr) Hitchc. Differs from the above in having no aerial rootlets; the canes smooth, and lighter colored; the tendrils dichotomous like the grape, and usually without disks; the inflorescence dichotomous rather than pinnate as in the former; flowering about two weeks earlier; the fruit maturing earlier and considerably larger. Woods, infrequent.

 

Other articles:
• Golden Checklist Flora:  Malva neglecta;  

Malva neglecta Wallr. “Common Mallow”

 

Literature Cited:
- Wallroth, Karl Friedrich Wilhelm, 1824.  

Wallroth (1824, v. 1, p. 140) ...

Original Text Comments and Interpretation
149.  
1. Malva neglecta Wallr.  
Char. spec. M. seminum arillis pubescentibus immarginatis dorso latusculisque laevigatis , calyculi foliolis linearibus , laciniis calycinis ovatis corolla duplo breviotibus receptaculum lenticulare attingentibus , pedunculis fructiferis declinatis , foliis suborbiculatis obsolete - 5 angularibus , caule diffuse prostrato.Flor. Dan. t. 721. Schk. t. 192. (opt.), Engl. bot. t. 1092.  
Syn. M. rotundifolia Linn. sp. pl. ex synon. Bauh. nec flor. suec. et herb. — Poll. Leyss. Hoffm. Leers . Willd. Schk. Roth. Spr. et omnium auct. Fl. germ. exclus. syn. Linn.  
M. sylvestris perennis decumbens, flore minore albo, folio rotundo Moris. hist. s. v. 521. t. 17. f. 7.  
M. sylv. folio rotundo C. Bauh. pin. 314. cum synonym.  
Patria: In suburbiorum areis desertis , juxta plateas spurcas , parietes , sepes , in quibus assurgit , in sterquiliniis et ruderibus abundat cum Lappis, Chenopodiis et Polygonis. Fl. Junio inde per totam aestatem. Hometown: In the deserted areas of outlying islands filthy streets , walls , hedges on which it rises, he abounds in manure and rubble Lappis, Chenopodius and Polygonis. Fl. June thence throughout the summer.
Descriptio: ... [omitted.]  

Karl Friedrich Wilhelm Wallroth (13 March 1792 in Breitenstein, Saxony-Anhalt – 22 March 1857 in Nordhausen) was a German botanist. His name is abbreviated Wallr. as a taxon authority.

 

Other articles:
• Golden Checklist Flora:  Sphaeralcea coccinea;  

Sphaeralcea coccinea (Nutt.) Rydb. “Scarlet Globemallow”

 

Literature Cited:
- Moulton, Gary E., 1999.

Locations: Marias River.  

Moulton (1999) presents two vouchers of Sphaeralcea coccinea from the Lewis & Clark herbarium. Both were of the same collection dated July 20, 1806, on the Marias River, Toole County, Montana. The material was separated and some stayed at the Americal Philosophical Society, while some was taken by Pursh to England and ended up in the Lambert Herbarium, which Pursh (1814) cited.

Literature Cited:
- Fraser, John, 1813.

Other articles:
• Golden Checklist Flora:  Fraser's Catalogue, publication details;  

Fraser's (1813) Catalogue, with at least some entries written by Nuttall, is the first publication of S. coccinea.

Original Text Comments
51 *Malva coccinea. ‡ Flowers scarlet, produced in dense spikes. Met with from the river Platte to the Rocky Mountains. The double dagger (‡) indicates that the plant is a perennial.

Literature Cited:
- Pursh, Frederick, 1816.  

Pursh (Vol. 2, 1814, p. 453-4) describes Cristaria coccinea from Fraser's (1813) Catalogue and the Lewis herbarium.

Original Text
553. CRISTARIA. Cavan. ic. 5. p 10.
1. C. undique cano-tomentosa et pilis stellatis obsita ; foliis 3-5-fidis, laciniis incisis acutis, racemis terminalibus, caule diffuso. coccinea.
Malva coccinea. Fraser. Catal.  
On the dry prairies and extensive plains of the Missouri. ♃. Aug. Sept. v. v. ; v. s. in Herb. Lewis. Flowers scarlet.  
Caluis teres, subpedalis, ramosus, ramis diffusis, tomentosis, pilis stellatis distinctis obsitis, uti tota planta. Folia alterna, petiolata, palmato 3-5 partita: laciniis sub-pinnatifida-incisis, lobis sinubusque acutis. Petioli teretes, longitudine folii. Stipulae lineares, erectae, marcescentes. Flores terminales, racemosi, coccinei, diametro pollicari. Calyx 5-fidus, laciniis lanceolatis acutis. Petala 5. oblique cuneato-obcordata, subunguiculata, ad ungues fasciculata-villosa, calyce duplo longiora. Filamenta monadelpha, numerosa, in columnam brevem connata, superne libera, patentia. Styli circiter 10. Fructus orbiculatus.  
The singularly stellated fasciculi of hair, placed in the close tomentum, and its bright scarlet flowers, make this plant particularly interesting.  

Pursh doesn't tell us where he saw this plant in the live condition, but we can guess that he saw it in Fraser's garden or that his benefactor, A. B. Lambert, was growing specimens obtained from Fraser's garden.

We wil

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Nuttall, 1818, publication details;  

Original Text
478. MALVA. L. (Mallow.)
Calix double; the exterior mostly 3-leaved. Petals 5. Capsules many, 1-seeded, disposed orbicularly.
Shrubby or herbaceous; leaves alternate and stipulate, undivided, or palmately lobed; flowers axillary or terminal, solitary, more or less aggregated or racemose. (Pubescence stellate.)
Species. 1. M. abutiloides. 2. caroliniana. A Sida? 3. rotundifolia Introduced. 4. triloba. In Carolina. 5. * coccinea. T. N. in Fras. Catal. 1813. A very beautiful species with scarlet flowers disposed in dense recemes (sic); outer calix wanting, leaves mostly trifid, canescently tomentose. Hab, From the confluence of the river Platte and the Missouri, often extending over the plains in such quantities as to communicate a brilliant redness to thousands of acres. This plant has no sort of affinity to Cristaria, and by the fruit is a genuine Malva.
A genus of more than 60 species, many of them indigenous to tropical America, to the Cape of Good Hope, and some to Europe.

Literature Cited:
- Gray, Asa, 1849.

Other articles:
• Golden Checklist Flora:  Gray, 1849, publication details;  

Original Text
81, 82. Malvastrum coccineum : humilis, incanum ; cailibus e basi sublignosa ramosis diffusis ; foliis trisectis tripartitisva, segmentis lateralibus saepius bipartis intermedio trifido, lobis oblongis linearibusve integris seu paucidentatis ; floribus inferioribus solitariis rariusve geminis in axillis foliorum, sperioiribus in racemum strictum digestis ; bracteis subulatis deciduis ; bracteolis involucelli 1-2 setaceis fugacibus ; coccis 9-12 clausis dorso subtiberculatis cano-tomentosis. — Gray, Gen. Ill. t. 219. Cristaria coccinea, Pursh ! Fl. 2. p. 453. Malva coccinea, Nutt. ! Gen. 2. p. 81 ; Bot. Mag. t. 1643. Sida coccinea. DC. Prodr. 1. p. 465 ; Hook. Fl. Bor.-Am. 1. p. 109; Torr. & Gray ! Fl. 1. p. 235. — Var. β dissectum : nanum ; foliis 3-5-partitis, segmentis 3-5-fidis lobisque anguste linearibus. Sida dissecta, Nutt. ! in Torr. & Gray, l. c.; Hook. & Arn. ! Bot. Beech. Suppl. p. 327. — Plains, &c., Santa Fe, and East to Rock Creek and Poñi Creek of the Canadian ; June to September. In flower and fine fruit. Some varying forms as to foliage are distributed under these numbers ; under one of them there are a few specimens of the var. dissecta, or of forms that evidently collect the Sida dissecta of Nuttall with the S. coccinea. I have for several years cultivated this species, from seed brought from the Upper Missouri by Mr. Sprague, and had ascertained that its radicle is inferior, as in Malva. — An account of the genus, of which this is one of the typical species, is given in the subjoined revision of the genera allied to Malva.  

Literature Cited:
- Rydberg, Per Axel, 1913.  

Rydberg (1913) argued that Gray's (1849) Malvastrum should be merged into Sphaeralcea.,

Original Text
Sphaeralcea coccinea (Nutt.) Rydb.

Malva coccinea Nutt. Fras. Cat. 1813.
Cristaria coccinea Pursh, Fl. Am. Sept. 454. 1814.
Sida coccinea DC. Prod. 1: 465. 1824.
Malvastrum coccineum A. Gray, Mem. Am. Acad. 4: 21. 1849

 

Other articles:
• Golden Checklist Flora:  Hypericum perforatum;  

Hypericum perforatum L. “Common St. John's Wort”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 2, p. 785) …

Original Text
14. HYPERICUM floribus trigynis, caule ancipiti, foliis obtufis pellucido-punctatis. Hort. cliff. 380. Fl. fucc. 635. Mat. med. 368. Roy. lugdb. 474. Hall. helv. 630. Dalib. parif. 233. perforatum.
Hypericum vulgare. Bauh. pin. 279.  
Hypericon. Dod. pempt. 76.  
Habitat in Europae pratis. ♃  

 

Other articles:
• Golden Checklist Flora:  Hybanthus verticillatus;  

Hybanthus verticillatus (Ortega) Baill. “Baby Slippers”

 

Literature Cited:
- Ortega, Casmiro Gomez, 1797.  

Ortega (1797, 4, p. 50) described Viola verticillata from plants grown at the Royal Gardens of Madrid from seeds sent from Mexico by D. Sesse.

Original Text Comments
Habitat in Nova Hispania. Habitat in Mexico.
Floret mense Septembri in Horto Reg. Matrit. e seminibus missis per D. Sesse. Flowers in Septermber in the Royal Garden of Madrid from seeds sent by D. Sesse.

Literature Cited:
- Baillon, Henri, 1867-95.  

Henri Baillon (1867-95, v. 4 [1873], pp.344-345) writing about the Violet family – Violacees &ndash places a group of plants with similar characteristics in Hybanthus.

Original Text Comments
Les propriétés des plantes de cette famillle sont assez homogenes. Leurs racines sont vomitives, à un faible degre dans les especes europeennes, a un degre assez prononce dans les especes de l'Amerique australe, pour qu'on les ait souvent employees comme Faux-Ipecacuanhas. La plus célèbre, a cet égard, est la plante qui donne le Faux-Ipecacuanha du Brésil et de la Guyane, médicament très-employé dans som pays natal aux memes usages 6 que les vrais Ipecacuanhas, auxquels on le substitute frequemment ; cette espece devra sans doute prendre le num d'Hybanthus Ipecacuanha La racine de Cuichunchilli ou Cuchunchully du Perou, autre vomitif puissant, appartient a une seconda espece du meme genre, l'H. microphyllus8 The properties of the plants of this family are quite homogeneous. Their roots are emetic, to a low degree in European species, to a fairly pronounced degree in species from southern America, so that they were often used as Faux-Ipecacuanhas. The most famous, in this respect, is the plant which gives the False-Ipecacuanha from Brazil and Guyana, a drug widely used in his native country with the same 6 uses as the real Ipecacuanhas, to which it is frequently substituted; this species will undoubtedly have to take the name of Hybanthus Ipecacuanha The root of Cuichunchilli or Cuchunchully from Peru, another powerful vomit, belongs to a second species of the same genus, l H. microphyllus 8
[p. 344, Footnote 6] 6. Evacuant, vomitif, purgatif, antidysenterique, etc.; il renferme de l'emetine. 6. Evacuating, emetic, purgative, anti-dysentery, etc .; it contains ememetine.
Les H. scandens, … verticillatus1 … quoique moins connus, aont autant d'especes decrites comme appartenant au genre Ionidium, et qui, possedant des proprietes vomitives plus ou moins accentuees, sont employees comme Ipecachuanhas faux ou blancs dans les regions les plus chaudes de l'Amerique. The H. scandens , … verticillatus 1 … although less known, have as many species described as belonging to the genus Ionidium , and that, possessing more or less accentuated emetic properties, are employed as false or white Ipecachuanhas in the hottest regions of America.
[p. 345, Footnote 1] 1. Viola verticillata Orteg., Dec., IV, 50 — Solea verticillata Spreng., in Schrad. Journ., II (1800), 190, t. 6. — Ionidium polygalaefolium Vent., Malmais., t. 27. — DC., Prodr., a. 13. — H. B. K., Nov. gen. et spec., V, 376, t. 496 (Mexique et Antilles).  

 

Other articles:
• Golden Checklist Flora:  Viola canadensis;  

Viola canadensis L. “Canadian White Violet”

(Syn: Viola rydbergii Greene, Viola scopulorum (A. Gray) Greene)

 

Other articles:
• Golden Checklist Flora:  Viola nuttallii;  

Viola nuttallii Pursh “Nuttall's Violet”

 

 

Other articles:
• Golden Checklist Flora:  Viola odorata;  

Viola odorata L. “Common Sweet Violet”

 

 

Other articles:
• Golden Checklist Flora:  Viola pedatifida;  

Viola pedatifida G. Don “ Prairie Violet”

 

Literature Cited:
- Don, George, 1831-1838.  

Don, George (1831, v. 1, p. 320) ...

3. V. pedati'fida : leaves pedately 3-parted, middle partition trifid, lateral ones all bifid, all lobed. Very like V. pedata, but differs in the two lateral petals being bearded. ♄ H. Native of North America. Flowers beautiful blue.
Pedatifid-leaved Violet. Fl. May, June. Clt. 1826. Pl. ¼ ft.

 

Other articles:
• Golden Checklist Flora:  Mentzelia albicaulis;  

Mentzelia albicaulis (Douglas ex Hook.) Douglas ex Torr. & A. Gray “White-Stem Blazing Star”

 

Literature Cited:
- Hooker, Sir William Jackson, 1829-1840.  

Hooker (1834, v. 1, p. 222) …

Original Text
3. B[artonia] albicaulis; petalis 5 obovatis (parvis), staminibus petaloideis nullis, bracteis nullis, caule laevissimo nitido albo. Douglas, MSS. apud Hort Soc. Lond. — Mentzelia albicaulis. Douglas, in Herb. nostr.
Long Latin description omitted.
Hab. On arid sandy plains of the river Columbia, under the shade of Purshia tridentata. Douglas. — Mr. Douglas seemed to be doubtful whether this new plant should be referred to Bartonia or Mentzelia, It partakes, indeed, in some characters of the one, and some of the other. Mr. Nuttall, who has perhaps alone of any living Botanist had the opportunity of examining both Bartonia and Mentzelia in a recent state, and who may be considered to have established the former genus, says that “ nothing essential separates Bartonia from Mentzelia, except the augmentation of petals, and the structure of the capsules and seeds.” In regard to the augmentation of petals, Mr. Nuttall himself observes that in Bartonia their number varies from ten to fifteen. We have shown in the two preceding species, that it is reduced to five, but with five petaloid stamens, or imperfect petals. In the present plant, the petaloid stamens are wanting ; and the number of petals exactly accords with Mentzelia. Again, in Bartonia, the seeds ought to be, according to Nuttall, “ numerous, arranged horizontally in a double series ;” in Mentzelia, the “ seeds are 3-6, longitudinally arranged.” In our plant, the seeds are numerous (25-40, Douglas), placed in a single series. — For a knowledge of the structure of the flowers and seeds of this plant, I am indebted to a beautiful drawing which accompanied the specimen lent to me by Professor Lindley, and made by that admirable Botanist.

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & A. Gray (1840, v. 1, p. 534) ...

Original Text
6. M. albicaulis (Dougl. mss.) : low, branching from the base; stem white and polished and nearly glabrous below ; leaves lanceolate, remote, deeply sinuate-pinnatifid, sessile, scabrous ; flowers (small) solitary or mostly in loose clusters, not bracteolate ; petals obovate, scarcely exceeding the short subulate-lanceolate calyx-segments ; filaments (20-30) all subulate-filiform ; capsules cylindrical, narrow, elongated ; seeds 20-40. — Bartonia albicaulis, Hook. ! fl. Bor.-Am. 1. p. 222. Trachyphytum albicaulis & T. gracile ! Nutt. mss.
Arid sandy plains of the Oregon, under the shade of Purshia tridentata, Douglas. Plains of the Oregon and Rocky Mountains, Nuttall ! June. — Stem 6-10 inches high. Petals about 2 lines in length, yellow. — The pubescence, although less in degree, is of two kinds, like that of M. Lindleyi, &c.

 

Other articles:
• Golden Checklist Flora:  Mentzelia dispersa;  

Mentzelia dispersa S. Watson “Bushy Blazing Star”

 
 

Watson (1871, v. 5, p. 114) ...

Original Text
Var. integrifolia. Usually branched ; leaves ovate, oblong or oblong- lanceolate, entire or sinuately toothed or pinnatifid ; flowers and fruit as in the ordinary M. albicaulis, but the seeds rather few (6—20) and some of the 15-20 filaments occasionally petaloid. — There seems very little to separate this from M. micrantha, H. & A., which has the same habit and foliage and is by no means constant in having a 3-seeded capside, having sometimes half-a-dozen or more seeds. This is 663 Geyer, 571 Hall & Harbour, and 195 Vasey. East Humboldt Mountains, Nevada, and Antelope Island and the Wahsatch, Utah; 4,500-0,000 feet altitude ; June, July. (430.)

Literature Cited:
- Watson, Sereno, 1871.
- Watson, Sereno, 1876.  

Watson (1876, v. 11, n.s., v. 3, p. 137) ...

Original Text
Mentzelia dispersa. A slender annual, usually about a foot high ; leaves narrowly lanceolate, sinuate-toothed or sometimes entire, rarely pinnatifid, the uppermost often ovate: flowers small, mostly approximate near the ends of the branches; calyx-lobes a line long, little shorter than the five spatulate or obovate petals : filaments not dilated : capsule narrowly linear-clavate, six to nine lines long : seeds very often in a single row, angular and somewhat rhombohedral, more or less grooved upon the angles, very nearly smooth, half a line long. — M. albicaulis, var. integrifolia, Watson, Bot. King's Rep. 114. From Washington Territory to Colorado and southward, frequent ; Yosemite Valley, Bolander ; Guadalupe Island, Palmer. Much resembling M. albicaulis, Dougl., with which it has been confounded almost from the first, but which is distinguished by its more pinnatifid leaves and slightly larger flowers, and especially by its rather strongly tuberculate seeds, irregularly angled with obtuse margins. The rarer allied species M. micrantha differs in its more leafy habit and small ovate leaves, and in its shorter, broader and few-seeded capsules, the seeds a line long.

 

Other articles:
• Golden Checklist Flora:  Mentzelia multiflora;  

Mentzelia multiflora (Nutt.) A. Gray. “Adonis Blazing Star”

 

Literature Cited:
- Nuttall, Thomas, 1818.

Other articles:
• Golden Checklist Flora:  Nuttall, 1818, publication details;  

 

Literature Cited:
- Nuttall, Thomas, 1848b.

Other articles:
• Golden Checklist Flora:  Nuttall, 1848, publication details;  

Nuttall (1848, p. 180) plants collected by William Gambel.

Original Text
BARTONIA.
B. *multiflora. Biennial ? Stem smooth, white and shining, corymbosely branched ; leaves narrow-lanceolate, sinuate, pinnatifid, attenuated below and sessile ; flowers subtended by one or two linear bractes; petals 10, oblong-oval, obtuse; capsule urceolate, with three to four valves ; segments of the calyx long and subulate ; seeds in a double series, winged.
Hab. Sandy hills along the borders of the Rio del Norte. Santa Fe, (Mexico.) Flowering in August.

Literature Cited:
- Gray, Asa, 1849.

Other articles:
• Golden Checklist Flora:  Gray, 1849, publication details;  

Gray (1849) placed the plant in Mentzelia in his Plantae Fendleriana.

Original Text
242. M. multiflora. Nutt. Pl. Gamb. in Jour, Acad, Philad. n. ser. 1. p. 180? under Bartonia. Santa Fe ; June, July. Flower seldom open in the day-time. Also along the Arkansas near Bent's Fort ; Sept. (I have not seen the latter specimens.) — The specimens are a foot of more in height, with most of the leaves rather deeply pinnatifid, and flowers about one fourth larger than those of an original specimen of Nuttall's Bartonia pumila in Dr. Torrey's herbarium. with which, except in the greater size, they accord tolerably well. The stems become bright white with age, as in other species. The petals apparently straw-color, perhaps white, are one half or one third of an inch in length, obovate or spatulate and rather obtuse, except with age, exceeding the subulate calyx-segments, and longer than the ovary. The outer filaments are conspicuously dilated. In have the same species, apparently, from Coulter's Californian collection, but with rather larger flowers, and mire interruptedly pinnatifid leaves. In Dr. Torrey's herbarium I notice specimens, for the most part imperfect, referable either to this plant or to M. pumila itself, gathered on the Upper Platte or Arkansas by Dr. James in Long's expedition, and in the valley of the Rio del Norte, New Mexico, by Lieut. Emory. I can hardly doubt that it is the Bartonia multiflora of Nuttall, although the petals are not quite so large as he describes them. There is a related species in Texas, which I have also flowered in cultivation.* This last species is M. wrightii A. Gray s. nov., which Gray goes on to describe.

Literature Cited:
- Hufford, Larry, John J. Schenk, and Joshua M. Brokaw, 2017.  

Hufford, et al. (2017) retain all of Mentzelia s.l. in a single genus, and place M. multiflora in section Bartonia.

 

Other articles:
• Golden Checklist Flora:  Mentzelia nuda;  

Mentzelia nuda (Pursh) Torr. & A. Gray “Bractless Blazing Star”

 

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 1, p. 328) …

Original Text
nuda. 2. B. germine nudo, seminibus alatis. Sims l. c.
  On the banks of the Missouri. Nuttall. ♂. July, Aug. v. v. This species has smaller flowers, and the leaves are not so glaucous as the foregoing : in every other respect the above description is applicable to the present one, with little deviation. The number of petals is variable, from to to fifteen.

The “foregoing” is Bartonia ornata Pursh. Pursh placed B. decapetala Sims in synonomy, which we now know as Mentzelia decapetala (Sims) Urb. M. decapetala is not known from Jefferson County; collections are from Boulder to the north and east, and Colorado Springs to the south and east.

Pursh's Index Auctorum has two entries for Sims.

  • Annals of bot. — Annals of Botany, by C. Konig and J. Sims. 2 vols. London, 1805, 1806. 8vo.
  • Bot. mag. — Curtis's Botanical Magazine, continued by J. Sims. London, 1801 et seq. 8vo.

Literature Cited:
- Torrey, John, and Asa Gray, 1838-1843.  

Torrey & Gray (1838, v. 1, p. 535) ...

Original Text
9. M. nuda : rough with a minute barbed pubescence ; leaves somewhat lanceolate, interruptedly pinnatifid, the segments obtuse ; flowers (large) terminating the branches, not bracteolate ; petals 10; stamens very numerous; the exterior filaments petaloid and often sterile, the others filiformn ; capsules 3-valved at the summit ; seeds numerous, winged. — Bartonia nuda, Nutt l. c. ; Pursh, Fl. 1. p. 328, & 2. p. 749 ; DC. I. c.
Gravelly hills of the Missouri, near the Great Bend, Nuttall, Dr. .James ! July-Aug. — We have drawn up the character of this, as well as of the preceding species, wholly from the excellent and complete description of Nuttall, as we possess only an imperfect specimen collected by Dr. James. The flowers are about half the size of those of M. ornata ; and the root, Mr. Nuttall thinks, is triennial or perhaps perennial.

The preceding was M. ornata now treated as a synonym of M. decapetala.

 

Literature Cited:
- Osterhout, George F., 1901b.

Other articles:
• Golden Checklist Flora:  Mentzelia speciosa;  

Mentzelia speciosa Osterh. “Plains Blazing Star”

 

Literature Cited:
- Osterhout, George F., 1901a.  

Osterhout (1901, v. 28, n. 11, p. 644) ...

Original Text
Mentzelia aurea
Perennial, the stem whitish, rather stout, finely pubescent, glandular on the pedicels and ovary ; corymbosely branched from near the base, the leaves linear to oblong, sinuate-dentate to sinuate-pinnatifid, the lower including the narrowly winged petiole, 10 to 15 cm. long, hispid on both sides : flowers numerous, vespertine, the earlier ones sessile, the others on pedicels 1-2 cm. long : petals 10, the five outer ones whitish on the outside, 2 cm. long and 8 mm. wide at the widest portion, narrowed to a claw which is nearly half the length of the whole petal ; the five inner ones narrow and not more than half the size of the outer, a number of the outer filaments also petaloid : the capsule a little more than 2 cm. long, striate: the seeds round, winged, mature ones smooth, i. e., not punctate.
Type specimens collected in Estes Park, Larimer county, Colorado, July 18, 1900, no. 2203. A. showy plant, ranging from the edge of the foothills to an altitude of 7000 feet. Before the flowers open they appear to be white on account of the whitish outside of the petals, but when open they are golden yellow.
 

Osterhout (1901, v. 28, n. 12, p. 689) ...

Original Text
A corrected Name
Mentzelia speciosa
Mentzelia aurea Osterhout, Bull. Torr. Club, 28 : 544.
Not Mentzelia aurea Nutt. Gen. 1 : 300 which is now a synonym Mentzelia oligosperma Nutt.
George E. Osterhout.

The 5 in the page number 544 has been crossed out and a 6 pencilled in.

   

Coryphantha (Engelm.) Lem.

 

Literature Cited:
- Engelmann, George, 1856.  

Engelmann, George. 1856. Synopsis of the Cactaceae of the Territory of the United States and Adjacent Regions. Proceedings of the American Academy of Arts and Sciences. v. 3 (1852-1857), p. 259-314.

p. 260 CACTACEAE  
p. 260 Tribus I. TUBULISAE, Miquel.  
p. 260 Subtrib. 1. Parallelae. Cotyledones margine hilum versus spectantes, lateribus seminis parallelae.  
p. 264 Subgen. 2. Coryphantha Flores e basi tuberculorum hornotinorum aculeiferorum sulcatorum, vel in vertice ipso oriundi : ovarium emersum. Flowers from the base of hornet's tubers of the prickly furrows, or at the very top of the rising : ovary emerged.
p. 264 § 2. Flaviflorae.  
p. 264 * Laxiflorae. (The originally central flowers are pushed aside by the continuous development of new tubercles.)  
p. 264 15. M. Nuttallii, E.; [Latin diagnosis …]  
p. 264 Var. α borealis (M. Nuttallii, E. l. c. Cactus mamillaris, Nutt. Gen., non Linn.): subsimplex ; aculeis setaceis 13-17 cum centrali saepe deficiente puberulis ; stigmatibus 2-5 ; baccis seminibusque minoribus.  
p. 268 § 3. Rubriflorae.  
p. 268 * * Sepalis fimbriatis  
p. 269 28. M. vivipara, Haw. …  

Literature Cited:
- Lemaire, (Antoine) Charles, 1868.  

Lemaire (1868) elevated Engelmann's subgenus of Coryphantha to genus.

3e Genre. — Les Coryphanthes (Coryphantha). 3rd Kind. — The Coryphanthes (Coryphantha)
ÉTYMOLOGIE. ETYMOLOGY.
Coryphè, tête, sommet; anthè, fleur, allusion à l’insertion des fleurs. Coryphe, head, apex; anthè, flower, allusion to the insertion of flowers.
Sous ce nom, l’excellent cactographe, M. Engelmann, a séparé du genre Mamillaria, en en faisant un sous-genre, les espèces pourvues sur les podaires (en dessus) d’une fossule longitudinale plus ou moins profonde, à la base de laquelle naissent les fleurs. Ce caractère, qui ne se remarque plus que chez les Echinocactes, ainsi que ceux que nous allons énumérer, nous ont engagé à admettre le Coryphantha, comme suffisamment distinct du Mamillaria. Under this name, the excellent cactographer, Mr. Engelmann, separated from the genus Mamillaria, making it a subgenus, the species provided on the pods (above) a more or less deep longitudinal fossula, at the base from which the flowers are born. This character, which is no longer noticeable except in the Echinocacti, thus that those we are going to enumerate, have committed us to admit the Coryphantha, as sufficiently distinct of Mamillaria.

   

Echinocereus Engelm. “Hedgehog Cactus”

 

Literature Cited:
- Engelmann, George, 1848.  

Engelmann(1848, p. 91) published echinocereus ....

Original Text
In the prairies about Wolf creek, in an elevation of between 6,000 and 7,000 feet, the smallest of a tribe of cactaceae was discovered, numerous species of which were found in the course of the journey south and southeast several others have also been discovered in Texas. I mean those dwarfish Cerei, some of which have been described with the South American genus Echinopsis, or have been referred alternately to Cereus or Echinocactus, and which I propose to distinguish from all these under the name of Echinocereus,7 indicating their intermediate position between Cereus and Echinocactus: they approach more closely to Cereus, in which genus they, as well as the genus Echinopsis, should perhaps be included as subgenera.

7 Echinocereus, n. gen. Perigonii tubus ultra germen productus ... [... Latin diagnosis omitted ...]

Globose, or mostly ovate; simple, or mostly branching from the base or cespitose; tubercles, forming few or mostly a great many ribs; bunches of short or long spines, distant or approximate, often very crowded; vertex never woolly; flowers lateral, produced from last year's growth, opening only in sunshine, but for two or three days in succession; closed at night, or in dark weather.

 

Other articles:
• Golden Checklist Flora:  Echinocereus viridiflorus;  

Echinocereus viridiflorus Engelm. “Nylon Hedgehog Cactus”

 

Literature Cited:
- Engelmann, George, 1848.

Locations: Wolf Creek.  

Engelmann then proposed E. viridiflorus ...

Original Text
The species mentioned above is distinguished from all others known to me by its yellowish green flowers, the others having crimson or purple flowers. I have named it, therefore, Echinocereus viridiflorus.8

8 Echinocereus viridiflorus, n. sp. ovato-globusus, humilis, … [… Latin diagnosis omitted …]

Prairies on Wolf creek, flowers in June; Santa Fe, flowers in May, (Fendler.) Body 1 to 1½ inch high, oval; spines 1 or 1½ to 3 lines long; central spine when present 6 to 7 lines long; flower 1 inch long and wide, outside green brown, inside yellowish green; petals only 2 lines wide, being about 5 lines long.

 

Literature Cited:
- Sanchez, Daiel, Balbina Vazquez-Benitez, Monserrat Vaquez-Sanchez, David Aquino, Salvador Aria, 2022.  

Escobaria Britton & Rose

 

 

Other articles:
• Golden Checklist Flora:  Escobaria;  

Escobaria Britton & Rose

 

Literature Cited:
- Britton, Nathaniel Lord, and J. N. Rose, 1919-1923.  

Britton & Rose (1919-1923, v. 4, p. 53) published the genus Escobaria to distinguish the cacti from the Coryphtanthae.

8. ESCOBARIA gen. nov.
Globose or cylindric, usually cespitose cacti, never milky; tubercles grooved abve, persisting as knobs at the base of old plants after the spines have fallen; spines both central and radial, never hooked; flowers small, regular, appearing from the top of plant at bottom of groove of young tubercles; stamens and style included; fruit red, naked (or with one scale), indehiscent, globular to oblong, crowned by the withering perianth; seeds brown to black; aril basal or subventral, oval.
Type species: Mammillaria tuberculosa Engelmann.
The two species of this genus known to Schumann were placed by him in the subgenus Escobaria of Mammilaria; they are like the Escobariae in having groved flower-bearing tubercles, but are otherwise different, especially in the flowers, fruit, and seeds.
Eight species are known from northern Mexico and southern Texas.
The genus commerated the work of two distinguished Mexicans, the Escobar brothers, Romulo and Numa, of Mexico City and Juarez.

However, they did not put our two local Escobaria, E. missouriensis and E. vivipara, in the new genus.

Literature Cited:
- Sanchez, Daiel, Balbina Vazquez-Benitez, Monserrat Vaquez-Sanchez, David Aquino, Salvador Aria, 2022.  

 

 

Other articles:
• Golden Checklist Flora:  Escobaria missouriensis;  

Escobaria missouriensis (Sweet) D. R. Hunt “Missouri Foxtail Cactus”

 

Literature Cited:
- Linne´, Carl von, 1753.
- Nuttall, Thomas, 1818.  

The first name that was applied to this cactus was Cactus mamillaris (Nuttall, 1818, v. 1, p. 295). This was an existing name applied by Linnaeus to Mammillaria mammillaris (L.) H. Karst, a native of Columbia and Venezuela (Linnaeus, 1753, v. 1, p. 466). Note that Mammillaria was spelled with two “m”s.

Original Text
Melocactus. roundish.
Species. 1. C. mamillaris. Tubercles ovate terete, bearded; flowers scarcely exserted; berries scarlet about equal with the tubercles. — On the high hills of the Missouri probably to the mountains. A species which was hitherto supposed solely indigenous to the tropical parts of America. It appears to be smaller than the West India plant.

This cactus was not listed in Fraser's catalogue. Why did Nuttall not indicate it was a new name with an asterisk?

It turns out that Cactus mamillaris is a Linnaean (1753) name, but Nuttall does not indicate so. C. mamillaris L. is a synonym of Mammillaria mammillaris (L.) H. Karst. So either Nuttall failed to recognize a new species or applied an illegitimate name to the new species.

Literature Cited:
- Sweet, Robert, 1826.  

Sweet (1826, p. 171) in a catalogue of plants raised in gardens of England placed our species in Mamillaria and gave it the specific epithet of missouriensis.

Original Text
MAMMILLARIA. H.S. Mammillaria. Icosandria Monogynia.
[Number] [Specific Epithet] [Common Name] [Location Found] [Year] [Bloom Month] [Characteristics]
14 missouriensis. Missouri. Missouri.

G. ♄
Cactus mamillaris N. nec aliorum.

“G. ♄” indicates that the plants are grown in a greenhouse and they are shrubby.

The phrase “nec aliorum” literally translates to “no other.” Does that mean there are no other names to be placed in synonomy? Or does it mean that … what?

Literature Cited:
- Britton, Nathaniel Lord, and J. N. Rose, 1919-1923.  

Britton & Rose (1919-1923, v. 4, p. 53) treated our plant as Neobesseya missouriensis (Sweet).

Literature Cited:
- Harrington, H. D., 1954.  

Harrington (1954 & 1964 2ed.) treated our cactus as Mamillaria missouriensis Sweet, Hort. Brit. 171. 1827. Synonyms given were M. similis Engelm., Coryphantha missouriensis (Sweet) Britton & Rose, C. similis (Engelm.) Britton & Rose, Cactus similis Britton & Rose, C. missouriensis (Sweet) Kuntze, and Neobesseya missouriensis (Sweet) Britton & Rose. “Reported as occurring throughout Colorado but our few records from the western and southern parts at 4500-7500 feet.

 

Hunt (1978, p. 13) placed Neobesseya missouriensis (Sweet) Britton & Rose into Escobaria Britton & Rose, saying in part,

...
The justification for combining Escobaria and Neobesseya was (i) that the two genera possess, in common grooved tubercles and pitted seeds; and (ii) the existence of the intermediate species such as Escobaria duncanii, ...
Though my generic concept is broader than that of Britton & Rose and many subsequent students of the Cactaceae, is not as broad, in this instance, as that of Professor Lyman Benson, who includes Escobaria and Neobesseya in oryphantha (Engelm.) Lem. My view accords with the belief, earlier expressed by Buxbaum, that Escobaria (with Neobesseya) represent an evolutionary lineage independent of Coryphantha, that it to saythey are distinct phyletic groups at approximate analogous stage. ...
...

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012, p. 148) treat this cactus as Coryphantha missouriensis (Sweet) Britton & Rose, noting “… very inconspicuous and probably infrequent, plains and outwash mesas …”

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015, p. 256) treated our plant as Coryphantha missouriensis (Sweet) Britton & Rose.

Literature Cited:
- Ackerfield, Jennifer, 2022.  

Ackerfield (2022, p. 276) treats our plant as Escobaria missouriensis (Sweet) D. R. Hunt

 

Other articles:
• Golden Checklist Flora:  Escobaria vivipara;  

Escobaria vivipara (Nutt.) Buxb. “Beehive Cactus”

 

Literature Cited:
- Fraser, John, 1813.

Other articles:
• Golden Checklist Flora:  Fraser's Catalogue, publication details;  

The first publication of Escobaria vivipara was in Fraser's Catalogue (Nuttall, 1813).

Original Text Comments
22 *Cactus viviparus. This species has much the appearance of C. mamillaris, but produces a red flower, like C. Flagelliformis, and a greenish edible fruit, about the size of a grape. Collected near the Mandan towns on the Missourie: lat. near 49°.  

The Cactus mamillaris that Nuttall refers to will eventually become E. missouriensis.

Literature Cited:
- Pursh, Frederick, 1814.  

Pursh (1814, v. 2, p. 735) ...

Cactus viviparus. — C. subrotundus multiplex, tuberculis cylindraceis barbatis supra sulca exaratis proliferis. p. 327.
C. viviparus. Fraser. catal. 1813.  
In Upper Louisiana. ♄ v. v. s. fl. In Hort. Lambert. Resembles C. mammillaris ; the flowers are large and red; the fruit the size of a grape, green and eatable. In the same collection are also two other species mentioned in Fraser's Catalogue ; but only seeing the plants without flowers, I could not define their characters.  

Literature Cited:
- Nuttall, Thomas, 1818.  

Nuttall's Genera of North American Plants (1818, v. 1, pp. 295-296) description of Cactus viviparus.

2. * viviparus. Cespitose; glomeruli subglobose; tubercles cylindric-ovate, bearded, marked above with a proliferous groove; flowers central large and exserted; exterior segments of the calix, ciliate; fruit ficiform, greenish. Hab. With the above [E. missouriensis], on the summits of gravelly hills; flowering from June to August; flowers large and bright-red, almost similar to those of C. flagelliformis. Obs. Nearly allied to the preceding in habit, but differing probably from every other species of this section by the remarkable proliferous tendency of its leaves. Which not unfrequently multiply to the destruction of the parent plant, it consequently never becomes so large as C. mamillaris; inhabiting a climate which is scarcely temperate, from the great elevation of the land above the level of the sea, these 2 species in this country produce long and somewhat fusiform roots, penetrating deep into the earth; towards the approach of winter the upper part of the plant becomes dry, excessively spiny, and almost juiceless, in the spring numerous shoots issue from the root, and those glomeruli which have withstood the intensity of the frost, thus the plants becomes cespitose, forming masses sometimes of 2 or 3 feet in breadth. In spite of its armature the wild antelope of the plains finds means to render it subservient to its wants by cutting it up with his hooves.
The flowers are generally central, more than an inch in length; segments of the calix linear, exterior ones revolute with a fringed margin; petals numerous, narrow, linear and acuminate; berry about the size of grape, smooth and eatable; seed small, cotyledones none, (in the seeds which germinated with me, merely a tubercle similar to those of the parent plant.)

Literature Cited:
- Haworth, Adrian Hardy, 1819.  

Haworth (1819, pp. 72-73) ...

vivipara.
9.
M. (Viviparous) subrotunda multiplex tuberculis cylindraceis barbatis supra sulco exaratis proliferis.
  Cactus viviparus. Pursh. americ. 2. 785.
  Habitat in Louisiana Superiore.
  Introd. 1813 a D. Frazer. G. H. ♃
  Flores magni rubri. Fructus magnitudine fructus vitis viniferae, viridis edulis. Pursh. l. c. — Nuttall's gen. n. amer. p. 295.
  Obs. Spinae plerumque niveae, at 1-3 in singulo fasciculo majores quam ceteris, atro-purpureae.
  Obs. 2. Mr. Nuttall (p. 295.) says this plant, and another, which he (perhaps wrongly) supposes the same as the West Indian Cactus mammillaris of authors, inhabit the summits of gravelly hills near the Missouri, probably to the mountains, and that the flowers of viviparus from June to August are large, bright red, and almost similar to those of C. flagelliformis.

Literature Cited:
- Britton, Nathaniel Lord, and J. N. Rose, 1919-1923.
- Wikipedia contributors, 2020a.  

Britton & Rose (1919-1923, 4 vols.) The Cactaceae is a monograph on plants of the cactus family written by the American botanists Nathaniel Lord Britton and Joseph Nelson Rose and published in multiple volumes between 1919 and 1923. It was landmark study that extensively reorganized cactus taxonomy and is still considered a cornerstone of the field. It was illustrated with drawings and color plates principally by the British botanical artist Mary Emily Eaton as well as with black-and-white photographs.

A black-and-white reprint of the second (1937) edition of The Cactaceae was published by Dover Publications in 1963. In 2006, Daniel Schweich undertook a project to digitize the entire book, and all four volumes can now be downloaded in full color (WikiPedia contributors, 2020a).

Original Text Comments
31. Escobaria vivipara (Nuttall) Britton and Rose in Britton and Brown, Illustr. Fl. Ed. 2. 2: 571. 1913.  

Cactus viviparus Nuttall, Fraser's Cat. No. 22. 1813.
Mammillaria vivipara Haworth, Suppl. Pl. Succ. 72. 1819
Mammillaria radiosa Engelmann, Bost. Journ. Nat, Hist. 6: 196 1850
Echinocactus radiosus Poselger, Allg. Gartenz. 21: 107. 1853.
Echinocactus viviparus Poselger, Allg. Gartenz. 21: 107. 1853
Mammillaria vivipara vera Engelmann, Proc. Amer. Acad. 3: 269. 1856.
Mammillaria vivipara radiosa Engelmann, Proc. Amer. Acad. 3; 269. 1856.
Mammillaria vivipara radiosa Engelmann, Cact. Mex, Bound. 15. 1859, as subspecies.
 
[Description not reproduced.]  
Type locality: “Near the Mandan towns on the Missouri, lat. Near 49°.”
Distribution: Manitoba to Alberta, Kansas, south to northern Texas and Colorado.
 
The group to which Escobaria vivipara belongs has always been very puzzling. Dr. Engelmann, our greatest suthority on this group, was sometimes of one opinion and sometimes of another. Schumann rejected the specific name vivipara of Haworth for this plant since he thought that it was not the same as the vivipara of Engelmann, but in this he must be wrong, for Mammillaria vivipara Haworth was based upon Cactus viviparus Pursh, a name previously used by Nuttall, and both Pursh's and Nuttall's descriptions were based on the specimens collected by Nuttall in “Upper Louisiana” in 1812. This is undoubtedly the plant which Engelmann had in mind and which he called the variety vera. We have not seen the type, but Pursh stated that he had seen flowers in Lambert's Garden.  
Engelmann's remakrs regarding the variability of the species are interesting. In the Proceedings of the American Academy (3: 269) he says:
“The extreme forms are certainly very unlike one another, but the transitions are so gradual that I can not draw strict limits between them.”
 
Escobaria vivipara and the three following species are closely related. They are C. neo-mexicana, C. arizonica, and C. deserti.
This plant is a day bloomer, and according to Engelmann the flowers become fully expanded about one o'clock in the afternoon.  
Hooker in Curtis's Botanical Magazine (pl. 7718) figures and describes a plant purchased from D. M. Andrews of Boulder, Colorado, in which all the spines are brown, the flower is rose-red, and the stigma-lobes are linear and white.  
[A list of illustrations by others is not reproduced.]  
 

Britton & Rose (1919-1923, v. 4, p. 43-44) treat our plant as Coryphantha vivipara (Nutt.) Britton & Rose, in Britton & Brown Illustr. Fl. ed. 2. 2: 571. 1913.

Literature Cited:
- Buxbaum, Franz, 1951.  

Buxbaum (1951) placed E. vivipara in Escobaria. While early volumes of the publication, Österreichische botanische Zeitschrift are available on Biodiversity Heritage Library, the subject volume (v. 98, 1951) is not available.

Literature Cited:
- Harrington, H. D., 1954.  

Harrington (1954 & 1964 2ed.) treats our cactus as Mamillaria vivipara (Nutt.) Haw., Syn. Pl. Succ. Supp. 72. 1819. Synonyms listed are Coryphantha radiosa (Engelm.) Rydb., C. vivipara (Nutt.) Britton & Rose, and Cactus viviparus Nutt.

Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.  

Weber & Wittmann (2012, p. 148) treated our plant as Coryphantha vivipara (Nutt.) Britton & Rose var. vivipara, noting that it is common on the plains and in the San Luis Valley, very rarely in the outer foothills.

Literature Cited:
- Ackerfield, Jennifer, 2015.  

Ackerfield (2015, p. 256) treated our plant as Coryphantha vivipara (Nutt.) Britton & Rose.

Literature Cited:
- Ackerfield, Jennifer, 2022.  

Ackerfield (2022, p. 276) treats our plant as Escobaria vivipara (Nutt.) Buxb.

   

Opuntia (L.) Mill.

 

Literature Cited:
- Majure, Lucas C., Paul Puente, M. Patrick Griffith, Walter S. Judd, Pamela S. Soltis, and Douglas E. Soltis, 2012.  

 

Literature Cited:
- Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P.M., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds., 2018.  

 

Literature Cited:
- Allred, Kelly W., 2020.  

 

Literature Cited:
- Ackerfield, Jennifer, 2022.
- Engelmann, George, 1850.
- Engelmann, Georg, and John M. Bigelow, 1856.
- Ferguson, David J., 1987.
- Pinkava, Donald J., 2003.

Other articles:
• Golden Checklist Flora:   Notes on Opuntia cymochila, Engelmann & Bigelow, 1856;   Notes on Opuntia cymochila, Ferguson, 1987;   Notes on Opuntia macrorhiza, Pinkava, 2003;   Notes on Opuntia macrorhiza, Ackerfield, 2022;   Notes on Opuntia tortispina, Engelmann & Bigelow, 1856;   Notes on Opuntia tortispina, Pinkava, 2003;  

Comparison of Selected Taxa of Opuntia

  Opuntia macrorhiza Engelm. Opuntia tortispina Engelm. & J. M. Bigelow 1 Opuntia cymochila Englem. & J. M. Bigelow
Character Engelmann, 1850 FNANM, Pinkava, 2003 Ferguson 1987 Ackerfield, 2022 Engelmann & J. M. Bigelow, 1856 FNANM, Pinkava, 2003 Engelmann & J. M. Bigelow, 1856 2 Ferguson, 1987
Growth Form     Shrubs, forming clumps, 7.5–16 cm, sometimes from tuberlike rootstocks     Plants 0.8-3 dm. [8-30 cm.]     Shrubs, low, to 0.4 m [40 cm.], creeping from clumps, sometimes from thickened rootstocks.     commonly forms clumps up to five or six feet [1.5 or 1.8 m.] across, rarely to a foot [30 cm.] in height, joints rooting where they touch the soil.
Root   Root a large and fleshy tuber, sometimes 2 or 3 inches [5 to 8 cm.] in diameter;              
Stem   joints 3-4 inches long, about 2½-3½ wide, hardly attenuate at the base. Areolae ¾ — 1 inch distant, more crowded toward the base and on the edges:   Stem segments not easily detached, dark dull green, often cross wrinkled when stressed, flattened, obovate to circular, 5–11 × 3.5–7.5 cm, fleshy (to flabby and cross wrinkled when stressed), tuberculate, glaucous; areoles 5–6(–8) per diagonal row across midstem segment, oval to subcircular, 2–4 mm diam.; wool tan.     segments 5-11 × 3.5-7.5 cm., 5-7 areoles per diagonal row across the midstem segment   Joints rounded, 6-8 inches long ;   Stem segments not easily detached, pale green to deep green, graying with age, wrinkled when stressed, flattened, broadly obovate to ovate, 6.5–15 × 4–10 cm, tuberculate, glossy, glabrous; areoles 6–9 per diagonal row across midstem segment, oval, obovate, or subcircular, 2.5–5 × 1.5–4 mm; wool tan, aging brown.   Joints 2½-3 inches in diameter, orbicular or very slightly obovate pulvillis 6-8 lines apart;   … joints typically a deep, oftern dull green, shrivelling and turning purplish in winter dormancy, when they lie prostrate on the ground … rounded and about 2.5-4 inches (6-10 cm) in length
Spines   spines (often wanting) 1 inch long, the smaller 4-6 lines long.   Spines (0–)1–4 per areole, usually in distal areoles, erect to spreading, white to red-brown, acicular, straight, terete or 1 flattened, ± stout (0.5 mm diam. At base), longest to 60 mm.   Key: few spines per areole in few areoles;   spines (0) 1-4 per areole, to 60 mm long   pulvilli 1-1½ inches apart; bristles short, except on the edges, where they are 2-3 lines long, but rather slender ; spines more numerous than in any other of our species, with juicy fruit, often 6-8, lower smaller ones ?-l inch, larger ones 1½-2½ inches long, entirely white or yellowish horn-colored at base and tip ; on the upper areolae one erect spine, the others spreading in different directions, lower ones deflexed.   Spines 1–9 on most areoles to only on distal 1/2 of stem segment, white to gray with pale brown tips and bases, sometimes brown throughout; central spines 1–3, all deflexed or 1–2 porrect or ascending, terete or flattened, occasionally spirally twisted, 25–70 mm; small spines (2–)3–6(–8) strongly deflexed, usually slender, even bristlelike, 5–15 mm.   the very light yellowish-brown bristles numerous, and conspicuous only on the older joints; only the lowest pulvilli of a joint unarmed, upper ones with 2-5 spines, 2 or 3 larger ones, often reddish-brown at lower half, 1-2 inches long, lower, smaller, paler ones 3-9 lines long.   Key: several spines per areole in most areoles

Desc: several spines per areole, 1-4, often flattened, main spines which mostly spread downward, but one or two may project upward or outward … 1 to several smaller, thin, short downward radiating spines … typically whitish, sometimes with yellowish or more often brownish bases … occasionally may be mostly browh or even blackish.

Glochids     Glochids in dense tuft, pale yellow, tan to red-brown, aging brown, to 5 mm.     glochids in a dense tuft, to 5 mm;     Glochids forming a well developed adaxial tuft, yellow to brownish white, to 6 mm.     glochids are yellow to reddish-brown
Leaves   Leaves subulate, about 5 lines long.               usually green, mostly under 1/4 inch (7 mm) long
Flowers   Flower 3 inches in diameter: ovary 1¼ inch long: petals 1 inch wide, 1½ inch long, pale yellow, red at the base.   Flowers: inner tepals yellow with red basal portions, 25–40 mm; filaments pale yellow; anthers yellow; style white; stigma lobes cream to yellowish.   Key: flowers yellow, but often red centers, stigma lobes pale whitish or yellowish   flowers yellow, 25-45 mm long; filaments yellow;     Flowers: inner tepals yellow to gold, commonly darker to red near base, broadly spatulate, 30–40 mm, apiculate; filaments usually pale yellow; anthers yellow; style whitish to pale green; stigma lobes greenish.     Key: flowers yellow, rarely with red centers, with stigma lobes usually rich green.

Desc: 2.5 to 3 inches (6.5-8 cm) across, with many ruffled yellow petals … toward base orangey or rarely bright orange or red 3 stigma lobes typically rich dark green, but may vary to pale

Fruit   Fruit 1½ inches long;   Fruits green to yellowish to dull red, stipitate, elongate-obovoid, 25–40 × 15–28 mm, fleshy, glabrous; areoles 16–28.     fruit fleshy, green to yellow or reddish, 25-40 mm long.   Fruit similar to that of last species, large, oval, not contracted at base, perhaps less juicy and with a somewhat smaller and deeper umbilicus, 1¾- 2 inches long, 1-1¼ in diameter.   Fruits purple-red, oval to broadly ovate, subspheric or short ovoid, bases not narrowed, 30 × 20–25 mm, fleshy, base not narrowed, glabrous, spineless or nearly so; umbilicus deep; areoles 18–30.   Fruit oval, 1-1¼ inches long, about 10 lines in diameter, purplish, pulpy, sweet, and edible, less contracted at base than O. Rafinesquii; [=O. compressa]   mostly ovoid and narrowed below the apex … areoles usually prominent with obvious glochids and often a few short spines dull purplish red (almost brown) … pulp juicy and purple (among the sweetest and tastiest prickly pear fruits, but are rather small and most of the volume is taken up by the seeds)
Seeds   the strongly margined seeds comparatively few, 2½ lines in diameter.   Seeds tan, subcircular, 4–5 mm, thickish, warped; girdle broad, protruding to 0.5 mm.       Seeds 2-3 lines across, thick and quite regular, with a very slight indentation at the hilum.   Seeds whitish to tan, irregularly shaped, flattened, 4–6 × 3–4 mm; girdle protruding 1–2 mm.   seed remarkably irregular and twisted, 2½ lines in diameter, with a wavy or twisted very sharp rim, whence the name which indicates the undulated border.   mostly about 3/8 inch (5-6 mm) in diameter, regularly discoid, pale tan in color, with a broud prominent rim
Chromosomes     2n = 44.         2n = 44, 66.    
Habitat         Dry, open places, 3600-7000 ft. May-Aug. (plate 39)         dry grasslands, in stable, fairly heavy soils, but westward, where it is drier, they often favor sandy soil.
Distribution       Key: Mostly in tall-grass prairie (esp. sandy areas) from Texas and the Midwest to the Rocky Mountains, and west across N. New Mexico and into SW. Colorado.           Key: Mostly in short grassland on the Great Plains and west across New Mexico.

Desc: Widespread from the Front Range in northern Colorado eastward into western and southern Nebraska4

Character                


Footnotes:

1 Opuntia tortispina is a highly variable taxon that has risen from hybridization of O. macrorhiza with O. polyacantha. Opuntia tortispina has greenish to yellow-green stigma lobes and white to pale green styles, while O. macrorhiza has yellowish stigma lobes and white styles and fruit that are narrowed at the base. It is not recognized as distinct here, but is in the Flora of North America treatment. Ackerfield, 2022, p. 277.

2 Treated as a form of O. rafinesqueii by Engelmann & Bigelow, 1856

3 One plant with deep pink flowers was found in ne. Colorado; probably resulting from hybridization with O. polyacantha.

4 … [thence] southward from there into central Texas and southern New Mexico. It ranges westward through much of New Mexico to near the Arizona border. I have also seen specimens from Santee, northeastern Nebraska, and the species might be expected in short grass areas across Nebraska, perhaps into southern South Dakota. Plants from the Rio Grande drainage are somewhat distinctive and may deserve varietal status.

 

Other articles:
• Golden Checklist Flora:  Notes on Opuntia macrorhiza;   Notes on Opuntia tortispina;  

Opuntia cymochila Engelmann & J. M. Bigelow

 

Literature Cited:
- Engelmann, Georg, and John M. Bigelow, 1856.

Other articles:
• Golden Checklist Flora:  Descr. Cact. [G. Engelmann & J. Bigelow] iv. 42 (1856);   Notes on Opuntia, comparison;  

Engelmann, George, M. D. and John M. Bigelow, M. D. (1856, pp. 42-43) described O. cymochila as a form of O. rafinesquei that was subsequently determined to a a superfluous name and a synonym of O. cespitosa.

a. O. cymochila : diffusa articulis orbiculatis pulvillis subremotis griseo-tomentosis stramineo s. fulvo-setosis, plerisque armatis ; aculeis 1-3 robustioribus elongatis teretibus s. subcompressis tortisque albidis basi saepe rufescentibus, patulis deflexisve, additis saepe 2-3 gracilioribus radiatim deflexis ; flore? Stigmatibus 8; bacca obovata umbilico plano s. parum depresso pulvillis 20-24 griseo-tomentosis parce setulosis, demum nudatis; seminibus irregularibns angulosis margine undulato acuto. (Plate XII, fig. 1-3.)
On the Camanche plains east of the Llaño Estacado, near the 100th degree of longitude, and from there to Tucumcari hill, 80 miles east of the Pecos. Joints 2½4-3 inches in diameter, orbicular or very slightly obovate pulvillis 6-8 lines apart; the very light yellowish-brown bristles numerous, and conspicuous only on the older joints; only the lowest pulvilli of a joint unarmed, upper ones with 2-5 spines, 2 or 3 larger ones, often reddish-brown at lower half, 1-2 inches long, lower, smaller, paler ones 3-9 lines long. Fruit oval, 1-1¼ inches long, about 10 lines in diameter, purplish, pulpy, sweet, and edible, less contracted at base than O. Rafinesquii; seed remarkably irregular and twisted, 2½ lines in diameter, with a wavy or twisted very sharp rim, whence the name which indicates the undulated border.
The orbicular joints, the numerous spines, the oval not clavate fruit, and curiously twisted seed, seem to distinguish this form sufficiently from O. Rafinesquii, but these characters may not be sufficiently constant or important to constitute specific difference. The characters of Opuntiae are not yet sufficently studied to permit us to form satisfactory conclusions about their diagnostic importance ; so we find a form collected on the Sandia mountains, near Albuquerque, which, in habit and appearance, does not differ from the common form of O. Rafinesquii, but which has the seeds of O. cymochila.
O. cymochila, β. Montana : articulis orbiculatis majoribus inermibus s. margine superiore solum aculeatis; pulvillis remotis stramineo-setosis ; aculeis singulis binisve validis albidis infra fuscis ; bacca obovata subclavata seminibus irregularibus acute undulateque marginatis. Joints 3-4½ inches in diameter, pulvilli 9-12 lines apart, spines 12-18, smaller ones 4-6 lines long, on some plants entirely wanting. Fruit 1½ inches long, much contracted at base, with a much depressed, almost funnel-shaped, umbilicus. Seeds cannot be distinguished from those of the plant of the plains.

Literature Cited:
- Ferguson, David J., 1987.

Other articles:
• Golden Checklist Flora:   Notes on Opuntia, comparison;  

 

 

Other articles:
• Golden Checklist Flora:  Opuntia macrorhiza;   Notes on Opuntia cymochila;   Notes on Opuntia tortispina;  

Opuntia macrorhiza Engelm. “Western Pricklypear”

 

Literature Cited:
- Engelmann, George, 1850.

Locations: Guadalupe River.  

Engelmann (1850, v. 6, p. 206) ...

Original Text Interpretation and Comments
O. macrorhiza (n. sp.) : prostrata ; articulis obovato-orbiculatis planiusculis ; pulvillis setis fuscis et saepe aculeis singulis binisve instructis ; aculeis teretibus validis porrectis s. paulo deflexis basi apiceque fuscis ceterum albidis cum adventitio inferiore graciliore reflexo saepe deficiente ; floribus sulphureis basi intus rubellis ; ovario sepalis subulatis deciduis 13 in axillis setulas fuscas brevissimas gerentibus stipato; sepalis interioribus 15-18 subulatis et (internis) ovatis acuminato-cuspidatis ; petalis 8 sepala superantibus late obovato-spathu.latis obtusis cuspidatis eroso-denticulatis ; stigmatibus 5 obtusis, adpressis, stamina numerosa aequantibus ; bacca subpulposa clavata glabrata ; seminibus marginatis. — Naked, sterile, rocky places on the Upper Guadaloupe. Flowers (in St. Louis) in June. Root a large and fleshy tuber, sometimes 2 or 3 inches in diameter; joints 3-4 inches long, about 2½-3½ wide, hardly attenuate at the base. Leaves subulate, about 5 lines long. Areolae ¾ — 1 inch distant, more crowded toward the base and on the edges: spines (often wanting) 1 inch long, the smaller 4-6 lines long. Flower 3 inches in diameter: ovary 1¼ inch long: petals 1 inch wide, 1½ inch long, pale yellow, red at the base. Fruit 1½ inches long; the strongly margined seeds comparatively few, 2½ lines in diameter. — I have found the same plant in similar situations in Western Arkansas ; and it is possible that it may be one of Nuttall's new species (O. mesacantha, O. caespitosa, or O. humifusa) of which I cannot find a description. — Nearly related to O. vulgaris.  

Literature Cited:
- Pinkava, Donald J., 2003.

Other articles:
• Golden Checklist Flora:   Notes on Opuntia, comparison;  

 

Literature Cited:
- Majure, Lucas C., Paul Puente, M. Patrick Griffith, Walter S. Judd, Pamela S. Soltis, and Douglas E. Soltis, 2012.  

Placed in Section Humifusa by Majure et al. (2012) along with O. tortispina and O. cymochila. O. tortispina is not usually accepted by Colorado authors, as it is thought to be a hybrid of O. macrorhiza and O. polyacantha. Most sources treat O. cymochila as a synonym of O. tortispina.

Literature Cited:
- Kilburn, Paul D., and Sally L. White, 1992.
- Weber, William A., and Ronald C. Wittmann, 2012.
- Zeise, Larry Steven, 1976.  

Observations of cacti identified as Opuntia compressa (Zeise, 1976 and Kilburn and White, 1992) have been assumed to be O. macrorhiza per Weber & Wittman (2012).

Literature Cited:
- Ackerfield, Jennifer, 2022.

Other articles:
• Golden Checklist Flora:   Notes on Opuntia, comparison;  

 

 

Other articles:
• Golden Checklist Flora:  Opuntia polyacantha;  

Opuntia polyacantha Haw. “Plains Pricklypear”

 

Literature Cited:
- Nuttall, Thomas, 1818.  

Nuttall (1818, v. 1, p. 296) published Cactus ferox, which he placed in the group Opuntiæ of Cactus L., apparently unaware that Willdenow (1814) published the same name for a cactus occurring in Cuba.

Original Text Interpretation and Comments
4. * ferox. Articulately proliferous; articulations larger, nearly circular and very spiny; spines double, larger spines radiate persistent; flowers numerous; fruit dry and spiny. Hab. In arid situations on the plains of the Missouri, common. Obs. A much larger plant than C. opuntia to which it is nearly allied; exterior spines radiate, with one of them central, solitary and erect; flowers aggregated, marginal, dilute sulphur yellow, rosaceous towards the base; petals subemarginate. Style thick, stigmas 8 to 10 greenish. Colytedones 2, distinct. Flowering in July. Upon this species I found the Coccus coccinelliferus. ferox” means fierce, a reference to the spiny plants.

Coccus coccinelliferus is a synonym for Dactylopius coccus, a scale insect from which cochineal dye is derived. This insect, a primarily sessile parasite, lives on cacti from the genus Opuntia, feeding on moisture and nutrients in the cactus sap. The insect produces carminic acid, which deters predation by other insects. The carminic acid can be extracted from the insect's body and eggs to make the red dye.

Literature Cited:
- Haworth, Adrian Hardy, 1819.  

Haworth (1819, p. 82) ...

Original Text Interpretation and Comments
poly-
cantha.
     8.
O. (Many-spined) articulis compressis obretuso- rotundatis, spinis horridis variabilibus albis, 2-3 senectis subuncialibus divaricato-deflexis.

Cactus ferox, Nuttall gen. n. americ. 296, nec Willd.

Habitat prope flumen Missouri, in America Boreali, in aridis locis.

Cult. in hort. Chels. A.D. 1814. H. ♄

Affinis O. spinosissimae, at longe distincta, et aere aperto viget.

“… to Mr Anderson the excellent Curator of the Physic Garden at Chelsea, which have not heretofore been cultivated in England; and which Mr. Anderson, with his usual zeal in favour of science, has afforded me every opporunity of examining and describing.”

Literature Cited:
- Majure, Lucas C., Paul Puente, M. Patrick Griffith, Walter S. Judd, Pamela S. Soltis, and Douglas E. Soltis, 2012.  

Placed in Section Xerocarpa by Majure et al. (2012) along with O. basilaris, O. erinacea, and O. fragilis. Our other Opuntia, O. macrorhiza, was placed in Section Humifusa along with O. tortispina and O. cymochila.

 

Other articles:
• Golden Checklist Flora:   Notes on Opuntia cymochila;  Notes on Opuntia macrorhiza;  

Opuntia tortispina G. Engelmann & J. M. Bigelow

 

Literature Cited:
- Engelmann, Georg, and John M. Bigelow, 1856.

Other articles:
• Golden Checklist Flora:  Descr. Cact. [G. Engelmann & J. Bigelow] iv. 42 (1856);   Notes on Opuntia, comparison;  

Engelmann, George, M. D. and John M. Bigelow, M. D. (1856, pp. 41)

9. Op. tortispina (sp. nov.): prostrata articulis majusculis adscendentibus obovato-orbiculatis pulvillis subremotis stramineo s. fulvo-setosis ; aculeis 3-5 majoribus compressis angulatis subinde canaliculatis smpe spiraliter tortis, albis basi apiceque saepe corneis, adjectis infra aculeolis 2-3 gracilibus albis ; flore ------ ; bacca ovata areolis sub-20 parvulis notata, late umbilicata, seminibus mujusculis regularibus crassis. (Plate V, fig 2-3.)
On the Camanche plains, near the Canadian river, east of the plateau of the Llaño Estacado. Similar in growth to the more western O. Camanchica. Joints rounded, 6-8 inches long ; pulvilli 1-1½ inches apart; bristles short, except on the edges, where they are 2-3 lines long, but rather slender ; spines more numerous than in any other of our species, with juicy fruit, often 6-8, lower smaller ones ⅓-l inch, larger ones 1½-2½ inches long, entirely white or yellowish horn-colored at base and tip ; on the upper areolae one erect spine, the others spreading in different directions, lower ones deflexed. Fruit similar to that of last species, large, oval, not contracted at base, perhaps less juicy and with a somewhat smaller and deeper umbilicus, 1¾- 2 inches long, 1-1¼ in diameter. Seeds 2-3 lines across, thick and quite regular, with a very slight indentation at the hilum.
I had observed that sometimes 2 plants are produced from the same seed ; this I found to be the case occasionally with Opuntia occidentalis, Engelmanni and dulcis, one of the young plants always much larger and more vigorous than the other. In examining different seeds of this speceis (sic), I succeeded in finding one with two embryos (see figure), one spirally coiled around the other, both together appearing like one large one.

Literature Cited:
- Pinkava, Donald J., 2003.

Other articles:
• Golden Checklist Flora:   Notes on Opuntia, comparison;  

 

 

Literature Cited:
- Britton, Nathaniel Lord, and Addison Brown, 1913.  

Pediocactus Britton & Rose

Original Text
2. PEDIOCACTUS Britton & Rose.
Stems globose, leafless, tubercled, the tubercles arranged in spiral rows bearing clusters of spines arising from areolae. Flowers borne on the tubercles, at or near areolae from which spines are developed. Calyx-tube prolonged beyond the ovary, its tube funnelform, bearing a few scales. Petals numerous, similar to the inner sepals, but larger, pinkish. Stamens numer- ous, borne on the tube of the calyx. Ovary green, globose; style columnar. Berry irregularly bursting, with a terminal scar, nearly or quite scaleless. Seeds tubercled, with a large sub- basal hilum. [Greek, Plains-cactus.]

Three species, natives of central and western North America, the following typical.

 

Other articles:
• Golden Checklist Flora:  Pediocactus simpsonii;  

Pediocactus simpsonii (Engelm.) Britton & Rose “Mountain Ball Cactus”

 

Literature Cited:
- Engelmann, George, 1863.

Locations: Mount Vernon.  

Engelmann (1863, p. 197) published an account of a new cactus in “Additions to the Cactus-Flora of the Territory of the United States” in Transactions of the Academy of St. Louis.

Original Text
2. Echinocactus Simpsoni, spec, nov. : e basi turbinata … [… Latin diagnosis omitted …]
Butte Valley, in the Utah Desert, and Kobe Valley, farther west; var. β in Colorado Territory, e. g. in coarse gravel or in crevices of rocks, abundant near Mount Vernon, at the base of the mountains, Parry, Hall & Harbour ; fl. in May, fr. in July and August. With the New Mexican E. papyracanthus,* the Mexican E. horripilus, Lem., and perhaps the South American E. Odierii, Lem., and E. Cummingii, Salm, this species forms a small section of Echinocacti with the appearance of Mamillariae, named by Prince Salm, (Hort. Dyck., 1849, p. 31,) Theloidei. Through the Escobariae they are nearly allied to Mamillaria, while our species at least, (the fructification of the others not being known,) by its dry fruit, its black tuberculated seeds, and especially the large and curved embryo and the presence of an albumen, proves itself a true Echinocactus, very closely connected with the regularly ribbed E. intertextus, Eng. Cact., Mex. Bound, t. 34. The similarity in all essential organs of these two species is such that no system ought to separate them, proving again of how little essential importance among Cactacese the external form must be regarded; another striking example, among many, is the rat-tail Cereus tuberosus, and its globular or oval allies, C. caespitosus, etc.
[… Additional description omitted …]

Literature Cited:
- Simpson, Captain J. H., 1876.

Locations: Butte Valley. Kobeh Valley.  

The cactus George Engelmann (1863) described was collected by his brother Henry Engelmann on an 1859 expedition across the Great Basin led by Captain Simpson. However, the expedition report was not published until 1876 (Simpson, 1876).

At the time, “Territory of Utah” included all of the present-day State of Utah, most of the present-day state of Nevada, much of present-day western Colorado, and the extreme southwest corner of present-day Wyoming.

The Butte Valley of Simpson's report is the present-day Butte Valley of White Pine and Elko Counties, Nevada. The Ko-Bah Valley is present-day Kobeh Valley, Eureka County, Nevada, along US Highway 50 northwest of Eureka.

The Var. β. minor, was collected near Mount Vernon, by Parry, Hall & Harbour (1862) making it a Jefferson County plant.

Literature Cited:
- Britton, Nathaniel Lord, and Addison Brown, 1913.  

Original Text
I. Pediocactus Simpsoni (Engelm.) Britton & Rose. Simpson's Cactus. Hedge-hog-thistle.
Echinocactus Simpsoni Engelm. Trans. St. Louis Acad. 2 : 197. 1863.
Stems single, globose or with a narrowed base, 3'-6' high, 3'-4' in diameter. Tubercles ovoid, somewhat 4-sided at base, 6"-8" long, arranged in spirals; central spines yellowish below, nearly black above, 5"-7" long, the exterior ones slightly shorter, whitish; flowers greenish pink, 8"-10" long and about as broad, borne to one side at the ends of the tubercles; petals oblong, crenulate and cuspidate at the apex ; berry dry, 3"-3½" in diameter, bearing near its summit 2-3 scales which sometimes have short spines in their axils.
Kansas (according to B. B. Smyth) ; Colorado to Wyoming, Utah, New Mexico and Nevada. April-May.

   

Elaeagnus angustifolia L. “Russian Olive”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Original Text
1. ELAEAGNUS foliis lanceolatis. Roy. lugdb. 250. Hort. upf. 31. anguftifolia
Elaeagnus. Cam. epit. 106. Hort. cliff. 38.  
Olea fylveftris , folio mollii incano. Bauh. pin. 473.  
Habitat in Bohemia, Hifpania, Syria, Cappadocia. ♄  

 

Other articles:
• Golden Checklist Flora:  Lythrum salicaria;  

Lythrum salicaria L. “Purple Loosetrife”

 

 

Other articles:
• Golden Checklist Flora:  Chamerion angustifolium;  

Chamerion angustifolium (L.) Holub. “Fireweed”

(Syn: Epilobium a. L , Epilobium angustifolium L var. intermedium)

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 347) ...

Original Text
1. EPILOBIUM foliis fparfis lineari-lanceolatis. anguftifolia.
Epilobium floribus difformibus , piftillo declinato. Fl. fuec. 304.  
Epilobium foliis lanceolatis integerrimis. Fl. lapp. 146. Hort. cliff. 154. Roy. lugdb. 250.  
Lyfimachia Chamaenerion dicta anguftifolia. Bauh. pin. 245.  
β. Lyfimachia Chamaenerion dicta latifolia. Bauh. pin.245.  
γ. Lyfimachia Chamaenerion dicta alpina. Bauh. pin. 245. prodr. 116.  
Habitat in Europa boreali. ♃  

Literature Cited:
- Holub, Josef, 1972.  

Chamerion angustifolium (L.) Holub, Folia Geobotanica & Phytotaxonomica. Prague. 7: 86 (1972). … unable to penetrate paywall.

 

Other articles:
• Golden Checklist Flora:  Circaea alpina;  

Circaea alpina L. “Small Enchanter's Nightshade”

 

Literature Cited:
- Linne´, Carl von, 1753.  

Linnaeus (1753, v. 1, p. 9) ...

Original Text Translation and Comments
2. CIRCAEA caule adfcendente, racemo unico. alpina.  
Circaea calyce colorato. Fl. lapp. 3.    
Solanifolia Circaea alpina. Bauh. pin. 168.    
Habitat ad redices montium in frigidis Europae. ♃   “She lives in the cold mountains of Europe.”
Planta vix digiti longitudine, magis procumbens, calyce colorado, uti corolla; differt quidem a priori , attamen nimim affinis.   “The plant is scarcely an inch in length, rather stooping down, the calyx colored as the corolla; it differs indeed from the former, but is very closely related.” The former was C. lutetiana canadensis.

 

Other articles:
• Golden Checklist Flora:  Epilobium;  

Epilobium L.

Epilobium (from the Greek) Willow-Herb It is the largest in the family and contains the great majority of species of Onagraceae found in the Old World. Most of the species are somewhat mesophytic in habitat preference. In this genus, the mode of vegetative propagation is variable, some species having above-ground stolons. Those of Iraq have below ground level, either turions (fleshy overwintering buds), soboles (pale elongate shoots) or leafy rosettes which are ± erect. The genus name derives from the Greek words "epi" meaning "upon" and "lobos" meaning "lobe", with reference to position of the petals above the ovary.

  • Perennial herbs, rarely annuals or woody at base.
  • Leaves usually opposite, at least below, and alternate above or in the inflorescence, rarely in whorls of 3.
  • Flowers epigynous or calyx tube absent.
  • Sepals 4, deciduous in fruit.
  • Petals 4, purplish or white, usually deeply notched at the apex.
  • Stamens 8, the epipetalous shorter.
  • Stigma 4-lobed or entire.
  • Ovary 4-locular, with numerous ovules.
  • Capsule slender, elongate, dehiscing loculicidally into 4 valves; seeds with a chalazal tuft of long hairs, wind-borne.
  •  

    Other articles:
    • Golden Checklist Flora:  Epilobium brachycarpum;  

    Epilobium brachycarpum C. Presl “Autumn Willow Herb”

     

    Literature Cited:
    - Presl, Carl B., 1830-1835.  

    Presl (1835, t. 2, p. 30) ...

    EPILOBIUM BRACHYCARPUM. presl.
    E. caule erecto glabro apice paniculato pubescente, foliis alternis petiolatis lineari-lanceolatis acutis remote et argute dentieulatis aveniis, spicis elongatis, bracteis pedicello fructifero longioribus , petalis obcordatis, stigmate indiviso.
    Habitat in Mexico. ♃.
    Long Latin description omitted.

     

    Other articles:
    • Golden Checklist Flora:  Epilobium ciliatum;  

    Epilobium ciliatum Raf. “Fringed Willowherb”

     
     

    Epilobium ciliatum Raf., Medical Repository. New York II. v. (1808) 361.

     

    Other articles:
    • Golden Checklist Flora:  Epilobium hirsutum;  

    Epilobium hirsutum L. “Hairy Willowherb”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p.347-8) ...

    Original Text
    3. EPILOBIUM foliis oppofitis lanceolatis ferratis. Hort. cliff. 145. Fl. fuec. 305. Gron. virg. 154. Roy. lugdb. 251. hirfutum.
    Lyfimachia filiquofa hirfuta magno flore. Bauh. pin. 245.  
    Lyfimachia purpurea. Fuchf. hift. 491.  
    β. Lyfimachia filiquofa hirfuta, parvo flore. Bauh. pin. 245. prodr. 116.  
    Habitat in Europae humidiufculis. ♃  

       

    Oenothera L.

     
     

    • Oenothera sect. Kleinia Munz, N. Amer Fl. ser. 2, 5: 110. 1965 — Type: Oenothera albicaulis Pursh

      • O. albicaulis Pursh “Whitest Evening Primrose” Type.
      • O. coronopifolia Torrey & A. Gray “Crownleaf Evening Primrose”

    • Oenothera sect. Anogra (Spach) W. L. Wagner & Hoch, comb. nov. Anogra Spach, Ann. Sci. Nat. Bot., ser. 2, 4: 164. Sep 1835. Baumannia Spach, Hist. nat. veg. 4: 351. Apr 1835, non Baumannia DC., 1834. Oenothera [unranked] Anogra (Spach) Endlicher, Gen. pl. 1190. 1840. Oenothera subg. Anogra (Spach) Reichenbach, Deut. Bot. Herb.-Buch. 170. 1841. — Lectotype, designated by Pfeiffer, Nomencl. Bot. 1(1): 200. 1872: Anogra douglasiana Spach [=Oenothera pallida Lindley].

      • O. nuttallii Sweet “Nuttall's Evening Primrose”

    • Oenothera sect. Oenothera
      • Oenothera sect. Oenothera subsect. Oenothera
        Onosuris Rafinesque, Fl. Ludov. 95. 1817. — Type: Onosuris acuminata Rafinesque [=Oenothera biennis L.].
        Pseudo-oenothera Ruprecht, Fl. Ingr. 365. 1860. — Type: Pseudo-oenothera virginiana Reprecht [=Oenothera biennis L.].

        • O. biennis L. “King's-Cureall”
        • O. villosa Thunb. “Hairy Evening Primrose”

    • Oenothera sect. Pachylophus (Spach) W. L. Wagner, Syst. Bot. 30: 340. 2005. Pachylophus Spach, Hist. nat. veg. 4: 365. 1835. Oenothera [unranked] Pachylophus (Spach) Endlicher, Gen. pl. 1190. 1840. Oenothera subg. Pachylophus (Spach) Reichenbach, Dwut. Bot. Herb.-Buch. 170. 1841 (as “Pachylophis”). — Type: Pachylophus nuttallii Spach [=Oenothera cespitosa Nuttall].

      • O. cespitosa Nutt. Tufted Evening Primrose.

    • Oenothera sect. Megapterium (Spach) W. L. Wagner & Hoch, comb. nov. Megapterium Spach, Hist. nat. veg. 4: 363. 1835. Oenothera [unranked] Megapterium (Spach0 Endlicher. Gen. pl. 1190. 1840. Oenothera subg. Megapterium (Spach) Reichenbach, Deut. Bot. Herb.-Buch. 170. 1841. — Lectotype, designated by Britton & Broan, Ill. fl. n. U.s., ed. 2, 2: 605. 1913: Megapterium nuttallianum Spach [=Oenothera macrocarpa Nuttall].

      • O. brachycarpa A. Gray. Short-Fruit Evening Primrose.
      • O. howardii (A. Nels.) W. L. Wagner “Howard's Evening Primrose”

    • Oenothera sect. Gaura (L.) W. L. Wagner & Hoch, comb nov. Gaura L., Sp. pl. 347. 1753 — Type: Gaura biennis L. [=Oenothera gaura W. L. Wagner & Hoch].

      • Oenothera sect. Gaura subsect. Schizocarya (Spach) W. L. Wagner & Hoch, comb. et stat. nov. Schizocarya Spach, Ann. Sci. Nat. Bot., ser. 2, 4: 170. 1835. Gaura [unranked] Schizocarya (Spach) Endlicher, Gen. pl. 1195. 1840. Gaura sect. Schizocarya (Spach) P. H. Raven & D. P. Gregory, Mem. Torrey Bot. Club 23: 23. 1973 ["1972"]. — Lectotype, designated by Raven and Gregory, Mem. Torrey Bot. Club 23: 23. 1973. ["1972"]: Schizocarya micrantha Spach [=Oenothera curtiflora W. L. Wagner & Hoch].

        • O. curtiflora W.L. Wagner & Hoch. (Syn: Gaura mollis James, nom. rej., Gaura parviflora Douglas ex Lehm. ) Velvet Butterfly-Weed.

      • Oenothera sect. Gaura subsect. Campogaura (P. H. Raven & D. P. Gregory) W. L. Wagner & Hoch, comb nov. Gaura sect. Campogaura P. H. Raven & D. P. Gregory, Mem.Torrey Bot. Club 23: 27. 1973 ["1972"] — Type: Gaura coccinea Pursh [=Oenothera suffrutescens (Seringe) W. L. Wagner & Hoch].

        • O. suffrutescens (Ser.) W. L. Wagner & Hoch. (Syn: Gaura coccinea Pursh, Gaura glabra Lehm.) Linda Tarde.

     

    Other articles:
    • Golden Checklist Flora:  Oenothera albicaulis;  Notes on Oenothera nuttallii;  

    Oenothera albicaulis Pursh “Whitest Evening Primrose”

     

    Literature Cited:
    - Nuttall, Thomas, 1813.  

    54 * ________ albicaulis. ‡ M.

    Nuttall's (1813) publication of Oenothera albicaulis is nom. inval. because there is no description.

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, Supplementum, p. 733-4) ...

    Original Text
    Oenothera albicaulis. — O. tenuissime pubescens ; caule nervisque foliorum albidis, foliis pinnatifidis, laciniis divaricatis, floribus subspicatis, germinibus sessilibus cylindraceis, staminibus corolla brevioribus, petalis obcordatis.
    O. albicaulis. Fraser. catal. 1813.
    In Upper Louisiana. Bradbury. ♂. v. s. in Herb. Bradbury. Not above four inches high ; flowers large, white. This species is intermediate between O. tetraptera and O. sinuata.

    Generally, the symbol of Mars “♂” denotes a biennial.

    Pursh's listing of “O. albicaulis Fraser catal. 1813” was an error on Pursh's part since he and Nuttall were looking at different plants. Pursh's O. albicaulis is an accepted name for a caespitose plant, whereas Nuttall's name would be replaced by O. nuttallii Sweet for a caulescent plant.

    Literature Cited:
    - Nuttall, Thomas, 1818.
    - Sweet, Robert, 1830, 2nd ed..

    Other articles:
    • Golden Checklist Flora:  Notes on Oenothera nuttallii, Sweet, 1830;  

    Nuttall (1818, v. 1, p. 245) ...

    Original Text
    7. * albicaulis. Perennial; stem simple, erect, white and polished, upper part branching; leaves linear-sublanceolate, rarely subserrulate, under side a little villous; flowers axillary, middle-sized, white; capsule prismatic; petals entire. — Oe. albicaulis. Fras. Catal. 1813. Pursh's synonym wrong, 2. p. 734. See the Herbarium of A. B. Lambert, esq. Hab. From the river Platte to the Northern Andes. Flowering in July and August. Stem about 3 feet high, calix somewhat villous; capsule about an inch long.

    This is considered nom. illeg. because it was previously validly published by Pursh (1814). Note also that the plant being described by Nuttall has a stem to 3 feet tall.

    Sweet (1830) would apply O. nuttallii to Nuttall's O. albicaulis separating it from Pursh's O. albicaulis. See the paragraph for O. nuttallii, below.

     

    Other articles:
    • Golden Checklist Flora:  Oenothera biennis;  

    Oenothera biennis L. “King's-Cureall”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p. 346>

    Original Text Translation and Comments
    biennis. 1. OENOTHERA foliis ovato-lanceolatis planis. Vir. cliff. 33. Hort. upf. 94 Gron. virg. 254. Roy. lugdb. 251. Gort. gelr. 78.  
      Oenothera foliis ovato-lanceolatis denticulatis, floribus lateralibus in fummo caulis. Hort. cliff. 144.  
      Lyfimachia lutea corniculata. Bauh. pin. 245. 516. * Morif. hift. 2. p. 271. f. 3. t. 11. f. 7.  
      Habitat in Virginia unde 1614, nunc vulgaris Europae. “The plant lives in Virginia from whence it was imported in 1614, is now a common European plant.”

    O. biennis is native to the lower 48 states and the southern Canadian provinces, except for the southern Rocky Mountain states.

    Literature Cited:
    - Walter, Thomas, 1788.  

    Oenothera biennis Walter, Fl. Carol. [Walter] 129 (1788).

    Listed in Walter (1788) in his Flora Caroliniana with a description but no distribution or ecological comments.

     

    Other articles:
    • Golden Checklist Flora:  Oenothera brachycarpa;  Gray, 1852-1853, Plantæ Wrightianæ;  

    Oenothera brachycarpa A. Gray “Short-Fruit Evening Primrose”

     
     

    Gray (1852, pt. 1, p. 70) ...

    Original Text
    Œ. (Lavauxia) brachycarpa (sp. nov.): acaulescens, pube brevi cinerea ; caudice incrassato ; foliis primariis ovatis oblongisve subintegerrimis, sequentibus lyrato-pinnatifidis lobo terminali oblongo-lanceolato vel lineari-lanceolato ; tubo calycis prselongo ; capsulis sessilibus ovatis tetragono-alatis lignescentibus laevibus ; seminibus angulatis testa suberosa incrassata. — Between Western Texas and El Paso ; in fruit. Also in the collection of 1851, in flower, and with a few capsules of the preceding year. — The branching, creeping caudex is from one third to half an inch in diameter. Leaves crowded, cinereous or canescent with a fine and close pubescence, glabrate above with age, petioled ; the earlier ones (as seen in the coll. of 1851) mostly entire or repand, with the lamina two inches or less in length and sometimes an inch wide ; the succeeding ones narrower and mostly pinnatifid ; the entire or toothed terminal lobe prolonged, 2-4 inches in length. Tube of the calyx 5 or 6 inches long, rather stout, much as in the section Pachylophis, as likewise, apparently, is the flower; except that the petals (18 lines long) seem to have been pale yellow, changing to rose-color in fading. Capsules closely sessile on the caudex, from half to three quarters of an inch in length, and nearly half an inch in diameter, of a thick, crustaceous texture, acute, but blunt at the apex, scarcely if at all reticulated, the sides broad and not at all ridged, the narrow wings or wing-like angles perfectly smooth and even, thickish, loculicidally 4-valved at the apex, the valves not pointed. Seeds numerous, large, closely packed in two rows in each cell, which they entirely fill, strongly angled by mutual pressure ; the testa very thick and corky, especially toward the chalaza ; Avhere in the young seeds is seen a sort of thick double crest with a denticulate edge, which is nearly obliterated when mature.

     

    Other articles:
    • Golden Checklist Flora:  Oenothera cespitosa;  

    Oenothera cespitosa Nutt. “Tufted Evening Primrose”

     

    Literature Cited:
    - Moulton, Gary E., 1999.

    Locations: Great Falls.  

    First collected July 17, 1806, by Lewis & Clark near the Great Falls of the Missouri, Cascade County, Montana (Moulton, 1999). However, the delay in publishing the results of this expedition until 1814 will leave an opening for other explorers.

     

    Thomas Nuttall ascended the Missouri River from Saint Louis in 1811, traveling at least as far as Fort Mandan, and possibly as far as the junction of the Missouri and Yellowstone Rivers. He collected Oenothera cespitosa, which he took back to England with him, leaving from New Orleans in December, 1811.

    It would seem that the primary material Nuttall took with him to England in 1811 were seeds that were grown in Fraser's garden. Those seeds became the source of the plants for sale in Fraser's Catalogue (Nuttall, 1813).

    Literature Cited:
    - Fraser, John, 1813.
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Fraser's Catalogue, publication details;

    Locations: Fort Mandan Historical Site. White River.  

    Nuttall (1813) first published Oenothera cespitosa in Fraser's (1813) Catalogue. Presumably, these were the only plants grown from seeds collected in 1811 along the Missouri River between the White River and Fort Mandan.

    Original Text
    53 *Oenothera cespitosa. ‡ This species is more perfectly stemless than OE. acaulis of Cavanilles, from which it is distinct. Flowers very large and white, with dilated obcordate petals.

    Note that Nuttall, as the author of Fraser's (1813) Catalogue spelled the specific epithet “cespitosa” and not “caespitosa.” Indeed some authors use caespitosa, such as Weber & Wittmann (2012), whereas others, such as Ackerfield (2015) use cespitosa. It is probably better Latin to use caespitosa. On the other hand, we could just treat the author's epithet cespitosa as valid, and caespitosa as an orthographic variant.

    Literature Cited:
    - Sims, John, M.D., 1814.
    Full Size ImageSims, No. 1593, Oenothera caespitosa.  

    Sims (1814, no. 1593) published an image and description of Oenothera cespitosa, noting that it was illustrated from a plant from Fraser's nursery, and probably originating from Thomas Nuttall.

    [ 1593 ]
    Œnothera cæspitosa. Matted Evening Primrose.
    ********************
    Clafs and Order.
    Octandra Monogynia.
    Generic Character.
    Cal. 4-fidis, tubulofus. Petala 4. Capf. 4-locularis, 4-valvis, infera. Sem. nuda.
    Specific Character.
    Œnothera cæfpitofa ; acaulis, foliis lanceolatis incifo-dentatis, dentatis, capfulis feffilibus, calycis tubo longiffimo, petalis bilobis diftantibus

    At firft fight, this fpecies appears to have a near affinity with Œnothera tetraptera (No. 468). The flowers, as in that, are perfectly white on their firft expanfion, and change in fading to a purplifh red. But the germens in this are feffile, not obovate, and nearly cylindrical, with fmall undulated wings at the four angles ; the tube of the calyx is three times longer than the petals, refembling in this refpect longiflora, No. 365. Root perennial
    In the evening Œnothera cæfpitofa is a beautiful flower, and agreeably fcented. Native of Upper Louifiana in North-America : difcovered we believe by Mr. Nuttall. Our drawing was taken from a plant communicated by Meffrs. Fraser, of the Sloane-Square Nurfery. Flowers in June. Has not, we believe, been defcribed before.

    Literature Cited:
    - Pursh, Frederick, 1814.
    - Sims, John, M.D., 1814.

    Locations: Great Falls.  

    Pursh (1814, v. 1, p. 263) first described our plant as Oenothera scapigera from a collection made by M. Lewis at the falls of the Missouri.

    14. O. scapis radicalibus uniflorus, foliis oblongo-lanceolatis repando-denticulatis decurrentibus, petalis obcordatis, staminibus corolla brevioribus. scapigera.
    On the falls of the Missouri. M. Lewis. ♃ July. v. s. Flowers large, purple, with dark veins : resembles O. acaulis Cavanill  
    All the species, excepting the last one, have yellow flowers.  

    Pursh (1814, v. 2, Supplementum, p. 735) revises his description of our taxon, crediting the illustrations in Sims, Curtis's Botanical Magazine.

    Oenothera caespitosa. — O. acaulis ; foliis lanceolatis inciso-dentatis, capsulis oblongis sessilibus, calycis tubo longissimo, petalis bilobis distantibus. Sims in Bot. mag. 1593. p. 263.
    O. scapigera. Fl. Amer. Sept. 1. p. 263.  
    On the banks of the Missouri. Lewis. ♃. June. July. v. v. in Hort. Fraser ; v. s. specimen imperfectum in Herb. Lewis. This singular species resembles O. acaule of Cavanilles. The flowers open in the evening, are white, changing to red, and of an agreeable scent. The erroneous description given in the place above quoted was owing to a very imperfect specimen in the Lewisian collection.  

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Locations: Fort Mandan Historical Site. White River.  

    Nuttall (1818) included his OE. cespitosa in his Gen. Am. noting the locations of the banks of the Missouri River from the White River north to Fort Mandan. Nuttall does not acknowledge Pursh's (1814) OE. scapigera.

    9. * cæspitosa Cespitose and stemless; leaves lanceolate, sinuately or repandly toothed, smooth, petals very large, dilatedly subbilobed; tube of the calix very long; capsule subconic-oblong, sessile, margin of the valves cristately muricate. Hab. On denudated and arid argillaceous hills on the banks of the Missouri, from White river to the Mandans, and in all probability to the commencement of the mountains. Obs. Segments of the calix carinate, appearing prismatic before flowering; seeds cylindric-ovate; plant 3 or 4 inches high, tube of the calix two and a half inches, flower often 3 inches in diameter. This species is considerably allied to Œ acaulis, which produces pinnatifid leaves and alated capsules, in this the leaves are entire, and the capsules considerable like those of Œ. biennis, cylindric-conic, and tuberculately crested along the margins of the valves; the flowers are white, of uncommon magnitude, and become tinged with red in withering: where its duration is long continued it produces numerous cespitose tufts, but from appearances of this kind its existence is seldom continued through more than 5 or 7 years.
     

    There is a Nuttall voucher in the British Museum (BM001024304) that is determined O. caespitosa Gill ex Hook & Arn. The voucher is annotated as a lectotype of Oenothera caespitosa Nutt. subsp. caespitosa with a synonym of Oenothera montana Nutt. in Torr. & A. Gray Fl. N. Amer. 1:500. 1840. This strongly implies to me that the collection was made on Nuttall's 1834 trip across the continent, rather than his 1811 trip up the Missouri River.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1838-1843.  

    23. Œ. cæspitosa (Nutt.) : almost stemless, caespitose; leaves lanceolate, acute, repandiy toothed or nearly entire, attenuate into a long margined petiole, nearly glabrous; tube of fhe calyx 4 times the length of the carinate acuminate segments ; petals (very large) deeply obcordate, longer than the declined stamens and style; anthers as long as the filaments; capsules, nearly sessile, oblong-conical, somewhat 4-angled ; the margin of the valves tuberculate-cristate. — Nutt. ! in Fraser, cat. ; Bot. mag. t. 1593 ; Pursh, fl. 2. p. 735 ; Nutt. ! gen. 1. p. 246. Œ. scapigera, Pursh, fl. 1. p. 263. Pachylophis Nuttallii, Spach, Onagr. p. 36, t. 30, f. 1.
    Dry and denuded argillaceous hills, on the banks of the Missouri and Platte, Lewis, Nuttall ! June-July. — Root very large- and succulent. Corolla often 3 inches in diameter : the petals very broad, white, with yellowish veins, reddish in withering. Seeds cylindric-ovate. (Nutt.)
    24. Œ. montana (Nutt. ! mss.) : “ stemless, somewhat cæspitose ; leaves broadly lanceolate, sinuate-toothed, pubescent on the margins, tapering into a short petiole ; tube of the calyx about twice the length of the linear-lanceolate acute (scarcely carinate) segments ; petals (large) broadly obcordate ; capsules sessile, cylindrical, conic, striated, even.
    “Plains of the Platte in the Rocky Mountains. — Nearly allied to Œ cæspitosa ; but the petioles and margin of tlie leaves pubescent, the capsules not muricate, &c. Tube of the calyxabout the length of the leaves.” Nuttall.

    Literature Cited:
    - Munz, Philip A., 1931.  

    Munx (1931) is the earliest reference I have seen to treat Oe. montana Nutt. ex Torrey & Gray as a synonym of Oe. cespitosa Nutt.

    Literature Cited:
    - Wagner, Warren L., Robert E. Stockhouse, and William M. Klein, 1985.  

     

     

    Other articles:
    • Golden Checklist Flora:  Oenothera coronopifolia;  

    Oenothera coronopifolia Torr. & A. Gray “Crownleaf Evening Primrose”

     

    Literature Cited:
    - Torrey, John, and Asa Gray, 1838-1843.  

    Torrey & Gray (1840, v. 1, p. 495) ...

    Original Text
    10. Œ. coronopifolia : perennial? minutely pubescent and strigose; stem ascending or erect ; leaves pectinate-pinnatifid, with linear acute segments ; the lowermost somewhat entire ; flowers (small) axillary; tube of the calyx filiform, abruptly dilated at the summit, villous in the throat ; the segments linear-lanceolate, shorter than the tube, longer than the entire roundish petals ; stamens shorter than the petals ; style exserted ; stigmas rather short and thick ; ovaries hirsute. — Œ. pinnatifida, Torr. ! in ann. lyc. New York, 2. p. 201, not of Nutt.
    Forks of the Platte, Dr. James! — Leaves crowded. Corolla about an inch broad when expanded. Fruit unknown.

     

    Other articles:
    • Golden Checklist Flora:  Oenothera curtiflora;  

    Oenothera curtiflora W.L. Wagner & Hoch “Velvet Butterfly-Weed”

     

    Literature Cited:
    - James, Edwin, ed., 1823.  

    James (1823, v. 2, p. 77) ...

    Original Text
    * A large undescribed species of Gaura is common about the banks of all the creeks we had seen since leaving the Arkansa. It attains, ordinarily, the size of G. biennis, but is clearly distinct both from that and all other North American species. It has a broader leaf than any other species of the genus met with in this country. The flowers are small, of a purple colour, and incline to form a terminal spike. The whole plant is covered with a dense silky pubescence, and is remarkably soft to the touch. We propose to call it Gaura mollis.
     

    Gaura parviflora Douglas ex Lehm., Nov. Stirp. Pug. [Lehmann] 2: 15 (1830).

    Literature Cited:
    - Wagner, Warren L., Peter C. Hoch, and Peter H. Raven, 2007.  

    Oenothera curtiflora W.L.Wagner & Hoch, Syst. Bot. Monogr. 83: 211 (2007).

     

    Other articles:
    • Golden Checklist Flora:  Oenothera howardii;  

    Oenothera howardii (A. Nels.) W. L. Wagner “Howard's Evening Primrose”

     

    Literature Cited:
    - Jones, Marcus E., 1893.  

    Jones (1893, v. 3, n. 4, p. 301) ...

    Original Text
    Œnothera Johnsoni Parry Am. Nat. 9, p. 270. This very poorly described plant is said to have elongated stigmas, petals an inch long, calyx tube not shorter than the leaves, capsules 9 to 12 lines long, somewhat 4-angled, strongly nerved, not crested, and to resemble Œ. primiveris, and to be very common at St. George, Southern Utah. The species which I have collected abundantly in Western Utah and Eastern Nevada is perennial, caespitose from a many-branched root, which is covered at the summit with the dead petioles of former leaves, acaulescent or stems an inch or two long; leaves lanceolate, gradually decurrent into the petiole, which is i to 3 inches long, and never more than ½ the length of the blade; blade entire or undulate, or irregularly and sparsely dentate with sharp teeth; whole plant hoary with a dense, soft, and very short pubescence; calyx tips free in the bud; calyx splitting on one side and reflexed in flower, lobes 1 to 1½ inches long, tube 3 to 5 inches long and erect, with 8 striae; petals rhomboidal, entire or slightly lacerate on the edge; 2 to 3 inches wide, and 2 to 2½ long, golden yellow, palmately veined with 3 very prominent and several intermediate veins, each feather veined in addition; the petals, in drying and fading, turn red, and resemble the meshes in the web of a frog's foot; stamens ½ line wide and 6 lines long, versatile, yellow; stigma lobes 4 to 6 lines long, ¼ line wide; capsule ovate, broadly winged, not nerved or veined, less than an inch long, not crested, hoary white; calyx also with scattered, fine, long, white hairs.

    This grows on sunny southern slopes In very dry places, blossoms in June, and is by far the handsomest species of the genus. It is vespertine. Rather common in Western Utah and Eastern Nevada at 6,000 feet altitude. Should it prove to be new, I name it Œnothera Howardi, after Mr. A. M. Howard, the gentleman in my party who saw it first.

    Literature Cited:
    - Nelson, Aven, 1902.  

    Lavauxia howardii A.Nelson, Bot. Gaz. 34(5): 368 (-369) (1902). Nelson (1902, v. 34, p. 368) …

    Original Text
    Lavauxia Howardi (Jones), n. comb. — Oenothera Howardi Jones, Zoe 3: 301. — For some reason this species has been completely ignored by recent writers on the allies of Oenothera, as has also Oenothera Johnsoni Parry, Am. Naturalist 9: 270. Jones suggests the possibility that the species he describes is O. Johnsoni, but at the same time points out characters that unmistakably distinguish the two. While the description of O. Johnsoni is very meager, yet in so essential a matter as the character of the capsule it is very explicit. Certainly no observer, least of all Parry, would have compared a large perennial such as Lavauxia Howardi with the small annual Lavauxia priminervis (Gray) Small, which has a capsule similar to that attributed to O. Johnsoni. The species to which Lavauxia Howardi is closely allied is L. brachycarpa (Gray) Britton Mem. Torr. Club 5: 235; Oenothera brachycarpa Gray, Pl. Wright 1 : 70, and Coulter, Contrib. U. S. Nat. Herb. 2: 116; but here again the character of the capsule serves to distinguish these two. In the latter it is smooth and acute, with narrow wings ; while in the former it is larger, oblong-obtuse, broadly winged, and cinereous pubescent. The leaves also are all oblanceolate, tapering into the petiole, from entire to coarsely and irregularly toothed, therefore the lamina continuous and not distinguished into lateral and terminal lobes.

    The species occurs on arid denudated hills from northern Colorado through Utah to Nevada.

    Literature Cited:
    - Wagner, Warren L., 1983.  

    Wagner, Warren L. 1983 New Species and Combinations in the Genus Oenothera (Onagraceae) Annals of the Missouri Botanical Garden v. 70, p. 194-196.

    Original Text
    Oenothera howardii (A. Nels.) W. L. Wagner, comb. nov.
    Lavauxia howardii A. Nels., Bot. Gaz. (Crawfordsville) 34:368. 1902, non M. E. Jones, Zoe 3:301. 1893. Type: “In clay, Vermillion,” Sevier Co., Utah, 1,600 m., 16 July 1894, M. E. Jones 5631c (RM-13996, lectotype here designated; BM, G, MO 2 sheets, MSC, NY, POM, UC, US, isolectotypes). Jones' publication of the name Oenothera howardii is invalid since he peoposed the name in anticipation of its future acceptance (Art. 34.1b International Code of Botanical Nomenclature, 1978). He placed his discussion of “Oenothera howardii” under the name Oenothera johnsonii Parry (=O. primiveris A. Gray) and stated tha should the entoty prove distince he named it O. howardii. Thus the first valid publication of the epithet was by Aven Nelson as Lavauxia howardii. I am grateful to Dan Nicolson for his helpful comments on this situation.
    Oenothera brachycarpa sensu Munz, Amer. J. Bot. 17: 367. 1930, proparte.
    Plants from Hamilton Co., Kansas, eastern Colorado, Utah, and Nevada formerly included in Oenothera brachycarpa are segregated here as O. howardii. This species if quite distinct from O. brachycarpa in its brilliant yellow petals that dry reddish brown, floral tubes 4.3-11 (12.5) cm long and in that in consists of entirely tetraploid, hexaploid, and octoploid plants whereas O. brachycarpa from west Texas to southeastern Arizona and northern Mexico has pale yellow petals that dry lavender, floral tubes (10.5-)12-22 cm long and is entirely diploid, n =7.

     

    Other articles:
    • Golden Checklist Flora:  Oenothera nuttallii;  Notes on Oenothera albicaulis;  

    Oenothera nuttallii Sweet “Nuttall's Evening Primrose”

     

    Literature Cited:
    - Nuttall, Thomas, 1813.  

    Nuttall (1813, item 54) listed Oenothera albicaulis in Fraser's Catalogue.

    Original Text
    54 *[Oenothera] albicaulis. ‡ M.

    This is considered nom. inval because there is no description or other means to distinguish this plant from any others.

     

    Pursh (1814) published O. albicaulis for a different plant, but included Nuttall's albicaulis as a synonym.

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, v. 1, p. 245) ...

    Original Text
    7. * albicaulis. Perennial; stem simple, erect, white and polished, upper part branching; leaves linear-sublanceolate, rarely subserrulate, under side a little villous; flowers axillary, middle-sized, white; capsule prismatic; petals entire. — Oe. albicaulis. Fras. Catal. 1813. Pursh's synonym wrong, 2. p. 734. See the Herbarium of A. B. Lambert, esq. Hab. From the river Platte to the Northern Andes. Flowering in July and August. Stem about 3 feet high, calix somewhat villous; capsule about an inch long.

    This is considered nom. illeg. because it was previously validly published by Pursh (1814). Note also that the plant being described by Nuttall has a stem to 3 feet tall.

    Sweet (1830) would apply O. nuttallii to Nuttall's O. albicaulis separating it from Pursh's O. albicaulis, which I guess is implied in the string “albicaulis, n. non Pursh.”

    Literature Cited:
    - Sweet, Robert, 1830, 2nd ed..

    Other articles:
    • Golden Checklist Flora:  Notes on Oenothera albicaulis, Nuttall, 1818;  

    Sweet (1830, 2ed, p. 199) published O. nuttallii noting that it is not the same as O. albicaulis Pursh.

      Syst. Name. Color. Eng. Name. Native. Introd. Flower. Habit. Ref. to Figures.
     
    59 Nuttallii. (wh.) Nuttall's N. America. 5, 8. H. ♃  
      albicaulis. N. non Pursh.

     

    Other articles:
    • Golden Checklist Flora:  Oenothera suffrutescens;  

    Oenothera suffrutescens (Ser.) W. L. Wagner & Hoch “Linda Tarde”

    (Syn: Gaura coccinea Pursh , Gaura glabra Lehm. )

     

    Nuttall (1813, p. 2) ...

    39 *Gaura coccinea. ‡ M.


    * New Species.
    ‡ Perennial.
    M. from the Missourie.

    Since there was no description or identifying information this name is not validly published.

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, p. 733) ...

    Gaura coccinea. — G. sericeo-pubescens ; foliis lineari-lanceolatis subdenticulatis, spica conferta, petalis longitudine calycis, stigmate subintegro.
    G. coccinea. Fraser. catal. 1813.
    In Upper Louisiana Bradbury. v. s. in Herb. Bradbury. About a span high ; flowers smaller than G. biennis, scarlet. In the Catalogue quoted this plant is marked as a perennial, but by the appearance of the specimen I should take it to be only annual.

    Literature Cited:
    - Wagner, Warren L., Peter C. Hoch, and Peter H. Raven, 2007.  

    Wagner, et al. (2007, p. 214) ...

    Oenothera suffrutescens (Seringe) W. L. Wagner & Hoch, comb. nov. Gaura ? suffrutescens Seringe in DC, Prodr. 3: 45. 1828. &mdsdh; Type: Fl. Mex. Icon. t. 374 (holotype: G-DC). [This plate is a copy of no. 0717 in the Torner Collection of Sessé and Mociño Biological Illustrations at the Hunt Institute (McVaugh 2000).]
    Gaura coccinea Pursh, Fl. Amer. Sept. 2: 733. 1814. — Type: U.S.A. North Dakota: Morton Co., near Mandan, [22 Jun to 5 Jul 1811], J. Bradbury s.n. (holotype: PH).

     

    Other articles:
    • Golden Checklist Flora:  Oenothera villosa;  

    Oenothera villosa Thunb. “Hairy Evening Primrose”

     

    Literature Cited:
    - Thunberg, Carl Peter, 1794-1800.  

    Thunberg (1794, v. 1, p. 75) ...

    OENOTHERA.
    O. foliis lanceolatis villofis , caule angulato hirfuto. villofa.

    Literature Cited:
    - Frean, Millicent, Kevin Balkwill, Cindy Gold, and Sharon Burt, 1997.  

    Frean (1997, v. 63, n. 6, p. 451) …

    OENOTHERA.
    O. villosa subsp. villosa … was the first taxon of the genus to be collected in southern Africa. It was collected by Thunberg between 1772 and 1775 and now shows scattered local distribution … in Free State, … Kwa-Zulu-Natal and … Western Cape. This scattered distribution suggests that natural dispersal is unlikely and human intervention is probably responsible for its dispersal from the Cape.

     

    Other articles:
    • Golden Checklist Flora:  Aralia nudicaulis;  

    Aralia nudicaulis L. “Wild Sarsaparilla”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Original Text
    nudicaulis. 4. ARALIA caule nudo. Hort. cliff. 113. Gron. virg. 34. Roy. lugdb. 94.
    Aralia caule nuda, redice repente. Cold. noveb. 66.
    Chriftophoriana virginia, zarzae radicibus furculofis & fungofis. Pluk. alm. 98. t. 238. f. 5.
    Habitat in Virginia, simillina (forte eadem) in Iava. ♄

     

    Literature Cited:
    - Sun, Feng-Jie, and Stephen R. Downie, 2010.  

    Apiaceae Apioidae

    Some current phylogenetic analysis (Sun & Downie, 2010) shows that among the western American Apioideae including Aletes, Cymopteris, Lomatium, and Musineon are highly polyphyletic and therefore likely to be significantly reorganized in the coming years.

       

    Aegopodium podagraria L. “Bishops Goutweed”

     
     

    Linnaeus (1753, v. 1, p. 265) ...

    ÆGOPODIUM.  
    1. ÆGOPODIUM foliis caulinis fummis ternatis. Hort. cliff. 107. Fl. fuec. 247. Roy. lugdb. 115. Podagraria.
    Angelica fylveftris minor. f. erratica. Bauh. pin. 155.  
    Herba gerardi. Dod. pempt. 320.  
    Habitat in Europa ad fepes inque pomariis.  

    Linnaeus (1753) says, “It lives in Europe in pastures and orchards.”

     

    Other articles:
    • Golden Checklist Flora:  Aletes acaulis;  

    Aletes acaulis (Torr.) J.M. Coult. & Rose “Stemless Indian Parsley”

     

    Literature Cited:
    - Coulter, John M., and Joseph N. Rose, 1888.  

    Original Text
    48. ALETES. — An acaulescent glabrous perennial, with pinnate leaves, broad sharply toothed or cut rather distant leaflets, mostly no involucre, and involucels of lanceolate bractlets about equalling the yellow flowers.
    1. A. acaulis. Cespitose, with peduncles 4 to 10 inches high, often much longer than the leaves: leaflets ovate, irregularly toothed and cut, sometimes almost pinnatifid: umbel 8 to 15-rayed; rays 5 to 10 lines long; fruit almost sessile, 1½ lines long. (Fig. 147.) — Deweya (?) acaulis Torrey, Pacif. R. Rep. iv. 94. Oreosciadium acaule Gray, Proc. Am. Acad. vii. 343. Seseli Hallii Gray, 1. c. viii. 288. Musenium Greenei Gray, 1. c. 387. Carum (?) Hallii Watson, Bibl. Index. Polypet. 416. Zizia Hallii C. & R. Bot. Gazette, xii. 137, foot-note.
    In the foot-hills of Colorado and New Mexico. Fl. May.
    The history of this species has been somewhat remarkable. Not only has it been referred to six genera, but it has stood under three of them at the same time. Referred first doubtfuliy to Deweya by Torrey in 1856, it was transferred by Gray to Oreosciadium in 1868. It then turned up in the Hall & Harbour collection and was described as Seseli Hallii Gray in 1870. A year or two later it was sent to Dr. Gray by E. L. Greene, and appeared in 1872 as a new species of Musenium, M. Greenei Gray. At this time the same plant was appearing in our publications under three names, Oreosciadium acaule, Seseli Hallii, and Musenium Greenei, all of Gray. In his Bibliographical Index, Watson referred it doubtfully to Carum, as C. (?) Hallii, at the same time recognizing the identity of Seseli Hallii and Musenium Greenei, and quoting them as synonyms. In 1887, however, in Proc. Am. Acad. xxii. 475, Watson records the identity of Oreosciadium acaule with his Carum (?) Hallii, and so the names were at last reduced to one. The fruit characters are those of Zizia, and if they are to dominate over every other consideration this piant must be a Zizia, as we suggested in Bot. Gazette, xii. 137. But no character should be used too arbitrarily, and the complete disimilarity of habit between the recognized species of Zizia and this species seems something that cannot be neglected. If such a thing is to be neglected in this ease, consistency would demand a consolidation of genera such as we are not at present disposed to accept. If this plant, then, is not a Zizia, nor any of the numerous genera proposed for it, a genus must be made for it, for surely, if a plant does not satisfy any genus, it must be sui generis. That it does not satisfy the demands of any recognized genus is to be inferred from its strange history; but it may be proper to point out a few of the reasons why it cannot belong to any of the genera heretofore proposed for it. It is hardly necessary to show why it cannot be a Veloea (Deweya). From Oreosciadium it differs in its prominent calyx-teeth, pinnate leaves, and yellow flowers; from Seseli in its laterally flattened fruit, yellow flowers, and whole habit; from Musenium in its much more prominent ribs, strengthening cells, solitary oil-tubes, almost plane seed-face, and simpler leaves; from Carum in its depressed stylopodium, yellow flowers, and its general habit. Taking its habit and fruit both into consideration it is more nearly related to Musenium than any other genus. For the reasons given, we propose for it a new genus, the name of which indicates its changing history.

     

    Other articles:
    • Golden Checklist Flora:  Berula erecta;  

    Berula erecta (Huds.) Coville “Cut-Leaf Water Parsnip”

     

    Literature Cited:
    - Hudson, William, 1762.  

    3. SIUM foliis pinnatis, umbellis axillaribus pedunculatis, involucro univerfali pinnatifido. erectum.
    Sium five apium paluftre foliis oblongis. Bauh. pin. 154. R. syn. 211.  
    Sium majus anguftifolium. Ger. Em. 256.  
    Sium minus alterum. Park. 1241.  
    Sium erectum foliis ferratis, D. Doody. R. Syn. 211.  
    Anglis, upright Water Parfenep.  
    Habitat in foffis, rivulus et ad ripas fluviorum. ♃ VIII.  

    Literature Cited:
    - Coville Frederick V., 1893.  

    Coville (1893, p. 115) described Berula erecta, noting “The Umbelliferae have been determined by Dr. J. N. Rose.”

    Berula erecta (Huds.) Fl. Angl. 103 (1762), under Sium.
    This is Berula angustifolia (L.) Koch. It was recorded at Resting Springs (No. 271), Winters's ranch, Pahrump Valley, Ash Meadows, Cottonwood Spring, Vegas Ranch, and at several points between Lone Pine and Indian Wells. The plant is one of common occurrence in rivulets of fresh water throughout the desert and within the Larrea belt.

     

    Other articles:
    • Golden Checklist Flora:  Harbouria trachypleura;  

    Harbouria trachypleura (A. Gray) J.M. Coult. & Rose “Whiskbroom Parsley”

     

    Literature Cited:
    - Coulter, John M., and Joseph N. Rose, 1888.  

    Coulter & Rose (1888, p. 125) proposed Harbouria trachypleura.

    Original Text
    47. HARBOURIA. — Glabrous perennials, with 1 to 3 leaves which are ternately decompound and with narrowly linear or filiform segments, involucre and involucels of few subulate bracts, and long-peduncled umbels (mostly in pairs) of yellow flowers.
    Hall & Harbour, to the latter of whom it is dedicated, as the name Hallia is preoccupied.
    H. trachypleura. A foot or more high: leaf-segments mucronulate: umbels (mostly 2 long-peduncled ones) 15 to 25-rayed ; rays an inch long; pedicels 2 to 3 lines long: fruit 2 lines long. (Fig. 146.) — Thaspium trachypleurum Gray, Proc. Acad. Philad. 1863, 63. Cicuta (?) trachyplera Watson, Ribl. Index. Polypet. 417.
    Hall & Harbour 215, Parry 159, etc., etc.) to New Mexico (Fendler 277). Fl. May to July.
    Thaspium and Cicuta. Its position in Thaspium must have been among the so-called "apterous" forms, which we have referred to Zizia, from which genus it is plainly separated by its much more prominent corky ribs, narrow and prominently corky-thickened commissural face, and whole general habit. It is distinct from Cicnta not only in these same characters, but also in its more laterally flattened fruit, equal ribs, yellow flowers. and habitat.

     

    Other articles:
    • Golden Checklist Flora:  Ligusticum porteri;  

    Ligusticum porteri J.M. Coult. & Rose “Porter's Licorice-Root”

     

    Literature Cited:
    - Coulter, John M., and Joseph N. Rose, 1888.  

    4. L. Porteri. Rather stout, 2 to 8 feet high, more leafy, with glabrous or puberulent inflorescence: leaves large, biternate then bipinnate; the numerous rather crowded segments lanceolate to lanceolate-ovate, laciniately pinnatifid, toothed or entire: umbel of numerous rays, mostly with neither involuere nor involucels; rays (fruiting) 1 to 2 inches long; pedicels 3 to 4 lines long; flowers white or pinkish: fruit (immature) oblong-ovate, 2 lines long, with more prominent winged ribs than in other species: oil-tubes 4 to 6 in the intervals, 8 to 10 on the commissural side: seed somewhat dorsally flattened, with very sharply angled back, and face with a broad shallow concavity and central longitudinal ridge.

    In the mountains and foothills of Colorado, Bear Creek (Vasey 223, in 1868), headwaters of Platte River (Coulter, in 1873, distribed (sic) as L. apiifolium, and described as such in Fl. Colorado); Arizona (Palmer 176, in 1877), Huachuca Mts. (Pringle, in 1884, distributed as L. filicinum); New Mexico, mountains near Las Vegas (G. R. Vasey, in 1881, distributed as L. apiifolium), near top of mesa, Raton (Tracy 52, in 1887). Fl. June and July.

    This is also the Colorado form referred doubtfully to L. apiifolium by Brewer & Watson in Bot. Calif, i. 264. It is a curious fact that this species has always been referred to L. apiifolium or L. filicinum, two species with which it cannot be confounded, and has never been referred to L. scopulorum, to which it is most nearly allied. It differs from that species chiefly in its more leafy habit, more numerous crowded smaller and narrower leaf-segments, naked often glabrous umbels, more ovate fruit, which is more prominently winged than in any other species, more numerous oil-tubes, and seed with a very sharply angled back.

     

    Other articles:
    • Golden Checklist Flora:  Lomatium orientale;  

    Lomatium orientale J.M. Coult. & Rose “Salt-and-Pepper”

     

    Literature Cited:
    - Coulter, John M., and J. N. Rose, 1900.
    - Moulton, Gary E., 1999.  

    Coulter & Rose (1900, v. 7, n. 1, p. 220) in their Monograph of North American Umbelliferae published Lomatium orientale.

    Original Text Comments and Interpretation
    20. Lomatium orientale C. & R. sp. nov,  
    Peucedanum nudicaule Nutt. in great part, and of all later authors.  
    Acaulescent or shortly caulescent, with short And soft pubescence, peduncles 1 to 3 cm. high, and a thick elongated root (often swollen in places); leaves bipinnate, the small oblong segments entire or toothed; umbel unequally 5 to 8-rayed, with involucels of scarious-margined (often purplish) lanceolate distinct bractlets; rays 1 to 3.5 cm. long; pedicels glabrous, 5 to 7 mm. long; flowers white or pinkish, with glabrous ovaries; fruit almost round, emarginate at base, glabrous, 5 mm. long, 4 mm. broad, with wings not as broad as body, and indistinct or obsolete dorsal and intermediate ribs; oil tubes solitary in the intervals (rarely 2 in the lateral intervals), 4 on the commissural side; seed face plane.  
    Type locality, plains around Denver, Colo.; collected by Bethel, May, 1895; type in U. S. Nat. Herb. Ellsworth Bethel (1863-1925) came to Colorado in 1888 and until 1914 was a teacher in th Denver High Schools. Receiving a Masters degree, in 1917 he started a new career as a plant pathologist. There are 16 collections attributed to Bethel from Golden s.l.
    On the plains from North Dakota to Kansas, and west to Arizona and Washington.  
    Specimens examined:  

    Kansas: Western Kansas, Herb. State Agric. Coll.

    Nebraska: Wilcox, in 1887; Long Pine, Brown County, Rutter, June 1, 1893.

    South Dakota: Aurora County, Wilcox, May 20, 1892.

    Montana: Blankinship, May 3-4, 1890; Warm Peak, Bear Lodge, V. Bailey, June 8, 1894.

    Wyoming: Fort Russell, Ruby, in 1885; Cheyenne, Havard, in 1893; Laramie plains, Nelson, May, 1893 and 1894.

    Colorado: Hall & Harbour 212, in 1862; Palmer Lake, Alice Eastwood, May 25, 1890; Cache la Poudre, Cowen, May 23, 1891; foothills, altitude 1,800 to 1,950 meters, Crandall, May, 1893 and 1894; near Windsor, Osterhout, May, 1894; plains about Denver, Bethel, May, 1895; Log Canyon and Rist Canyon, Holzinger 3, May 31, 1896.

    New Mexico: Mangus Springs, Rusby 148, February 25, 1S80.

    Arizona: MacDougal 5, June, 1891 ; Flagstaff, altitude 1,650 meters, MacDougal 5, May 31, 1898; Clifton, Davidson, in 1899.

    Utah: Palmer 181, in 1877.

    Idaho: Allen, in 1873; boundary of Idaho and Washington, Canby, in 1891.

     
    In 1818 Nuttall transferred Pursh's Smyrnium nudicaule to Ferula, and cited with the type of Pursh (a Lewis & Clark plant from the Columbia River) a more eastern plant, “on the high plains of the Missouri, commencing about the confluence of the river Jauk [Jaune= Yellowstone or Jacque= Dakota],” a plant said by Nuttall to be associated on the eastern plains with L. foeniculaceum. Since then the name nudicaulis has been associated with the plant of the eastern plains. It was a puzzle to us to find that the type locality of Smyrnium nudicaule was entirely to the west of its present range, but Messrs. Robinson and Greenman have cleared up the matter by examining Pursh's type, and discovering that it is the same as the abundant Seseli leiocarpum of Hooker, from the same region. This leaves the eastern plant without a name. The Lewis & Clark collection that Coulter & Rose refer to is currently determined Lomatium nudicaule (Pursh) Coulter & Rose and is showin as f. 96 in Moulton (1999).

     

    Other articles:
    • Golden Checklist Flora:  Musineon divaricatum;  

    Musineon divaricatum (Pursh) Raf. “Leafy Wildparsley”

     

    Literature Cited:
    - Pursh, Frederick, 1816.  

    Pursh (v. 2, supplement, p. 732) adds S. divaricatum to Seseli on page 197, which heretofore contained only S. triternatum (=Lomatium triternatum (Pursh) J.M.Coult. & Rose) from the Lewis & Clark herbarium.

    Original Text
    p. 197 Seseli divaricatum. — S. caule ramosissimo divaricato, foliis bipinnatifidis, laciniis lanceolatis incisis, involucris nullis, involucellis linearibus.

    In Upper Louisiana. Bradbury, v. s. in Herb. Bradbury. Resembles S. tortuosum very much.

    Literature Cited:
    - Rafinesque, C. S., 1819.  

    Rafinesque (1819, p. 101) published Marathrum divaricatum based upon Seseli divaricatum Pursh.

    Original Text Translation
    22. Marathrum. (Ombellif.) Fleurs hermaphrodites a involucelles, sans involucres. Calice ovale 5-dente. 5 petales obcordes. 5 etamines longues. 2 longs styles caducs. Semences ovales, a dos convexe ou gibbeaux, legerement anguleux. — Caulescent. Feuilles pinnatifides , involucelles polyphylles , fleurs jaunes. — Le type de ce genre est le Sesili divaricatum de Pursh et Nuttall ; mais il fiddere evidemment du genre Seseli par son dente, ses petales obcordes, ses semences anguleuses et ses fleurs jaunes. 22. Marathrum. (Umbellif.) Hermaphroditic to involucral flowers, without involucres. Calyx oval 5-toothed. 5 obcorded petals. 5 long stamens. 2 long obsolete styles. Seeds oval, with convex or gibbeau back, slightly angular. — Caulescent. Pinnatifid leaves, polyphylous involucelles, yellow flowers. & mdash; The type of this genus is the Sesili divaricatum of Pursh and Nuttall; but it evidently belongs to the Seseli kind by his teeth, his obcorded petals, its angular seeds and yellow flowers.

    This turned out to be an unavailable name because Matathrum was unavailable having been previously used by Robert Brown.

    Literature Cited:
    - Rafinesque, C. S., 1820.  

    Rafinesque (1820, p. 71) published a correction for his previously published Marathrum.

    Original Text Translation
    1. Dans le prodrome de 50 nouveaux genres de plantes d'Amerique, j'ai decrit deaux nouveaux genres sous les noms de marathrum et de pythagorea ; je me suis apercu depuis lors, que ces noms avaient deja ete employes, le premier par Robert Brown, le second par Louriero; et comme je suis convaincu de l'importance et de la necessite d'eviter des doubles emplois en Botanique, je m'empresse de rectifer cette erreur, et de proposer les noms suivans en place. 1. In the prodrome of 50 new genera of plants from America, I have described new genera under the names of marathrum and pythagorea; I have noticed since then that these names had already been used, the first by Robert Brown, the second by Louriero; and as I am convinced of the importance and the need to avoid duplication in Botany, I hasten to correct this error, and propose the following names in place.
    Mon G. marathrum devra se nommer musineon. Ces deux noms sont des synonymes de fenouil. My genus marathrum should be called musineon. These two names are synonyms for fennel.
     

    Torrey & Gray (1838, v. 1, pt. 4, p. 642) published Musenium and species from a manuscript by Nuttall.

    Original Text
    47. MUSENIUM. Nutt. mss.
    “Margin of the calyx 5-toothed ; the teeth persistent. Petals obovate ; the point inflexed. Styles slender, reflexed, rather long. Fruit ovate or ovate-oblong, laterally compressed. Carpels more or less minutely scabrous, with 5 filiform acute slightly prominent ribs. Intervals with 2-3 vittae. Commissure with 4 vittae. Carpophore 2-cleft. Seed with the sides moderately incurved. Perennial dwarf rather foetid resiniferous (North American) herbs, with fusiform roots, and a short caudex, or branching dichotomously from the base. Leaves 2-3-pinnatifid. Involucre none. Involucels unilateral, of a few rather rigid narrow leaflets. Flowers yellow or white.” Nutt.
    § 1. Stem dichotomous : flowers yellow.
    1. M. divaricatum (Nutt. ! mss.) : decumbent ; stem short, dichotomously branching from the base ; leaves bipinnatifid ; divisions confluent with the winged rachis ; segments short, rather acutely toothed ; fruit somewhat glabrous. — Seseli divaricatum, Pursh, fl. 2. p. 732? ; Nutt. gen. 1. p. 194 ; DC. prodr. 4. p. 146.
    β Hookeri : rachis narrow ; fruit scabrous, with elevated points. — M. Hookeri, Nutt.! mss. Seseli divaricatum, Hook.! fl. Bor.-Am. 1. p. 264 ; Sims, bot. mag t. 1742. (ex Hook.)
    Naked and arid hills and plains of the Upper Missouri, Nuttall ! β. Plains of the Upper Platte, near the Rocky Mountains, Nuttall ! On the Saskatchawan, Drummond! Douglas. May. — Plant about a span long. Leaves all, except the radical ones, opposite, glabrous and shining : petiole and rachis distinctly winged : lamina with an ovate outline ; primary and secondary divisions 3-4 pairs ; the segments about one-third of an inch long, 3-4-toothed. Peduncles 4-5 inches long, scabrous, naked, rigid, stout. Umbels 10-20-rayed ; the rays (in fruit) about half an inch long. Fruit 2 lines long, oblong-ovate : pericarp thin : vittae filled with a strong terebinthine oil. — The plant exudes small drops of resin spontaneously.

     

    Other articles:
    • Golden Checklist Flora:  Cornus sericea;  

    Cornus sericea L. “Creek Dogwood”

     

    Literature Cited:
    - Linne´, Carl von, 1771.
    - Plukenet, Leonard, 1696.  

    Original Text Comments and Interpretation
    fericea CORNUS arborea, cymis nudis, foliis fubtus fericeis.  
      Cornus femina laurifolia, fructu nigro-caeruleo, officulo compreffo. Pluk, alm. r 21, t. 169. f. 3. Leonard Plukenet (1641-1706) Almagestum botanicum. 1696, 1705. One of several pre-Linnean checklists.
      Habitat in America feptentrionali. ♄. H. U.  
      Frutex altior : Ramis rubicundis . laevibus. Ramuli ultimi obfolete ancipites , a Petioli dorfo , vix manifefto , decurrente.  
      Folia oppofita (nec bifaria) , ovata . plicato - lineata ; fubius alba (non pallide viridia) , fericeo-nitentia.  
      Cymae terminales, folio breviores.  
      Petala ovata f. oblongiufcula , alba , patentia. Stamina alba , corolla duplo longiora. Receptaculum purpurafens.  
      Simillima C. fanguineae et albae , at difficile diftinguatur , fed Baccae atro-purpureae. “Similar to C. sanguineae and C. alba, but difficult to distinguish, except for black-purple berries.”

     

    Other articles:
    • Golden Checklist Flora:  Arctostaphylos uva-ursi;  

    Arctostaphylos uva-ursi (L.) Sprengel “Bearberry”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Original Text
    4. ARBUTUS caulibus procumbentibus, foliis integerrimis. Fl. lapp. 162. t. 6. f. 3. Fl. fuec. 339. Mat. med. 211. Hort. cliff. 163. Roy. lugdb. 440. Hall. helv. 415. Uva urfi
    Uva urfi. Cluf. hift. 1. p. 63.  
    Radix idaea putata & Uva urfi. Bauh. hift. 1. p. 524.  
    Arbutus noveboracentibus Bearberry. Cold. nov. 104.  
    Habitat in Europa frigida, Canada. ♄  

    Literature Cited:
    - Sprengel, Curt Polycarp Joachim, 1826.  

    Sprengel 1825, v. 2., p. 287 ...

    Original Text
    1542. Arctostaphylos Adanf.
    Uva urfi* 1. A. caule procumbenta fruticofo, foliis obovatis integerrimis coriaceis nitidis, floribus fafciculatis. Europ. et Amer. bor. (Arbutus L.)

     

    Other articles:
    • Golden Checklist Flora:  Orthilia secunda;  

    Orthilia secunda (L.) House “Sidebells Wintergreen”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Original Text
    fecunda. 3. PYROLA racemo unilaterali. Fl. fuec. 332.
      Pyrola floribus uno verfu fparfis. Fl. lapp. 168.
      Pyrola folio mucronato ferrato. Bauh. pin. 181.
      Pyrola 2 tenerior. Cluf. pan. 506.
      Habitat in Enropae (sic) borealis fylvis. ♄

    Literature Cited:
    - House, Homer D., 1921.  

    House (1921) moved our plant to Orthilia Raf. from Pyrola without comment.

    Original Text
    Orthilia Raf. Aut. Bot. 103. 1840
    Ramischia Opiz, Seznam 82. 1852.
    Actinocyclus Klotzsch,, Akad. Monats Berlin (1857): 14. 1857.
    Orthilia secunda (L.) House, comb. nov.
    Pyrola secunda L. Sp. Pl. 396. 1753.
    R. secundiflora Opiz, l. c.
    Actinocyclus secundus Klotzsch, l. c.
    R. secunda Garcke, Fl. Deuts. Ed. 4, 222. 1858.
    O. parvifolia Raf., l. c.
    In our northern swamps, merging into the var. obtusata (Turcz.) House, comb. nov. (pyrola secunda var. obtusata Turcz.), originally described from northern Asia and Europe, but which appears to be practically the same as described by Paine (Cat. Pl. Oneida County 135. 1865), as var. pumila. All intermediate forms between the typical species and the var. obtusata, occur in a swam near Newcomb, New York.
    An Additional species of this genus occurs in Mexico.
    Orthilia Elatior (Lang) House, comb. nov. (Actinocyclus secundus elatior Lange; Ramischia elatior Rydberg).

     

    Other articles:
    • Golden Checklist Flora:  Pyrola asarifolia;  

    Pyrola asarifolia Michx. “Liverleaf Wintergreen”

     
     

    There is a collection of P. asarifolia from “draw W of Little Scraggy Peak,” Philip V. Woodhead, #20-W, 29 June 1916, RM402926.

     

    Other articles:
    • Golden Checklist Flora:  Androsace occidentalis Pursh “Western Rockjasmine”;  

    Androsace occidentalis Pursh “Western Rockjasmine”

     

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 1, p. 137) ...

    Original Text
    1. A[ndrosace] tenuissime pubescens ; coliis involucrisive olongo-spathulatis integerrimis, perianthiis angulatis, capsulis calyce brevioribus. occidentalis.
    On the banks of the Missouri. Nuttall. ☉ v. s. in Herb. Nuttall. Approaches near to A. elongata, according to specimens of Pallas's in the Herbarium of A. B. Lambert, Esq.  

     

    Other articles:
    • Golden Checklist Flora:  Androsace septentrionalis;  

    Androsace septentrionalis L. “Pygmyflower Rockjasmine”

    (Syn: Androsace septentrionalis L. ssp. subumbellata (A. Nelson) G. T. Robbins)

     

    Other articles:
    • Golden Checklist Flora:  Dodecatheon pulchellum;  

    Dodecatheon pulchellum (Raf.) Merr. “Beautiful Shootingstar”

    (Syn: Exinia pulchella Raf. , Primula pauciflora (Greene) Mast & Reveal )

     

    Other articles:
    • Golden Checklist Flora:  Lysimachia ciliata;  

    Lysimachia ciliata L. “Fringed Loosestrife”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Habitat in Virginia, Canada (Linnaeus, 1753, v. 1, p. 147).

     

    Other articles:
    • Golden Checklist Flora:  Fraxinus americana;  

    Fraxinus americana L. “White Ash”

     

     

    Other articles:
    • Golden Checklist Flora:  Ligustrum vulgare;  

    Ligustrum vulgare L. “Privet”

     

     

    Other articles:
    • Golden Checklist Flora:  Eustoma grandiflorum;  

    Eustoma grandiflorum (Raf.) Shinners “Showy Prairie Gentian”

     

    Literature Cited:
    - Shinners, Lloyd H., 1957.  

     

     

    Other articles:
    • Golden Checklist Flora:  Frasera speciosa;  

    Frasera speciosa Griseb. “Elkweed”

     

    Literature Cited:
    - Hooker, Sir William Jackson, 1829-1840.
    Full Size ImageHooker's illustration of Frasera speciosa.  

    Hooker, William Jackson, Sir, 1785-1865 Flora boreali-americana, or, the botany of the northern parts of British America : compiled principally from the plants collected by Dr. Richardson & Mr. Drummond on the late northern expeditions, under command of Captain Sir John Franklin, R.N. To which are added (by permission of the Horticultural society of London,) those of Mr. Douglas, from north-west America, and of other naturalists v. 2, p. 66.

    The Gentianaceae was authored by Dr. Grisebach, see footnote on page 54.

    Original Text
    2. F. speciosa (Dougl. ms.); caule erecto tereti sulcato glabro, foliis quaternis oblongo-linearibus acutiusculis, racemo elongato densifloro, calyce corollam aequante, foveis elliptico-oblongis binis. (Tab. CLIII.)
    Longer Latin description omitted.
    Hab. On the low hills near Spokan and Salmon Rivers and subalpine parts of the Blue Mountains, near the Kooskooska River, N.W.C. Douglas. — [This is quite distinct from the preceding (F. carolinensis), much larger in every part, with longer petals and constantly 2 oblong foveae upon each of them. — Hook.]
    Tab. CLIII. — Fig. 1, Flower; fig. 2, Petal: – magnified.

     

    Other articles:
    • Golden Checklist Flora:  Apocynum cannabinum;  

    Apocynum cannabinum L. “Indian Hemp”

     

     

    Other articles:
    • Golden Checklist Flora:  Apocynum ×floribundum;  

    Apocynum ×floribundum Greene “Dogbane”

     
     

    Greene (1893, p. 151) published A. floribundum without mention of it being a hybrid.

    Original Text
    Apocynum floribundum. Glabrous, pallid and glaucous, 2 feet high, with numerous ascending and somewhat fastigiate branches rising to about the same level, each ending in a cyme: leaves about 2 inches long, from ovate to elliptical, mucronate, the margins sparsely serrulate-scabrous: cymes not dense, many-flowered, erect: corollas erect, lurid-purplish, nearly cylindrical, the lobes erect, or only a little spreading. Dry ground bordering pine woods, in the higher mountains west of the Mohave Desert, in Kern Co., Calif., 1889. Nearest A. cannabinum, but of very different habit; the fastigiate branches, each with its own many-flowered cyme, uniting to form a compound corymb often a foot broad. A. cannabinum, common in many parts of California, was found in wet ground by streamlets, in the same region, and true to its character, the dense terminal cyme of small greenish flowers, being greatly surpassed by those of the single pair of lateral branches.

    Literature Cited:
    - Johnson, Samuel A., Leo P. Bruederle, and Diana F. Tomback, 1998.  

    Johnson, et al. (1998) confirmed that A. ×floribundum is a hybrid and that the occurrences they examined were clonal.

     

    Other articles:
    • Golden Checklist Flora:  Asclepias speciosa;  

    Asclepias speciosa Torr. “Showy Milkweed”

     

    Literature Cited:
    - Torrey, John G., 1828.  

    Torrey (1828, p. 218-219) described James' collection as A. speciosa ...

    Original Text
    260. A. speciosa, tota planta tenuiter tomentosa ; foliis ovato-cordatis obtusis, oppositis, brevi-petiolatis ; umbellis axillaribus, terminalibusque solitariis ; coronae foliolis erectis, lanceolatis, antheridiis triplo longioribus, basi intus obtuse bidentatis.

    Desc. Stem erect, branched ? LeavesUmbels axillary and terminal, pedicels white-lanuginous, thick. Flowers twice as large as in A. syriaca. Calyx woolly ; segments lanceolate, spreading. Corolla pale purple ? Segments reflexed, oblong, obtuse. Leaflets of the nectary lanceolate, erect, straight, acute. Horn short, incurved, compressed. Fruit not seen.

    Hab. On the Canadian ?

    Obs. Flowers larger than in any other North American species of this genus.

     

    Other articles:
    • Golden Checklist Flora:  Asclepias incarnata;  

    Asclepias incarnata L. “Swamp Milkweed”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Original Text
    8. ASCLEPIAS foliis lanceolatis, caule fuperne divifo, umbellis terminalibus congeftis.
    Afclepias caule erecto ramofo annuo, foliis lanceolatis, umbellis terminalibus erectis pluribus. Vir. clif. 20. hort. cliff. 78. Roy. lugdb. 411. Gron. virg. 27.
    Apocynum minus rectum canadenfe. Corn. canad. 9. t. 93. Barr. rar. 8. t. 72. Raj. hift. 1089. Habitat in Canada, Virginia. ♃
    Caulis ubi dividitur folia e regione tria producit.
    incarnata.

     

    Other articles:
    • Golden Checklist Flora:  Asclepias pumila;  

    Asclepias pumila (A. Gray) Vail “Plains Milkweed”

     

    Literature Cited:
    - Gray, Asa, 1876a.  

    Gray (1876a, p. 70) in an extended footnote revising Asclepias published Asclepias pumila as a variety of A. verticillata.

    Original Text
    A. verticillata, L. A widely distributed species, including A. galioides, HBK., of Mexico. Var. pumila is a singularly dwarf or depauperate form, of the western dry plains, from Nebraska to New Mexico. Var. subverticillata (A. verticillata var. galioides, Torr. Bot. Mex. Bound. 164, chiefly), is a marked form, with single stems, simple or branched, the leaves mostly in pairs and threes, and their margins little revolute, the horns of the hoods rather less exserted. Decaisne's A. verticillata var. linifolia may include this ; but it is evidently a mixture of A. verticillata (to which the specimens from "Florida and Georgia" may belong) ; of A. virgata, Lag. (A. angustifolia, Roem. & Schult. &c.), which, from Kunth's character of opposite leaves and little exserted horn, may be A. linifolia, HBK.; and of A. Mexicana, Cav. (from which may come the character of leaves 4-6-nate), which must be identical with A. fascicularis of Decaisne. Here also A. linearis, Scheele in Linntea, xxi.

    Literature Cited:
    - Britton, Nathaniel Lord, and Addison Brown, 1898.  

    Britton & Brown (1898, v. 3, p. 12) published Asclepias pumila as an isonym. Text of the section on Asclepiadaceae was revised by Miss Anna Murray Vail.

    Original Text
    22. Asclepias pumila (A. Gray) Vail. Low Milkweed. (Fig. 2921.)

    Asclepias verticillata var. pumila A. Gray, Proc. Am. Acad. 12: 71. 1876.

    Stems 4'-10' high, tufted from a woody root. Leaves very numerous, crowded, sometimes obscurely whorled, filiform-linear, 1'-2' long, smooth or minutely roughened, the margins revolute; umbels 2-several, short-peduncled, few-flowered; pedicels filiform, puberulent, 3"-4" long; corolla greenish white, its segments oblong, 1½"-2" long; column short; hoods white, erect, oblong, entire, equalling the anthers, shorter than the slender incurred horn; follicles erect on erect fruiting pedicels, narrowly spindle-shaped, l½'-2' long, finely puberulent.

    Dry plains. South Dakota to Arkansas, Colorado and New Mexico.

    Anna Murray Vail (January 7, 1863 – December 18, 1955) was an American botanist and first librarian of the New York Botanical Garden. She was a student of the Columbia University botanist and geologist Nathaniel Lord Britton, with whom she helped to found the New York Botanical Garden. In her Wikipedia entry are three links to her studies in the Asclepiadaceae, which may show her original publication of Asclepias pumila

     

    Other articles:
    • Golden Checklist Flora:  Asclepias stenophylla;  

    Asclepias stenophylla A. Gray “Narrow-leaved Milkweed”

     

    Literature Cited:
    - Nuttall, Thomas, 1834b.  

    Nuttall (1834b, p. 201) ... Proposed Polyotus angustifolius noting that Acerates Elliott was already employed for another genus … some sources, IPNI in particular, show the publication date as 1836.

    4. P. *angustifolius. Caule erecto, foliis linearibus sub-oppositis, umbellis caulinis subsessilibus; lepanthium antheridio sublongius, sessile, foliolis apice tridentatis.
    Descript. Perennial. Stem simple, herbaceous and slender, the lower part naked, about eighteen inches high ; leaves very long, narrow and acute, somewhat revolute and scabrous on the margin, one to one and a half lines wide, opposite and alternate, nerves confluent below the margin ; umbells globular, small, three or four to eight ; pedicells about the length of the flowers ; flowers greenish, with a mixture of white ; segments of the corolla oblong, reflected ; lepanthium sessile, divisions linear, longer than the antheridium, tridentate above, the central denture minute, concave and auriculate below; clefts of the antheridium salient from the base to the summit; cusps broad and membranaceous, concealing the stigma; follicles two. — Hab. In dry prairies from Fort Smith to Red river. Flowering in June. — Obs. This species appears to be very nearly related to Gomphocarpus, and also very nearly to Asclepias, particularly A. cinerea, from which it merely differs in the absence of the short internal awn ; it may, however, be considered as present in the central, acute and shorter third denture of the segments of the lepanthium.

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1844.  

    DeCandolle in his Prodromus vol. 8 (1844, p. 522-523) ... proposed Acerates angustifolia ... the chapter on Order Asclepiadeae was written by J. Decaisne.

    9. A. angustifolia, caule erecto, foliis suboppositis linearibus acutis interdum marginibus anguste revolutis scabrisque, umbellis multifloris globosis subsessilibus, coroll. laciniis oblongis reflexis, cucullis gynostegio sublongioribus apice tridentatis. ♃ in Am. sept. graminosis prope Fort Smith ad Red-River. Species habitu Asclep. cinerae ex cl. Nutt. Polyotus angustifolius Nutt. Trans. phil. am. vol. 5, p. 201.

    From Wikipedia:

    Joseph Decaisne (7 March 1807 – 8 January 1882) was a French botanist and agronomist. He became an aide-naturaliste to Adrien-Henri de Jussieu (1797-1853), who served as the chair of rural botany. It was during this time that he began to study plants brought back by various travelers like those of Victor Jacquemont (1801-1832) from Asia. Decaisne used applied research, most notably on the agronomy of the madder, the yam and the ramie. He was also interested in algae.

    Literature Cited:
    - Gray, Asa, 1876a.  

    Gray (1876a, p. 72) …

    A. stenophylla. Polyotus angustifolius, Nutt. in Trans. Amer. Phil. Soc. v. 201. Acerates angustifolia, Decaisne in DC. l. c. Nuttall noted the clear transition to Asclepias.

     

    Other articles:
    • Golden Checklist Flora:  Asclepias viridiflora;  

    Asclepias viridiflora Raf. “Green Comet Milkweed”

     

    Literature Cited:
    - Rafinesque, C. S., 1808.  

    Original Text
    18. Asclepias viridiflora green flowered asclepias ; leaves lanceolated, obtuse, hirsute, umbells axillar, bending down, corniculas, without appendices. I have found it in several parts of Maryland and Pennsylvania, mostly in fields.
     

    Pursh (1814, v. 1, p. 181) also published A. viridiflora, though his name is considered illegitimate because it was previously published by Rafinesque (1808). I think it more likely that Pursh chose Rafinesque's name but failed cite Rafinesque by oversight or intent. We'll never really know. Pursh does tell us that he had seen the plant alive (“v. v.”)

    Original Text
    9. A. caule simplici erecto hirsuto, foliis lanceolato-oblongis obtusiusculis subsessilibus utrinque tomentoso-hirsutis, umbellus lateralibus solitariis subsessilibus nutantibus subgloboso-densifloris, appendicibus nullis

    In dry fields : Pensylvania to Virginia. ♃ June, July. v. v. Flowers green.

    viridiflora.

     

    Other articles:
    • Golden Checklist Flora:  Evolvulus nuttallianus;
    • Field Notes:  Coll. No. 2319, 6 Jun 2020;
    Full Size ImageFlower of Coll. No. 2319, Evolvulus nuttallianus
    Full Size ImageColl. No. 2319, Evolvulus nuttallianus  

    Evolvulus nuttallianus Roem. & Schult. “Shaggy Dwarf Morning Glory”

    Evolvulus nuttallianus is native to Colorado and is found from Montana and North Dakota south to Texas and Arizona. It is usually described as occuring on sandy and rocky prairies and plains, chaparral, pinyon-juniper and oak woodlands. It has also been found on limestone glades and bald knobs in the Ozark region of Missouri. It is a non-vining morning glory-like perennial that grows as a densely hairy subshrub to 20” tall with prostrate to decumbent stems that root at the nodes as they go. It has very small, bell-shaped, lavender to blue morning glories (to 1/2” across) from spring to mid-summer on stems with oblanceolate, hairy, silvery-green leaves (to 1” long). Flowers usually close up at night and on cloudy days. There is some confusion in the taxonomy of the genus Evolvulus. This species is sometimes listed as synonymous with E. pilosus.

    The genus name comes from the Latin word “evolvo” meaning to untwist or unravel because this genus does not twine like many species in the Convolvulus family.

    The specific epithet honors British botanist Thomas Nuttall (1786-1839) who traveled extensively in the U.S., as discussed below.

    For gardeners, E. nuttallianus is winter hardy to USDA Zone 4. It is easily grown in organically rich, consistently moist, well-drained soils in full sun to part shade. It also performs well in sandy soils with good drainage. But they may be difficult to find in commerce.

    Literature Cited:
    - Nuttall, Thomas, 1813.  

    Published in Fraser's Catalogue as:

    32 *Evolvulus sessiliflorus. ‡ M.
    However, this is nomen nudum because it has no description.

    Literature Cited:
    - Graustein, Jeannette E., 1967.
    - Pursh, Frederick, 1814.

    Other articles:
    • Glossary:  v. s.;  

    Thomas Nuttall traveled up the Missouri in 1811, partially duplicating the trip of Lewis and Clark, spending the summer in and around Fort Mandan (Graustein, 1967). I have been unable to determine where Nuttall's collection of E. nuttallianus is located. Graustein notes that many of Nuttall's collections are in the British Museum, and few of their holdings are available online. Regardless, Pursh (1814) published the first description in his Flora of North America. Pursh arrived in London in November 1811. Nuttall and Pursh met in London in the spring of 1812. Nuttall must have had his collections with him.

    Pursh (1814) described Nuttall's morning-glory as E. argenteus.

    Latin My Interpretation
    230. EVOLVULUS. Gen Pl. 524.   230. EVOLVULUS. Gen Pl. 524.  
    1. E. erectus ; foliis oblongis utrinque sericeo-tomentosis, pedunculis unifloribus brevibus. argenteus. 1. Evolvulus that is erect, leaves oblong silky-felted both sides, peduncles short with a single flower. argenteus.
    On the banks of the Missouri. Nuttall. v. s. Flowers yellow.   On the banks of the Missouri. Nuttall. In have seen it in the dried condition. Flowers yellow.  

    Full Size Image
    Pursh desription of Evolvulus argenteus
    The name E. argenteus was unavailable because is was previously used by Robert Brown, 1810. Pursh notes that he saw the collection “v. s.” but it was not in the Lambert herbarium. Pursh characterizes the flowers as yellow, but since he worked from dried material, the blue to purple color may have faded.

    Literature Cited:
    - Nuttall, Thomas, 1818.
    Full Size ImageNuttall (1818) description of Evolvulus nuttallianus.  

    Nuttall (1818) published his own description of E. nuttallianus in his Genera of North American Plants. He writes both Pursh's name argenteus and his own name of pilosus.

    251. EVOLVULUS. L.

    Calix 5-parted. Corolla rotate-campanulate; lobes subemarginate. Styles 2, deeply bifid; segments capillary and divergent. Stigma simple. Capsule 2-celled, 4-valved, 2 to 4 seeded.

    Stem creeping, procumbent, or erect. Leaves alternate entire; flowers pedunculate, small; peduncles solitary, bibracteate, capsule perfecting 1, 2, or 4 seeds.

    Species. …

    3. argenteus, Ph. (pilosus.) Perennial; stems simple, erect, and low, many from the same root: the whole plant densely hairy and shining: leaves cuneate-oblong, acute, crowded; peduncles l-flowered, subsessile; bibracteate, bractes sessile in the axill; segments of the calix linear. — Obs. Allied to E. Commersoni. Stems 4 to 6 inches high; leaves 6 to 8 lines long, 2 to 4 wide, extremely hairy as well as the stem; flowers solitary, appearing sessile, purple, edge of the plaits hairy, capsule 4-valved, often perfecting only a single seed. — Hab. On arid gravelly hills near the confluence of Rapid river and the Missouri; flowering in May. This genus, with the above exceptions, exists exclusively within the tropical regions of India, Australia, and America.

    By current standards of publication, one would think that Nuttall was putting his pilosus in synonomy with Pursh's argenteus.

    Literature Cited:
    - Ro¨mer, Johann Jacob, Joseph August Schultes, Julius Hermann Schultes, Jurt Polycarp Joachim Sprengel, and J. G. Cotta, 1820.
    Full Size ImageRoemer & Schultes description of Evolvulus nuttallianus  

    In the meantime, Roemer and Schultes (1820) published the first valid name for Nuttall's collection.

    LatinMy Interpretation
    18. E. Nuttallianus; erectus, foliis onlongis utrinque sericeo-tomentosis, pedunculis unifloris brevibus. E. argenteus Pursh Fl. Sept. Americ. I. p. 187.
    Flores flavi. Ad ripas Missouri. Nuttall.
    Obs. Nomen mutandum orat, cum sit antiquior argenteus R. Brown.
    18. Evolvulus nuttallianus; erect, leaves oblong silky-felted, single flowers on short pedicels. (E. argenteus Pursh Fl. Sept. Americ. I. p. 187.)
    Flowers yellow. On the banks of the Missouri River. Nuttall.
    The name was changed, because argenteus was used earlier by R. Brown.

    This was just a name change because Pursh's proposed E. argenteus was previously used by R. Brown (1810).

    Literature Cited:
    - Nuttall, Thomas, 1834b.
    Full Size ImageNuttall (1834) description of E. pilosus  

    Then, in 1834, Nuttall first publishes his name of E. pilosus in his descriptions of plants of Arkansas territory.

    2. E. pilosus. Erectus, foliis lineari-oblongis utrinque sericeo-pilosis, pedunculis uniflorus brevibus. Nuttall's Gen. Am. Pl. 1, p. 174 (E. Nuttallianus, erectus, foliis oblongis utrinque sericeo-tomentosis, pedunculis unifloris brevibus. Schultes, Syst. Veg. vol. 6, p. 198; E. argenteus, Pursh, 1, 187). — Obs. Flowers purplish, coming out in the middle of the stem; peduncle shorter than the calix; calix segments partly linear and acuminate. — Hab. On the high hills of Red river near Kiamesha,

    This name is, of course, superfluous and therefore illegitimate because Roemer and Schultes previously validly published nuttallianus.

     

    Other articles:
    • Golden Checklist Flora:  Aliciella pinnatifida;  

    Aliciella pinnatifida (Nutt. ex A.Gray) J.M.Porter. “Sticky Gilia”

     

    Literature Cited:
    - Don, George, 1831-1838.  

    G. Don (1831-1838, v. 4., p. 245) …

    Original Text Comments
    7 G Se'ssei; stem dwarf, branched, clothed with glandular down ; leaves pinnatifid ; segments cuneated, with pointed teeth ; brecteas ovate-lanceolate, mucronatem longer than the calyx ; corolla with a filiform tube, double the length of the calyx ; flowers faccicled. (.) [circle with a dot inside. H. Native of Mexico. Gilia pinnatifida, Sesse et Mocino, in herb. Lamb. Stamens inclosed.
    Sesse's Gilia. Pl. ¼ foot.
     

    Literature Cited:
    - Gray, Asa, 1863.

    Other articles:
    • Glossary:  Gilia;  

    Gray (1863) Enumeration of the Plants Collected by Parry, Hall, and Harbour notes Gilia pinnatifida ined. I assume it was ined. in Gray's hands as he published it in a revision of Polemoniaceae (Gray, 1870).

    Original Text Comments
    456. Gilia pinnatifida, Nutt. ined.  

    Literature Cited:
    - Gray, Asa, 1870.  

    Original Text Comments
    3. GILIA, Ruiz & Pav.  
    § 10. GILLIANDRA. …  
     
    48. G. Pinnatifida, Nutt. In herb.; Gray Enum. Pl. Parry. … [Latin diagnosis not reproduced] … — N. New Mexico and Colorado to Snake River, &c., in or near the Rocky Mountains, Nuttall, Fendler, and various collectors. A part of Geyer's 42 and 25, referred to G. inconspicus, much exserted stamens three lines long. Seeds with a close coat, wholly unchanged when wetted.  

    Literature Cited:
    - Porter, J. Mark, 1998.  

    Porter (1998) recircumscribed Aliciella and placed G. pinnatifida in that group.

    Original Text Comments
    1. Aliciella pinnatifida (Nutt. ex Gray) J. M. Porter, comb. nov.  
    Gilia pinnatifida Nutt. ex Gray, Proc. Amer. Acad. Arts 8: 276. 1870, (basionym). Navarretia pinnatifida (A. Gray) Kuntze, Revisio, Gen. Pl. 2: 433. 1891 Gilia vixcosa Woot. & Standl, Contr. U. S. Natl. Herb. 16; 161, 1913. TYPE: — U.S.A. New Mexico: (without location, but presumably near Santa Fe) 1847, A. Fendler 655 [lectotype: (here designated) GH!].  
     
    Although the Nuttall collection [as a candidate for lectotype] may seem a logical choice, it presents problems because it lacks flowers, a diagnostic feature of this species. In addition, the collection locality is vague (Lewis River), referring to three different rivers in the mid 1800s. By contrast, the Fendler collection is clearly consistent with Gray's description, posessing flowers, fruit, and a basal rosette. … also the collection locality of the Fendler collection is less ambiguous.

    Literature Cited:
    - Porter J. Mark, and Leigh A. Johnson, 2000.  

    Original Text Comments
    Formerly included in Gilia ... [See publications by Grant] ... Aliciella is more closely related to Loeselia and Ipomopsis than to Gilia ... [See publications by Johnson and Porter] ..., based on DNA sequence data. Morphologically, too, Aliciella differs from Gilia in chromosome number (most, but not all, species of Aliciella are N = 8) and seed morphology (seeds do not produce copious mucilage; large seeded species tend to have an irregular wing).  

     

    Other articles:
    • Golden Checklist Flora:  Collomia linearis;  

    Collomia linearis Nutt. “Tiny Trumpet”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;

    Locations: Cheyenne River.  

    Nuttall (1818) described a new genus Collomia and placed C. linearis therein.

    Original Text
    194. *COLLOMIA. †
    Calix Cyathiform, rather large, border 5-cleft, acute. Corolla funnelform, 5-lobed, lobes oval-oblong, very short, tube straight, long, and slender. Capsule 3-cornered, 3-celled, 3-valved, 3-seeded, valves obcordate. Seed oblong, angular, enveloped by a tenacious mucilaginous integument, (visible when moistened.)
    Annual, leaves alternate, simple, and entire; flowers small and inconspicuous, conglomerated in a terminal fascicle, resembling a capitulum, subtended by several bractes which are broader than the leaves.
    A genus appertaining to the Natural Order Polemonideae and intermediate with Phlox and Polemonium
    Species. 1. C. linearis. Minutely and pulverulently pubescent; leaves oblong-linear, or sublanceolate; involucrate leaves, ovate-lanceolate, acute; bractes and calix viscid.
    Phlox linearis? Cavan. __ 6. p. 17. t. 527.
    ... [Long description not reproduced] ...
    Hab. Near the banks of the Missouri, about the confluence of Shian river, and in the vicinity of the Arikaree village, in moist places. Flowering in June; flower violaceous. It appears to be the same plant figured by Cavanilles, and first discovered in Chili. In upper Louisiana, or above the confluence of the Platte and the Missouri, we no longer meet with any species of Phlox. To this genus probably also belongs Phlox biflora of Chili, which is also annual, but the habit appears to be different.

    I think the “Shian river” is the Cheyenne River.

    Full Size Image
    Coll. No. 713, Collomia linearis

     

    Other articles:
    • Golden Checklist Flora:  Gilia ophthalmoides;  

    Gilia ophthalmoides Brand “Pinyon Gilia”

     

    Literature Cited:
    - Brand, A., 1907.

    Locations: Gold Mountain.  

    A. Brand. IV. 250 Polemoniaceae. Heft 27, in Engler, Adolf, 1844-1930 Das Pflanzenreich: regni vegetablilis conspectus Leipzig Verlag von Wilhelm Engelmann. 1907. pg. 108. https://www.biodiversitylibrary.org/bibliography/250

    30. G. ophthalmoides Brand n. sp. — Caulis e basi ramosissimus, parce glandulosus, 25 cm altus. Folia radicalia subrosulala, pinnatifida, segmentis brevibus ovatis mucronulatis; caulina minima, trifida. Flores numerosi, sparsi, pedicellati, pedicellis terminalibus calyce longioribus, axillaribus et lateralibus multo minoribus; calyx campanulatus, glaber, dentibus subulatis tubo subbrevioribus; corolla (ex sicco) lilacina, ad faucem 5 oculis obscuris ornata, 1 2 mm longa, calyce triplo longior, tubiformis, fauce tubo duplo breviore; stamina sinubus affixa lobis corollae breviora; Stylus longitudine corollae; ovarium ovoideum, loculis 3 — 4-ovulatis. Capsula ovoidea, longitudine calycis ; semina parva, orbicularia, alata, sub aqua dense mucilaginosa.
    Nevada: Washes-Gold Mt. 2000 m (Purpus n. 5990, Herb. San Francisco).

     

    Other articles:
    • Golden Checklist Flora:  Gilia sinuata;  

    Gilia sinuata Benth. “Rosy Gilia”

    (Syn: Gilia inconspicua (Sm.) Sweet var. sinuata (Douglas ex Benth.) A. Gray)

    Literature Cited:
    - Bentham, George, 1845.  

    Douglas ex Bentham in DeCandolle (1845, v. 9, p. 313) ...

    19? G. sinuata ( Dougl.! mss. ), erecta glaberrima superne viscida , foliis linearibus pinnatifidis laciniis brevibus paucis, floribus pedicellatis dissitis numerosis, corolla calyce pubescente duplo v. triplo longioie tubo breviter exserto. (I) in arenosis aridis pr. Oakanagon ad flum. Columbia Americae boreali-occid. (Douglas!). Herba pedalis multicaulis ramosa, hinc G. inconspicuae , illinc G. tenuiflorae affinis et cum priore confusa. Cor. infundibuliformes 5 lin. longae , laciniis ovatis. ( v. s. )

     

    Other articles:
    • Golden Checklist Flora:  Ipomopsis aggregata aggregata;  

    Ipomopsis aggregata (Pursh) V. E. Grant ssp. aggregata

     

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 1, p. 147) ...

    Original Text
    2. C[antua] pubescens ; foliis tenuissime pinnatifidis, floribus terminalibus lateralibusque aggregato-subcapitatis subsessilibus, laciniis corollae angustatis oblongis sensim acutis, stylo incluso brevissimo. aggregata.
    On the banks of the Mississippi. M. Lewis. ♂. June. v. s. in Herb. Lewis. Flowers scarlet, more slender than the former species : the segment of the corolla not spreading, but erect, and small in proportion to the tube.  

    Literature Cited:
    - Grant, Verne, 1956.  

    Grant (1956) published our new combination without comment.

    Literature Cited:
    - Moulton, Gary E., 1999.  

    Moulton (1999) notes that Lewis' collection of I. aggregata occurred in Hungery Creek, Lolo Trail, Idaho County, Idaho, June 26, 1806.

     

    Other articles:
    • Golden Checklist Flora:  Ipomopsis aggregata candida;  

    Ipomopsis aggregata (Pursh) V. E. Grant ssp. candida (Rydb.) V. E. Grant & A. D. Grant “Scarlet Gilia”

     

    Literature Cited:
    - Rydberg, Per Axel, 1901a.

    Locations: La Veta.  

    Rydberg (1901, v. 28, p. 29-30) ...

    Gilia candida sp. nov.
    A glandular perennial with short caudex. Stems often more than one, ascending or erect, simple, 4-6 dm. high, leafy : leaves rather fleshy, divided into linear-filiform, spinulose-tipped segments ; the upper gradually smaller : inflorescence a very narrow panicle : branches usually very short and few-flowered : flowers subsessile : calyx glandular-pubescent, about 8 mm. long ; its lobes lanceolate, bristle-tipped : corolla white, or rarely pale pink, 2-3 cm. long, salverform : its lobes oval or obovate, obtuse : stamens inserted unequally below the throat, included.
    This species is nearest related to G. aggregata ; but differs in the white corolla and its rounded obtuse lobes. It is also lower and the upper leaves more reduced. It grows on dry hillsides at an altitude of 2000-2700 m.
    Colorado : Mesas near La Veta, 1900, F. K. Vreeland, 602 (type); Veta Pass, 1900, Rydberg & Vreeland, 5730 ; Calham, 1893, De A. Saunders.

     

    Other articles:
    • Golden Checklist Flora:  Ipomopsis spicata;  

    Ipomopsis spicata (Nutt.) V.E. Grant. “Spiked Ipomopsis”

     

    Literature Cited:
    - Nuttall, Thomas, 1848b.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1848, publication details;

    Locations: Scotts Bluff.  

    Nuttall (1848) described some of his own collections along the Platte River while describing Gambel's collections.

    Original Text Comments
    GILIA.  
    §. Perennials or biennials, with the leaves often sparingly pinnatifid towards the extremity, or entire and linear, fleshy. Flowers in condensed clusters, capitate or in spikes, generally white. Corolla tubular, with a deeply 5-cleft, spreading border. Stamens shortly exserted or even with the summit of the tube. Stigmas very short. Ovaries 2 to 4 in a cell, rarely 1. — *Elaphocera.  
    ...  
    G. *spicata. Perennial ; leaves linear, fleshy ; flowers in clusters, spiked ; stem and calyx lanuginous, segments of the calyx linear acute and ciscid ; tube of the corolla exserted ; stamens at the summit of the tube.

    Hab. On the hills near Scott's Bluffs of the Platte. Flowers white, segments oblong. (Nuttall.)

     
    G. *trifida. Biennial ; radical leaves linear ; cauline trifid towards the extremity, fleshy and smooth ; flowers clustered in spikes ; stem and calyx pubescent, segments of the calyx linear and very acute ; tube of the corolla exserted ; stamens at the summit of the tube.

    Hab. With the above, which it greatly resembles, except in the leaves ; cells of the capsule each with three or four ovules. About a span high. (Nuttall.)

     
    ...

    Literature Cited:
    - Grant, Verne, 1956.  

    Grant (1956) separated Ipomopsis from Gilia and placed I. spicata into section Microgilia (Benth.) V. Grant.

     

    Other articles:
    • Golden Checklist Flora:  Microsteris gracilis;  

    Microsteris gracilis (Douglas ex Hook.) Greene “Slender Phlox”

     

    Literature Cited:
    - Greene, Edward L., 1898b.  

    Greene (1898, v. 3, n. 18, p. 300) ...

    MICROSTERIS.
    Small much branched annuals, with entire leaves, all except the floral ones opposite. Flowers minute, scattered, singly or in pairs in the axils of the raraeal alternate leaves. Calyx, corolla, stamens and capsule wholly as in Phlox. Seeds few, large, the coat when moistened developing a this glutinous mass, this not spirilliferous.
    1. M. gracilis. Collomia gracilis, Dougl. in Hook. Bot. Mag. t. 2924 (1829), and Gilia gracilis, Hook. l. c. Phlox gracilis, Greene, Pitt. i. 141 (1887). That this, as published by Hooker, was an aggregate, is apparent from the fact that, in the Flora Boreali-Americana, he prints as synonyms of this, two species which Douglas had segregated and named in manuscript.1 In other words, Douglas had made out three species, which Hooker, in publishing, merged in one ; merely printing the other two names as synonyms. The figure in the Botanical Magazine appears to represent altogether only one of Douglas's three, and this must accepted as the type of the species. This type came from the Spokane River region, and is beautifully represented in several herbaria by specimens which Mr.Leiberg distributed in 1891, from Lake Pend d'Oreille, under n. 119. Heller's n. 3278 is also the same. Mr. Leiberg has also collected in the same region a second species, the distinctness of which he has insisted upon. I have no doubt that this is one of Douglas' segregates, and I take up the name which he assigned it.

    1 See Hook. Fl. ii. 76.

       

    Phlox

     

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Original Text

    5a. Stems solitary or a few together, never forming mats; leaves usually over 2.5 cm long; flowers distinctly pedicelled. P. longifolia Nuttall. Dry Grasslands and sagebrush, lower Arkansas Valley drainage; North Park.

    5b. Stems numerous from a stout taproot, forming mats; leaves less than 2 cm. long; flowers without obvious pedicels. ... (6)

    6b. Leaves not at all glandular. ... (7)

    7b. Leaves not especially short or imbricate, nor prominently ciliate; not in alkaline soils. ... (8)

    8b. Inflorescence glabrous or ± pubescent, but not [with long crinkly hairs]. P. multiflora A. Nelson. Common in forested areas from the foothills to the montane.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Original Text

    1a. Flowers 3-12 in a cyme, on conspicuous, spelder pedicels; stems with well-developed internodes, the longer ones well over 10 mm long; leaves mostly over 20 .. long; plants erect to tufted but not mat- or cushion-forming, usually over 10 cm tall ... 2

    2a. Hyaline membranes between the calyx lobes carinate, forming a distinct raised ridge, bulging towards the base; plants glabrous or often glandular-hairy, especially in the in florescence; flowers pink to white; not restricted to southwestern counties ... P. longifolia

    1b. Flowers 1-3(5), sessile or short pedicellate (on pedicels 3-10 mm long) at the ends of stems; stems with shorter internodes often hidden by the leaves, the longer ones to 10 mm long; leaves mostly 2-30 mm long; plants caespitose, often mat- or cushion-forming, usually to 10 cm above the ground ...3

    3b. Leaves glabrous, granular-scabrid, glandular, or arachnoid-pubescent with tangled hairs, but not stiffly ciliate or sometimes ciliate on the margins but also arachnoid-pubescent at the base or granular-scabrid throughout; calyx glabrous or villous- to arachnoid-tomentose; plants variously distributed, usually below 9500 ft in elevation (but up to 10,500 ft) ... 6

    6b. Calyx 5-15 mm long; leaves linear to lance-subulate, 2-30 mm long, less crowded and spreading; plants forming loose to dense mats, sparsely arachnoid-pubescent in the leaf axils or on the calyx to glabrous or granular-scabrid, the plants green or sometimes grayish; corolla lobes 4-11 mm long ... 7

    7b. Hyaline membranes between the calyx lobes flat, not carinate and forming a ridge; plants of the eastern or western slope ... 8

    8b. Leaves 10-30 mm long and 1-2 mm wide; internodes usualllt visible; leaf axils arachnoid-woolly or not; calyx 6-15 mm long, arachnoid-pubescent or not; style 5-9 mm long; flowers sessile to shortly pedicellate on pedicels 3-10 mm long ... 9

    9b. Plants above 6000 ft, variously distributed but absent from the northeastern plains; calyx 10-15 mm long, glabrous or sparsely hairy but the hairs not tangled; flowers white, pink, or bluish ... P. multiflora

     

    Literature Cited:
    - SEINet, 2019+.

    Other articles:
    • Golden Checklist Flora:  Phlox longifolia;  Phlox longifolia, Nuttall, 1834;  

    Phlox longifolia Nutt. “Longleaf Phlox”

    There were two PhloxPhloxes ? — that were reported for Golden s.l. They are Phlox longifolia Nutt. ldquo;Longleaf Phlox” and P. multiflora A. Nelson “Mountain Phlox.” As it happened, both were found only on the north slopes of North Table Mountain. And with two doubtful exceptions, these are the only Phlox collected in Jefferson County. See collection data in SEINet (2019+). The first exception is an undated collection of Phlox caespitosa by E. S. Greene on “… barren ground above trees … Golden” (PAC29495) If this collection was truly above the trees, as P. caespitosa often is, then it was collected at higher elevations, west of and outside Golden s.l. The collector was more likely E. L. Greene. The second exception is a collection by C. S. Crandall, #1589, May 19, 1894, Platte Canyon, Jefferson County, some distance south of Golden s.l. One voucher (RM16725) is determined P. patula A. Nelson. A second voucher (CS16725 -- the same ascension number at two herbaria also seems suspicious) is determined P. multiflora A. Nelson.

    Literature Cited:
    - Nuttall, Thomas, 1834a.
    - SEINet, 2019+.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1834a, publication details;  Notes on Phlox longifolia;
    • U. S. Highway 20:   at ID 33 W;
    • Wyoming Highway 354:   at Old Ft Bonneville;

    Locations: Henrys Fork. Old Fort Bonneville Site.  

    Historically, Phlox longifolia was described by Thomas Nuttall (1834a) from plants collected by Nathaniel Wyeth in July 1833 between the Henry Fork of the Snake River and and Bonneville's Fort at the junction of Horse Creek and the Green River. The plant is widely distributed in the American Cordillera.

    Original Text Comments
    73. Phlox *longifolia. Subcaespitosa-multocaulis, foliis subulatis longissimis angustissimis glabrus, cauliculi pauciflori brevissimi puberuli irregulariter trichotomi, pudunculis filifornibus elongatis, laciniis calycinis acuminatis, corollae laciniis oblongo-cuneatis integris.  
    ♃ The stems almost a span high, many from the same root, clothed below with the withered vestiges of former leaves. The leaves smooth, and narrow as threads, two and a half or so inches in length, those of the sterile branches extending nearly the length of the short and slender flower stems; peduncles very long and slender. Flowers apparently white. Allied to P. Hoodii, but very distinct.  
    Hab. In the valleys of the Rocky Mountains generally, flowering for the most part of the summer.  

     

    Other articles:
    • Golden Checklist Flora:  Phlox multiflora;  

    Phlox multiflora A. Nelson. “Mountain Phlox”

     

    Literature Cited:
    - Nelson, Aven, 1898a.  

    Nelson (1898a, p. 278) described Phlox multiflora as a segregate from P. longifolia when describing new plants from Wyoming.

    Original Text Comments
    Phlox multiflora  
    Suffrutescent as to the caespitose, much branched, prostrate base, the numerous herbaceous stems nearly erect, 6-10 cm. high the annual branches simple, one-flowered ; leaves broadly linear, glabrous, apiculate, 1-2 dm. long, opposite of fascicled ; peduncles finely pubescept, 1-3 cm. long ; calyx angled by the prominent midrib of the lobes, membranous in the sinus only, lobes linear-spiculate, equalling the tube ; tube of the corolla exceeding the calyx, lobes obovate, entire, 1 cm. long ; style equalling the calyx.  
    The affinities of this plant are with the P. longifolia group. It has been distributed under no. 182 from Laramie Hills, mostly as P. longifolia Nutt, but from which, I now think, it is wholly distinct. Its more caespitose growth, shorter broader leaves, simple branches, less membranous calyx and numerous flowers easilt separate it. It is very abundant on the moister slopes and valleys of the Laramie Hills at 8000 to 9000 ft., flowering from late May till late June.  
    Type specimen in Herb. University of Wyoming, no. 3175, June 16, 1897.  

     

    Other articles:
    • Golden Checklist Flora:  Polemonium foliosissimum;  

    Polemonium foliosissimum A. Gray “Towering Jacob's Ladder”

     

    Literature Cited:
    - Bentham, George, 1845.  

    Bentham in DeCandolle (1845, v. 9, p. 317) ....

    δ? pterosperma , seminibus apice et basi latiscule membranaceo-alatis. In montibus Americae boreali-occidentalis (Drummond !). An sp. propria ? (v. s.) δ? pterosperma , seeds laterally membraneous-winged at apex and base. In northwestern American mountains (according to Drummond). It own species ? I have seen a dried specimen.

    Literature Cited:
    - Gray, Asa, 1870.  

    Gray (1870, p. 281) ...

    2. P. caeruleum, L. Common from the arctic regions and Alaska to California and through the Rocky Mountains, also through Northern Asia to Europe; very rare eastward (in New York and New Jersey). — P. acutiflorum, Willd., which is reduced by Ledebour to a variety of this species, is an Alaskan form, with ovate acute lobes to the corolla (Pallas, Chamisso, &c). All the North American, like the Himalayan, forms of this species incline to have wing-angled seeds, — quite as much so as in
    Var. foliosissimum (P. caeruleum, var. pterosperma, Benth. in DC. Prodr.). Valde viscido-pubescens ; caulibus bipedalibus usque ad apicem cum ramis floridis corymbo-is foliosissimis; foliolis in rhachin alato-marginatam saepe confluentibus ; floribus minoribus; staminibus styloque corolla (calycem 2 - 3-plo superante) saepius brevioribus. — Through the Rocky Mountain region, Geyer, Fendler, Parry, Vasey, Watson, &c. This approaches
    3. P. Mexicanum, Cerv. (Mexico ?) This is distinguished by its …

    Literature Cited:
    - Gray, Asa, 1878-1895.  

    Gray (1878, v. 2, pt. 1, p. 151) elevated our plant to rank of species ...

    P. foliosissimum. Very viscid-pubescent throughout and strong-scented : stem a foot or more high, very leafy throughout; leaflets from lanceolate to ovate-lanceolate (seldom an inch long) : flowers corymbose-cymose, smaller than those of the preceding : corolla commonly white or cream-color, sometimes violet, twice the length of the calyx, which is 5-cleft to or beyond the middle: style and stamens not protruding. — P. caeruleum, var.? pterosperma, Benth. in DC. Prodr. ix. 317; but the seeds, as in P. caeruleum, are either marginless or wing-margined. P. carruleum, var. foliosissimum. Gray, Proc. Am. Acad. viii. 281. — Rocky Mountains of New Mexico, Colorado, and Wyoming, and west to Utah and Idaho. Some forms approaching the preceding species [P. caeruleum] ; but it is more like P. Mexicanum, Cerv., which is loosely branched, and has the violet corolla little exceeding the calyx, the lobes of the latter barely half the length of the tube.
    → → Ovules only 3 or 4 in each cell : stem lax or with diffuse branches and open corymbiform or paniculate inflorescence: leaflets fewer (5 to 15) and rather large, membranaceous, only the ultimate at all confluent: herbage glabrous or slightly pubescent, neither viscid nor glandular: style and stamens rather shorter than the corolla.

       

    Boraginaceae

     

    Literature Cited:
    - Greene, Edward L., 1887.  

    In a footnote to his “Some North American Asperifoliae,” Greene (1887) explains about his use of the name for a family we now universally call Boraginaceae

    Original Text
    1. Ordinal names in botany, no less than the generic and specific ought, it seems to us, to be received according to priority. The one here written was proposed by Haller, accepted by his contemporaries, including unch men as Dillenius, and has never yet been quite displaced by the more recent Jussiaean name Borragineae ; for even that most eminent writer on the order, Lehmann, continued always to use the older name, notwithstanding that the DeCandolles in their great general work, destined to wield such wide and lasting influence, chose to support the more modern one. I am glad that, among contemporary authorities, one of Baron von Muller's great fame adopts the original and, I may add, the most appropriate and convenient ordinal appellative for these plants. It is one which, like Cruciferae, Compositae and others, has the literary advantage of not ending in that awkward combination of successive vowles which is a serious objection against many of the names of comparatively recent date.

     

    Other articles:
    • Golden Checklist Flora:  Asperugo procumbens;  

    Asperugo procumbens L. “German Madwort”

     
     

    Linnaeus (1753, v. 1, p. 138) ...

    Original Text Interpretation
    procumbens. 1. ASPERUGO. Fl. lapp. 76. Fl. fuec. 159. Hort. cliff. 44. Roy. lugdb. 404. Hall. helv. 522. Dalib. parif. 61  
      Bugloffum fylveftre, caulibus procumbentibus. Bauh. pin. 257. Morif. hift. 3. p. 439. f. 11. t. 26. f. 13.  
      Habitat in Europae ruderatis pingvibus. ☉ It lives in fertile waste places. Annual.

     

    Literature Cited:
    - Gmelin, Johann Friedrich, 1788-1793.
    - Walter, Thomas, 1788.

    Other articles:
    • Golden Checklist Flora:  Notes on Lithospermum incisum, Pursh, 1814;  

    Batschia J. F. Gmel.

    Gmelin (1791, t. 2(1), p. 315) published Batschia and B. carolinensis in his 13th edition of Linneaus' Systema Naturae

    Original Text Translation and Comments
    BATSCHIA. Cor. hypocrateriformis: tubo baft lanofo. Stigma capitatum. Nuces? 4 imperforatae, fubovatae, arcuatae, acuminatae. BATSCHIA. Corolla hypocrateriform [salverform] : tube base lanose. Stigma capitate. Seeds 4, imperforate, subovate, arcuate, acuminate.
    carolini-
    enfis.
    1. Batschia. Walt. flor. Carol. p. 91. carolini-
    ensis.
    1. Batschia. Walter, Thomas. 1788. Flora Caroliniana. London: 1788. p. 91.

    Batschia caroliniensis was first published as Anonymos caroliniensis by Walter (1788). We now treat B. caroliniensis as Lithospermum caroliniense (J.F.Gmel.) MacMill.

     

    Other articles:
    • Golden Checklist Flora:  Cryptantha virgata;  

    Cryptantha virgata (Porter) Payson “Miner's Candle”

     

    Literature Cited:
    - Porter, Thomas C., 1870.

    Other articles:
    • Golden Checklist Flora:   Hayden Report;  

    Porter (1870, pp. 479-480) … in what is generally known as the “Hayden Report” …

    Eritrichium virgatum, sp. nov. Sesquipedale, hispidissimum ; caule simplici tereti erecto; foliis radicalibus oblongo-spathulatis, caulinis linearibus; cymis plurimis in axillis foliorum conglomeratis pedunculatis superne subsessilibus confertisque ; calycis 5-partiti segmentis lanceolatis tubum corollae aequantibus ; nuculis 4-ovatis dorso infra partem mediam convexo supra lateribus depresso laevibus nitidis. — Near Denver, Colorado Territory, 1869, B. H. Smith. The stout virgate spike made up of numerous glomerate cymes, crowded in the axils of the linear cauline leaves, which much exceed them in length. Limb of the corolla 4 lines in diameter, its lobes rounded. Nutlets smooth and shining, the lower half of the back convex, the upper depressed on the sides, leaving a central vertical ridge. According to Dr. Gray this plant is the same as Parry's 288, and Hall and Harbour's 438 (in part.) It has passed for a form of E. glomeratum, DC, but seems more nearly allied in its fruit and habit of growth to E. leucophaeum, A. DC. (Myosotis leucoplicea, Dougl., in Hook. Fl. Bor. Am. 2. p. 82. t. 163,) and I have ventured to give it a name.

    Literature Cited:
    - Porter, Thomas C., and John M. Coulter, 1874.

    Other articles:
    • Glossary:  virgate;

    Locations: Clear Creek Canyon. Colorado Springs. Ute Pass.  

    Porter & Coulter (1874, p. 102-103) in their Synopsis of the flora of Colorado treated our plant as a variety of Eritrichium glomeratum.

    Original Text Comments
    Var. VIRGATUM, Porter, (E. virgatum, Porter, Hayden's Rep. 1870, p. 479.) Stout; stem erect, 2°-3° high, very hispid; inflorescence much elongated, virgate, spike-like, 1°-2° in length; cymes many, conglomeated, few-flowered, lower ones short-peduncled, upper ones sessile, very much shorter than the long, spreading, bracteal leaves, which become narrowly linear above. &mdasdh; This variety is verv common on the plains at the base of the monutains and among the foot-hills. — Colorado Springs, Monument Park, and Ute Pass, Porter. Clear Creek Cañon, Coulter. There are five locations known as Ute Pass in Colorado. Assuming this collection was made in Teller County at the town now known as Divide.

    There are three Monument Parks in Colorado. It will be necessary to trace out the route of Hayden's party to narrow down which park they were in which Porter made his collection.

    Coulter's s.n. collection in Clear Creek Canyon is NY3224955, an image of which is available via SEINet. The elevation is given as 9000 ft., so the collection was probably made well above Golden s.l.

    Literature Cited:
    - Payson, Edwin Blake, 1927.  

    Payson (1927, p. 270) published Cryptantha virgata new comb. While the material is copywrited, it is licensed by Creative Commons (CC BY-NC-SA 3.0). The description is lengthy (2½ pages) and informative parts are reproduced here.

    Payson (1927, v. 14, p. 270-272) …

    Original Text
    13. C. virgata (Porter) new comb. Plate 26, figs. 32-34
    Eritrichium virgatum Porter, Hayden Rept. 479. 1870.
    E. glomeratum var. virgatum Porter in Porter & Coulter, Syn. Fl. Colo. 102. 1874.
    Krynitzkia virgata (Porter) Gray, Proc. Am. Acad. 20: 279. 1885.
    Oreocarya virgata (Porter) Greene, Pittonia 1: 58. 1887.
    O. spicata Rydb. Bull. Torr. Bot. Club 36: 678. 1909. (Type: Artist's Glen, Pikes Peak, Colorado, Aug. 1, 1901, Clements 102.)
    O. virgata forma spicata (Rydb.) Macbr. Proc. Am. Acad. 51:546. 1916.
    [Description omitted.]
    Distribution: Transition Zone, eastern foothills of the Rocky Mountains, southeastern Wyoming to south-central Colorado; also in North Park, Colorado. Type: "near Denver, Colorado Territory, 1869," B. H. Smith.
    Specimens examined: … [Omitted]
    C. virgata is one of the most conspicuous herbaceous plants on the eastern foothills of the Rocky Mountains in eastern Colorado. The strictly erect, rod-like stems with the closely set white flowers are unique. The numerous elongated leaf-like bracts of the in- florescence add to the peculiar appearance and serve to separate the species from its relatives. Its relationship to C. setosissima is decidedly problematical. Although it was for a time held to be a variety of Bradburiana there is no reason to believe it is really related to that plant.
    Additional evidence is at hand to support Macbride's contention (Proc. Am. Acad. 51: 546. 1916) that O. spicata Rydb. is not even deserving of varietal rank. It is simply to be regarded as a form of virgata. Thanks to the help of Mr. W. T. Penfound, of the Alpine Laboratory on Pikes Peak, specimens have been examined that prove the existence of typical virgata from that region as well as the smooth-fruited form.

    Literature Cited:
    - Mabry, Makenzie E., and Michael G. Simpson, 2018.  

    Mabry & Simpson (2018) found continued phylogenetic support for treatment of Oreocarya at the rank of genus, separate from Cryptantha s.l.

     

    Other articles:
    • Golden Checklist Flora:  Cynoglossum officinale;  

    Cynoglossum officinale L. “Gypsyflower” or “Houndstongue”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p. 134) ...

    Original text Comments
    officinale. 1. CYNOGLOSSUM flaminibus corolla brevioribus, foliis lato-lanceolatis tomentofis feffilibus.  
      Cynoglofum foliis lanceolato-ovatis, corollis calycem aequantibus. Roy. lugdb. 406. Fl. fuec. 154. Mat. med. 154.  
      Cynoglofum foliis ovato-lanceolatis. Hort. cliff. 47.  
      Cynoglofum majus vulgare. Bauh. pin. 25.  
      Habitat in Europae ruderatis. The Latin “ruderatis” literally translates to “rubbish.” It probably means “waste places” in this context. This suggests it was recognized as weedy in its native habitat. POWO states the plant is native is Europe, to west and south Siberia, and Iran.

     

    Other articles:
    • Golden Checklist Flora:  Ellisia nyctelea;  

    Ellisia nyctelea (L.) L. “Aunt Lucy”

     

    Literature Cited:
    - Gronovius, Jan Frederik, and John Clayton, 1738-1743.  

    Linnaeus (1752, v. 1, p. 160) ...

    Original Text Translation and Comments
    Nyctelea. 2. IPOMOEA foliis pinnatifidis finuatis acutis.

    Habitat in Virginia. D. Gronovius.

    Gronovius, Jan Frederik, and John Clayton. 1738-1743. Flora Virginica. 2 vols. I do not see our plant in Gronovius's volume.
     

    Linnaeus (1762, v. 1, p. 232) ...

    Original Text Translation and Comments
    5. POLEMONIUM ? foliis pinnatifidis acutis dentatis, caule diffufo.

    Ipomoea Nyctelea. Sp. pl. 1. p. 160.

    Scorpiurus humilis virginianus , foliis rutaceis. Morif. hift. 3. p. 451. f. 11. t. 28. f. 3.

    Habitat in Virginia.

    Caulis herbaceus, dochotomus, angulatus, diffufus. Folia alterna, petiolata, pinnatifida, fere Hydrophylli: lacinis acutis, utrinque dente angulatis. Pedunculi oppofitifolii, uniflori, pubefentes, elongati. Calyx monophyllus, quinquefidus ultra medium, acutus, corolla longior, patens. Corolla monopetala, campanulata, quinquefida. Stamina 5. Pistillum 1. Fructus fphaericus. Facies Solani peruviani f. radicantis, fed corolla campanulata; forte proprii generis planta.

    Nyctelea. Perhaps its own genus.

    Literature Cited:
    - Linne, Carl von, and Lars Salvius, 1763.  

    Linnaeus (1763, v. 2, Appendix, p. 1662) ...

    Original Text Translation and Comments
    Nyctelea. 271. ELLISIA. (Pentandra, Monogyna)
    Polemonium Nyctelea. Sp. pl. 2. p. 231.
    Proprii generis planta eft.
    This plant in the correct genus.

     

    Other articles:
    • Golden Checklist Flora:  Heliotropium curassavicum;  

    Heliotropium curassavicum L. “Seaside Heliotrope”

     

     

    Other articles:
    • Golden Checklist Flora:  Hydrophyllum fendleri;  

    Hydrophyllum fendleri (A. Gray) A. Heller “Fendler's Waterleaf”

     

     

    Other articles:
    • Golden Checklist Flora:  Lappula occidentalis;  

    Lappula occidentalis (S. Watson) Greene “Flatspine Stickseed”

     

    Literature Cited:
    - Watson, Sereno, 1871.  

    Watson (1871, v. 5, p. 246)

    Original Text
    Echinospermum Redowskii, Lehm. DC. Prodr. 10. l37. Stem erect, pubescent, paniculately branched; leaves linear or sublanceolate, hoary with spreading hairs; calyx-lobes narrow-linear, equaling the corolla-tube; nutlets compressed, surrounded by a single row of barbed prickles, muriculate-rugose upon the back and sides, shorter than the enlarged calyx. Var. occidentale. (E. Redowskii, Gray.) The tubercles, which are irregularly and thickly scattered over the faces of the nutlet, very sharply acute instead of rounded- obtuse as in Asiatic specimens. — Quite variable in its habit ; from 3'— 2° high, much branched at base and ascending, or with a single erect virgate stem; leaves and brads usually linear-oblong, not unfrequently ovate-oblong or spatulate, always obtuse: flowers small, but little exceeding the calyx, blue; the prickly margin more or less contracted over the back of the nutlet, and the prickles more or less confluent. E. patulum, Lehm., of Western Asia, to which this plant was at first referred, differs from E. Redowskii (as shown by specimen's in Herb. Gray.) only in the tuberculations upon the fruit, which in the former species are few in number, arranged regularly in longitudinal rows upon the back and upon the outer edge of the sides, and armed with curved points. The differences are represented with tolerable accuracy in the plate. From Western Texas to Arizona and northward to the Saskatchewan, Bear Lake and Fort Youkon. Frequent in the valleys and on the mountains from the Sierras to the Wahsatch ; 4-8,000 feet altitude ; May-July. Plate XXIII. Figs. 9, 10. Achenium of E. Redowskii, Var. occidentale. Figs. 11, 12. Achenium from Asiatic specimen of E. Redowskii. Figs. 13, 14. Achenium of E. patulum ; all enlarged eight diameters. (861.)
    Var. strictum. (E. strictum, Nees., App. Neuwied's Trav. 17; not of Ledebour.) The prickles united over the back into a strongly inflexed border. — An extreme state, but various intermediate forms are frequent. New Mexico to Colorado and westward. Valleys and foot-hills of Western Nevada, with the last. (S62.)

    Literature Cited:
    - Greene, Edward L., 1899.  

    Greene (1899, v. VI, p. 97) in a review of western aAmerican Asperifoliae (Boraginaceae),

    Original Text
    In naming and defining the following species of Lappula, several of which are of what may be called the cupulate group, I make no attempt to continue in use Gray's varietal name cupulatum, for that was made to include, as one variety of L. Redowskii, a number of easily definable species; and there is no determining to what one of the segregate species the name should be applied rather than to another. Moreover, the earliest known cupulate species obtained specific rank far anteriorly to the publication of L. Redowskii, var. cupulatum ; I refer to
    ... [Seven species descriptions omitted.] ...
    L. occidentalis. Echinospermum Redowskii, var. occidentale, Wats. Bot. King Exp. 246, in part at least. This is very widely dispersed from toward the Rocky Mountains almost to the Pacific coast ; is well marked among the species here defined by its 7 to 11 strongly developed marginal prickles usually of triangular-subulate form, lightly or not at all connected at base, and, I believe always, grooved or channeled down the whole inner face ; the surface of the nutlet being tuberculate, not muricate.
    There is no evidence of the occurrence, anywhere in America, of the true L. Redowskii.

    Literature Cited:
    - Baldwin, Bruce G., Douglas H. Goldman, David J. Keil, Robert Patterson, and Thomas J. Rosatti, 2012.  

    Baldwin, et al. (2012) in The Jepson Manual: Vascular Plants of California, Second Edition treat our plants as varieties of Lappula redowskii, and my collections from California continue to reflect that treatment.

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittmann (2012) treat our plant as L. redowskii without infraspecific nomenclature.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015) treats our plant as L. occidentalis with infraspecific names of occidentalis and cupulata.

    Literature Cited:
    - Khoshsokhan-Mozaffar, Maryam, Mahboubeh Sherafati, and Shahrokh Kazempour-Osaloo, 2018.  

    Khoshsokhan-Mozaffar, et al. (2018) prepared a molecular phylogeny of the tribe Rochelieae with special reference to Lappula. Their text does not specifically mention the relationship between Lappula occidentalis and L. redowskii. However, their trees place the North American Lappula in their own clade including L. occidentalis, separate from the Asian clades including L. redowskii.

    Literature Cited:
    - POWO, 2021 - 2023.  

    Plants of the World (2021) treats L. occidentalis at the specific level, with varieties occidentalis and cupulata.

     

    Other articles:
    • Golden Checklist Flora:  Lithospermum arvense;  

    Lithospermum arvense L. “Corn Gromwell”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p. 132) ...

    Original Text Comments
    arvenfe. 2. LITHOSPERMUM feminibus rugofis, corollis vix calycem fuperantibus. Hort cliff. 46. Fl. fuec. 152. Roy. lugdb. 405. Hall. helv. 518. Dalib. parif. 59.  
      Lithofpermum arvenfe, radice rubra. Bauh. pin. 258. Fl. lapp. 73.  
      Habitat in Eurpoae agris & arvis. Habitat in European fields and countryside.

     

    Other articles:
    • Golden Checklist Flora:  Lithospermum incisum;  

    Lithospermum incisum Lehm. “Narrowleaf Stoneseed”

     

    Literature Cited:
    - Pursh, Frederick, 1814.

    Other articles:
    • Golden Checklist Flora:  Notes on Batschia, Gmelin, 1791;  

    Pursh (1814, v. 1, p. 132) published the first name for our plant, Batschia longiflora.

    Original Text Translation
    longiflora. 3. B. sericeo-villosa ; foliis linearibus, calycibus longis linearibus, laciniis corollae crenato-incisis, tubo elongato.

    On the banks of the Missouri. Nuttall. ♃. July. v. s. in Herb. Nuttall. Flowers yellow.

    longiflora. 3. Batschia sericeous-villous ; leaves linear, calyx long linear, corolla tube elongated, lobes crenate and incised.

    On the banks of the Missouri. Nuttall. Perennial. Flowers July. I have seen it dried in Nuttall's Herbarium. Flowers yellow.

    Unfortunately, the specific epithet longiflora will not be available in Lithospermum having been previously used by Salisbury (1786).

    Literature Cited:
    - Lehman, Johann G. C., 1818.  

    Lehman (1818, v. 2, 303) …

    Original Text Translation
    217. LITHOSPERMUM INCISUM. 217. LITHOSPERMUM INCISUM.
    L. caule herbaceo, foliis linearibus sericeo-villosis, corollae tubo elongato, limbi laciniis crenatopincisis. Lithospermum with herbaceous stems, leaves linear silky-villous, tube of the corolla elongated, lobes crenate-incised.
    Batschia (longiflora) sericeo-villosa, foliis linearibus, calycibus lingis linearibus, laciniis corollae crenato-incisis, tubo elongato. Pursh. Fl. Americae septentr. Vol. 1. pag. 132. Batschia ( longiflora ) silky-villous leaves are linear, ealyx lobes linear, corolla lobes crenate and cut, the tube elongated. Pursh. Fl. Americae septentr. Vol. 1. pag. 132.
    Habitat ad Missuri fluvium Americae borealis. ♃ Lives along the Missouri River of North America. Perennial.
    Obs. Ab omnibus satis distincta videtur corollis crenato-incisis. Speciei nomen longuflorae mutandum esse putavi, quum nunc ad Lithospermorum genus relatae flores habeant longiasimos. It seems quite distinct from all pale crenato-serratum saplings. Longuflorae type a name change to be expected. [Because it was previously used by Salisbury in 1786. — Ed.] Lithospermum kind of in a manner related to the flowers when they have a longiasimos now.
    Lithospermum incisum Lehm., Pl. Asperif. Nucif. 2: 303 (1818). Plantae e Familiae Asperifoliarum Nuciferae

    Replaced synonym Batschia longiflora Pursh, Fl. Amer. Sept. (Pursh) 1: 132 (1813).

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Locations: Prairie du Chien.  

    Nuttall (1818, v. 1, p. 113-114) …

    Original Text
    171. BATSCHIA. Gmelin. Michaux.
    Calix 5-parted. Corolla salverform, rather large, tube straight, much longer than the calix, closed at the base by a bearded ring; orifice naked or partially closed; border orbiculate, nearly flat, segments rounded. Seed indurated, shining, (as in Lithospermum.)
    Flowers yellow or fulvous, axillary, almost fastigiate, in short bracleate spikes; leaves narrow, without collateral nerves.
    Species.

    1. B. Gmelini.

    2. canescens. Flowers fulvous.

    3. *longiflora. Hirsutely villous, erect; leaves approximating, long and linear, margin reflected; flowers in a fastigiate fascicle; tube of the corolla somewhat pentangular, (often from 10 to 15 lines long) border flat, segments fimbriate-crenate. — Flowers sulphur yellow. In open plains; around the Prairie du Chien, Missisippi, and on the banks of the Missouri to its sources.

    4. *decumbens. Hirsutely villous; stem decumbent; segments of the calix and leaves linear; flowers scattered; lobes of the corolla fimbriate-crenate, shorter than the tube. — Around the Mandan village. Nearly allied to the preceding. In both these species the orifice is partly closed by 5 arched protuberances.

    All the species of this genus afford a crimson lac from the root.

    Prairie du Chien is on the Mississippi River in southwest Wisconsin.

    Batschia decumbens is now treated as a synonym of Lithospermum incisum Lehm.

    I would hazard a guess that Nuttall actually first applied the name B. longiflora to our plant, but that Pursh (1814) beat him into print. In the meantime, Lehman (1818) of the Hamburg Botanical Garden published L. incisum without stating whether he had seen the plant or not.

     

    Other articles:
    • Golden Checklist Flora:  Lithospermum occidentale;  

    Lithospermum occidentale (Mack.) Weakley, Witsell & D. Estes “Western Gromwell” nee “Marbleseed”

     

    Literature Cited:
    - Gray, Asa, 1878-1895.  

    Gray (1878, v. 2(1), p. 206) published O. Carolianum var. molle

    Original Text
    O. Carolinianum, DC,, l. c. Stout, 2 or 3 feet high, shaggy-hispid : leaves ovate-lanceolate and oblong-lanceolate, acute, 5-9-ribbed, generally hairy both sides : flowers nearly sessile : corolla short (4 or 5 lines long), yellowish-white, oblong-funnelform ; its ovate-triangular acute lobes very hairy outside, and nearly half the length of the tube. — Lithospermum Carolinianum, Lam. Ill. & Diet. Suppl. ii. 837. Purshia mollis, Lehm. Asper. 383. — Alluvial grounds, Upper Canada to Georgia and Texas.
    Var. molle. A foot or two high ; the pubescence shorter and less spreading or appressed : leaves mostly smaller (about 2 inches long), when young softly strigose-canescent beneath. — Onosmodium molle, Michx. Fl. i. 133, t. 15 ; Gray, Man. ed. 5, 362. Purshia mollis, Lehm. Asper. 382. — Illinois to Saskatchewan, Utah, and Texas.

    Literature Cited:
    - Mackenzie, Kenneth Kent, 1905.  

    Kenneth Kent Mackenzie (1905, v. 32, p. 502-3) published what is now the basionym of Lithospermum occidentale.

    Original Text
    5. Onosmodium occidentale nom. nov.
    Onosmodium carolimaniun molle A. Gray, Syn, Fl. 21 : 205. 1878, and later American authors; not Onosmodium molle Michx., vide supra.
    Usually only 3-6 dm. tall, but occasionally 10-12 dm., branching above or often from near the base, the branches spreading or almost erect ; plant clothed all over with a silvery-white or sometimes a slightly yellowish appressed or more or less spreading canescent pubescence (in northern and mountain plants sometimes quite green and noticeably spreading) usually not exceeding 2 mm. in length, the pubescence of the branches and leaves usually appressed: leaves typically about 5 cm. long and 1.5 cm. wide, acutish, prominently 5-7-nerved on both surfaces, the pubescence sometimes strongly spreading and papillose, but most typically soft, appressed and hardly papillose : bracts 6-24 mm. long, often 2-ranked, resembling the leaves in outline : fruiting pedicels from very short to 6 mm. long : calyx-lobes lanceolate, acute, acutish or obtusish, rarely linear-oblong, 6- [2 mm. long, canescent or with somewhat spreading hairs; corolla 12-20 mm. long, canescent all over outside, the broadly triangular acute lobes (3-4 mm. long) usually with apical tuft of hairs not strongly noticeable: nutlets 3.5-4 mm. long, ovoid, acutish or acute, not constricted at base, dull in color, little if at all pitted.
    This is the common Onosmodium of the western part of the country and the most common species in herbaria. It has heretofore been referred to O. molle Michx., a species which as already related I believe to be distinct. It ranges from Illinois to Athabasca, Alberta, Montana, New Mexico and Texas, and seems to be a fairly common plant in all this vast extent of country.
    Specimens examined. [Reduced to Colorado specimens only. — Ed.]
    Colorado: Hulse, 1870; Britton, 8 October 1882, Golden ; Osterhout 2461, 25 June 1901, Boulder ; J. H. Cowen, 10 July 1S93, Ft, Collins ; Jones, 12 June 1878, Platte River, Denver ; Engelmann, 6 August 1874, Empire ; Brandegee 9 July 1873, Cañon City ; Cowen, 25 June 1895.
     

    O. molle Michx. var. occidentale (Mack.) I. M. Johnston

    I. M. Johnston (1924, v. 70, p. 18) published O. molle var. occidentale ...

    Original Text
    7. O. molle Michx. Fl. Am. Bor. i. 133, t. 15 (1803); Mack. Bull. Torr. Bot. Cl. xxxii. 499 (1905). Lithospermum molle Muhl. Cat. 19 (1813). Purshia mollis Lehm. Asperif. ii. 383 (1818). O. cariolinianum, var. molle Gray, Synop. Fl. N. A. ii, pt. 1, 206 (1878). — In its extreme form known only from the vicinity of Nashville, Tenn.
    7a. O. molle, var. occidentale (Mack.), comb. nov. O. occidentale Mack. Bull. Torr. Bot. Cl. xxxii. 502 (1905). O. occidentale, var. sylvestre Mack. l. c. 504. — Minnesota and Illinois, westward to Texas, New Mexico, Utah, and Saskatchewan. This is the common Onosmodium west of the Mississippi River. The plant is not accorded specific rank due to the material from Illinois which shows repeated tendencies to intergrade with O. molle. The variety usually differs from the typical form of the species in its coarser stems, rough spreading pubescence, larger calyces (8-12 mm. long), and unpitted non-constricted nutlets, but these characters and their complements occur in various combinations in the collections from Illinois.

    Literature Cited:
    - Harrington, H. D., 1964, 2nd ed..  

    Harrington (2nd ed., 1964) treats our plant as Onosmodium molle Michx. var. occidentalis (Mack.) Johnston.

    Literature Cited:
    - Cochrane, Theodore S., 1976.  

    O. molle subsp. occidentale (DC.) Cochrane

    Onosmodium molle subsp. occidentale (DC.) Cochrane, Michigan Bot. 15: 104 (1976).

    Literature Cited:
    - Turner, Billie L., 1995.  

    O. bejariense A. DC. var. occidentale (Mack.) B. L. Turner

    B. L. Turner (1995, v. 78, p. 46-47) ...

    Original Text
    ONOSMODIUM BEJARIENSE A. DC. VAR. OCCIDENTALE (Mack.) B.L. Turner, comb. nov. BASIONYM: Onosmodium occidentale Mack., Bull. Torrey Bot. Club 32:502. 1905. Onosmodium molle Michx. var. occidentale (Mack.) I.M. Johnst., Contr. Gray Herb. n.s. 70:18. 1924. Onosmodium molle Michx. subsp. occidentale (Mack.) Cochrane, Michigan Bot. 15:104. 1976. TYPE: U.S.A. Nebraska: Kearney Co., "Prairie", 13 Jun 1891, P.A. Rydberg s.n. (LECTOTYPE: NY!, selected by T. Barkley but first published here).
    My concept of this taxon is essentially the same as that of Das (1976). It occurs in the drier, more northwestern regions of the broad distribution of Onosmodium bejariense. Var. occidentale appears to intergrade into the bejariense in northcentral Texas, central Oklahoma, western Arkansas, northern Missouri, Illinois, and Minnesota.
    REPRESENTATIVE SPECIMENS: This variety, for the most part, is well-marked; in Texas, it is known only from Wichita and Tarrant counties (cf. discussion under var. bejariense).
    DISTRIBUTION (Figure 2) AND ECOLOGY: Northcentral Texas westwards to Colorado, Montana, and Canada, eastwards to Illinois and Wisconsin, mostly silty-clay or sandy soils; flowering May- July.

    Literature Cited:
    - Weigend, M., M. Gottschlinf, F. Selvi, and H. H. Hilger, 2009.  

    Weigend, et al. (2009) placed all the Onosmodium in Lithospermum to make that group monophyletic.

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittmann (2012) treat our plant as Onosmodium molle Michaux subsp. occidentale (Mackenzie) Cochrane.

     

    Ackerfield (2015) treats our plant as Onosmodium bejariense DC. ex A. DC. var. occidentale (Mack.) B. L. Turner (Syn: Onosmodium molle Michx. var. occidentale (Mack.) I. M. Johnst.)

     

    Other articles:
    • Golden Checklist Flora:  Mertensia lanceolata;  

    Mertensia lanceolata (Pursh) DC. “Prairie Bluebells”

     
     

    Pursh (v. 2, Supplementum, p. 729) ...

    Original Text
    Pulmonaria lanceolata. — P. glabra, erects ; foliis radicalibus longissime pedicellatis lanceolatis, caulinis linearipoblongis, floribus subpaniculatis, calycibus abbreviatis.

    In Upper Louisiana. Bradbury.v. s. in Herb. Bradbury. Flowers small, resembling those of P. paniculata.

    p. 131.

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, v. 1. p. 115) ...

    Original Text
    173. PULMONARIA. L. (Lungwort.)
    Calix prismatic-pentagonal. Corolla funnel-formed, somewhat 5-lobed, orifice open. Stigma emarginate.
    Flowers almost disposed in terminal corymbs, sometimes racemose. Leaves in the American species and in the P. maritima smooth and glaucous, calix short, about half the length of the corolla tube. (Mertensia Persoon.)
    Species. 1. P. virginica. 2. paniculata. 3. *marginata. Small, glabrous, erect; leaves hispid on the margin, on the lower part of the stem oblong'-spalhulate or oblong-ovate, upper leaves ovate, acute, semiamplexicaule; flowers subpaniculate; fasciculi few-flowered, axillary and terminal; calix acute, about half ihe length of the corollii tube.
    P. lanceolata. Pursh, Flor. Am. 2. p. 729.
    Perennial. Stem nearly erect, and somewhat branched, scarcely a foot high. Radical leaves petiolate, stem leaves sessile, uppermost semiamplexicaule, all glabrous, glaucous, and somewhat carnose. Fascicles or ramuli, pedicellate, a little longer than the leaves, 4, 5, 8, or more flowered. Calix 5-parted, smooth, acute, segments somewhat hispid on the margin. Corolla funnel-formed, much like that of P. virginica, and of the same colour, but smaller, lobes entire. Stamina seated around the orifice of the tube.
    Nearly allied to P. virginica, and also probably to the P. paniculata, but the leaves are never acuminated or hairy, but perfectly smooth, except on the margin where there are a few scattered hooked, hispid hairs, often only visible in a dried state. The name of lanceolata is so very inapplicable that it appeared to me necessary to alter it.
    On arid hills near the confluence of Teeton river, Missouri. Flowering in June.
    4. paniculata. 5. sibirica.
    The only 3 genuine species of Pulmonaria described are confined to Europe; the American species all arrange under the subdivision Mertensia of Persoon.

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1836.  

    DeCandolle (1846, v. 10, p. 88) ...

    Original Text
    2? M. lanceolata (DC. mss.) , caule erecto glabro , foliis glaucescentibus carnosulis ciliatis , iaferioribus oblongo-spathulatis vel oblongo-ovatis , sup. ovatis acutis semi-amplexicaulibus , raoemis 4-8-floris subfasciculatis folio sublongioribus, calyce 5-partito ciliato lobis acutis, corolla infundibuliformi tubo calycem duplo superante. ♃ in Missouri (Nutt.), Louisiana sup. (Pursh) et frequens ad Oregon (Dougl.). Pulmonaria lanceolata Pursh fl. 2. p. 729. Pulmonaria marginata Nutt. gen. 1. p. 115. Lithospermum marginatum Spreng. syst. 1. p. 547. Hook. et Arn. fl. n. Amer. 2. p. 86. Casselia lanceolata Dumort. obs. p. 24. Caulis vix pedalis. Flores pedicellati. Corolla colore et fere forma corollae M. Virginicae. Stamina fauci inserta. Nomen specif. mutavit cl. Nutt. quia folia non lanceolata , sed non melius marginata ( nempe margine revoluta aut nervosa) et lobi calycis fere lanceolati videntur. (Alph. DC. ex auot. oit.)

     

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1836.  

    Onosmodium bejariense DC.

    A. P. Decandolle (1846, v,. 10, p. 70) ... described O. bejariense from a manuscript by his uncle(?)

    Original Text Translation and Comments
    § 2. Flores minores. Filamenta gracilia glabra anthera multo breviora. Antherae glabrae, lobis divergentibus. Styli longius exserti.  
    … [Description of O. Virginianum omitted.] …  
    4. O. Bejariense (DC. mss.), caule pilis patentibus hispido , foliis oblongo-lanceolatis acutiusculis supra strigillosis subtus pilis brevissimis adpressis pubescentibus , nervis pinnatis utrinque 3 , lobis calycinis linearibus obtusius-culis pilis subpatentibus hispidis corolla duplo fere brevioribus , corolla adpresse villosa, tubo ad spicem dilatato , lobis lanceolatis acutis. ♃ in Mexico boreali prope Bejar ad Rio de la Trinidad (Berlandier pl. exs. n. 1681 ! ). Folia 2½ poll. longa 6-7 lin. lata. Bracteae longe ciliatae. Corolla 4 lin. longa. Nuces juniores rugulosse, adultse ovoideae laeves albae nitidae hinc inde punctatae. (v. s.)   There are two records of Berlandier no. 1681 in SEINet. One from GH described the collecting locality as “[San Antonio] de Bejar al Rio de la Trinidad [San Antonio to the Trinity River]” and therefore anywhere in between those two endpoints. Annotated, “Collected in May of 1828 at Trinity River crossing in eastern Texas. B. L. Turner, 1994.” The other, from F, appears to be an image of a voucher containing two specimens, one of which is Berlandier's no. 1681, from the herbarium of Moïse-Étienne Moricand, a Swiss botanist and malacologist (1779–1854).

    Literature Cited:
    - Gray, Asa, 1878-1895.  

    Gray (1878, v. 2(1), p. 206) accepted DeCandolle's O. bejariense ...

    Original Text
    O. Bejariense, DC. Stems 1 to 3 feet high, rather stout, hispid with spreading bristles : leaves oblong-lanceolate, 5-7-ribbed (the lower obtuse, upper acutish); upper surface appressed strigose-hispid, the lower more or less canescent with fine and soft pubescence : flowers short-pedicelled : corolla funnelform (6 to 9 lines long), about twice the length of the calyx, white; the greenish ovate-triangular acuminate lobes about one quarter the length of the tube, minutely pubescent externally and with some long hirsute hairs. — Prodr. x. 70. O. Carolinianum, Torr. Bot. Mex. Bound. l. c, not DC. — Border of thickets, nearly throughout Texas ; first coll. by Berlandier.

     

    Literature Cited:
    - Michaux, Andre, 1803.  

    Onosmodium molle Michx.

    Michaux (1803, v. 1, p. 132) ...

    Original Text
      ONOSMODIUM.
      Herbae foliis nervosis , nervis sublongitudinalibus.
      Cal. profunde 5-partitus : laciniis erectis , angusto-linearibus.
      Cor. oblongiuscule subcampanulata ; fauce nuda : limbo ventricoso , semi-5-fido ; laciniis erecto-conniventibus , subsemilanceolatis, acutis ; harum et incisurarum margine inflexo.
      Stam. antherae sessiles , inclusae , sagittatae , acutae.
      Pist. Onosmae ; stylo promisse exerto.
      Obs. Genus affine Onosmae : differt corolla multo breviore ; limbo profundius et acute diviso , connivente : antheris sessilibus nec ita mucronatis.
      [Describes O. hispidum with synonym of Lithopsermum virginianum L.]
    MOLLE.

    tab. 15

    O. molliter candicanti-villosum : foliis oblongo-ovalibus , subtriplinerviis , ferre tomentosis : corolla laciniis subovalibus.
      Hab. in Tennassee, circa Nashville.

     

    Other articles:
    • Golden Checklist Flora:  Phacelia heterophylla;  

    Phacelia hastata Dougl. ex Lehm.“Silverleaf Phacelia”

     

    Literature Cited:
    - Lehman, Johann G. C., 1828-1857.  

    Lehman (1830, v. 2. p. 21) describes P. hastata in great detail from Douglas' manuscript, though he gives no information about when and where Douglas saw the plant.

     

    Other articles:
    • Golden Checklist Flora:  Phacelia heterophylla;  

    Phacelia heterophylla Pursh “Varileaf Phacelia”

     

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 1, p. 140) ...

    Original Text
    heterophylla. 2. P. erecta, hispida ; foliis petiolatis pinnatifidis, laciniis lanceolatis integerrimiis nervosis : impari elongata : floralibus simplicibus lanceolatis petiolatis, racemiis bifidis densifloris, laciniis corollae integerrimis.
      On dry hills on the banks of the Kooskoosky. M. Lewis. ♂ June, July. v. s. in Herb. Lewis. Flowers pale blue.
      Caulis erectus, ramosus : rami elongati. Folia petiolata : inferiora pinnarifida : laciniis lateralibus lanceolatis, 1-2-jugis : lacinia terminali longo-lanceolata ; superiora et ramea simplicia, lanceolata. Petioli marginati. Racemi terminales pedunculati dichotomi, spirales, secundi. Pedicelli brevissimi. Calyx laciniis linearibus. Corolla subcampanulata, calyce duplo longior: laciniis oblongis, obtusis, integerrimis. Filamenta corollae duplo longiora. Sylus staminibus longior, bifidus.
     

    P. heterophylla and P. hastata clustered into a megellanica/humiles complex alon with P. imbricata, nemoralis, egena, californica, breweri, humilis, divaricata, and douglasii.

     

    Ackerfield (2015, p. 503) notes that P. hastata and P. heterophylla “… belong to a large polyploid complex and might be better treated as a single species [P. magellanica (Lam.) Cov.] with many infraspecific taxa.”

     

    Other articles:
    • Golden Checklist Flora:  Plagiobothrys scouleri hispidulus;  

    Plagiobothrys scouleri (Hook. & Arn.) I.M. Johnst. var. hispidulus (Greene) Dorn “Scouler's popcornflower”

    Found only in emphemeral ponds on North and South Table Mountains, Plagiobothrys scouleri (Hook. & Arn.) I.M. Johnst. var. hispidulus (Greene) Dorn “Scouler's popcornflower” is common along drying pond margins or in moist muddy soil in open meadows. The only other Jefferson County collection is at Ranson/Edwards Homestead Open Space by the author. Colorado collections are mostly west of the Front Range wherever suitable habitat is found.

    Literature Cited:
    - Greene, Edward L., 1887.  

    Greene (1887, v. 1, p. 17) ...

    Original Text
    10. A. HISPIDULA. Diffusely branching, 4 — 8 inches high, canescent with a short, setose-hispid pubescence ; racemes naked or leafy-bracted : calyx not accrescent: akenes ovate, opaque, ¾-line long, carinate on both sides, the back very lightly so and beset with a minute muriculation, the transverse rugulae few and not prominent ; scar almost basal, ovate-oblong.
    From the San Bernardino Mountains, Cal. (Parish, No. 1470) northward to Oregon (T. J. Howell), referred to “Eritrichium Californicum” from which plant it differs in its rough pubescence, and muriculate rather tlian granulate akenes.
     

    Plagiobothrys scouleri var. hispidulus (Greene) Dorn, Vasc. Pl. Wyoming 295 (1988): (1988).

     

    Other articles:
    • Golden Checklist Flora:  Phyla cuneifolia;  

    Phyla cuneifolia (Torr.) Greene “Wedgeleaf”

     

    Literature Cited:
    - Greene, Edward L., 1899b.  

    Greene (1899, v. 4, p. 47) ...

    P. cuneifolia. Zapania cuneifolia, Torr. Ann. N. Y. Lyc. ii. 234 (1826). Lippia cuneifolia, Steud. Nom. ii. 54 (1841), Torr. Marcy's Report 261, t. 17. A very strongly marked halophilous species, common on moist subsaline or alkaline plains of the Rocky Mountain region, ranging westward to California, southward to Arizona and perhaps Mexico, and with a near relative or two in South America.

     

    Other articles:
    • Golden Checklist Flora:  Verbena aristigera;  

    Verbena aristigera S. Moore “Moss Verbena”

     
     

    Verbena aristigera (syns. Glandularia aristigera and Verbena tenuisecta), variously called the moss verbena, desert verbena, fine leafed verbena, wild verbena, tuber vervain, South American mock vervain, Mayne's curse and Mayne's pest, is a species of flowering plant in the family Verbenaceae. It is native to Bolivia, southern Brazil, northern Argentina, Paraguay and Uruguay. It has been widely introduced to the rest of the world's drier tropics and subtropics, including California, Guatemala, Honduras, Nicaragua, Venezuela, Greece, Nigeria, eastern and southern Africa, India, and all of Australia except Tasmania.

     

    Other articles:
    • Golden Checklist Flora:  Verbena bracteata;  

    Verbena bracteata Lag. & Rodr. “Large Bract Vervain”

     

    Literature Cited:
    - Lagasca, Mariano, and Joseph Rodriguez, 1801.  

    Described by Lagasca & Rodriguez (1801) who say, “Se ignora su patria …[Homeland is unknown].”

    Literature Cited:
    - Perry, Lily M., 1933.  

     

    Literature Cited:
    - O'Leary, N., M. E. Mulgura, & O. Morrone, 2010.  

    O'Leary, et al. (2010, pp. 369-371) ...

    Distribucion y ecologia. Especie muy ampliamente distribuida en Estados Unidos de America, principalmente en los estados del sudoeste como Arizona, Nevada y Colorado; tambien en Mexico (Perry, 1933). Se la encuentra en campos abiertos y tambien en orillas de rios, formando grandes matas que se esparcen sobre la tierra alrededor de 1 m de distancia.

     

    Other articles:
    • Golden Checklist Flora:  Verbena hastata;  

    Verbena hastata L. “Swamp Verbena”

     

     

    Other articles:
    • Golden Checklist Flora:  Dracocephalum;  

    Dracocephalum parviflorum Nutt. “American Dragonhead. ”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Nuttall (1818) described D. parviflorum from material seen at Fort Mandan. I have not found any reference to his material, such as a type.

    Original Text
    5. * parviflorum. Flowers verticillate, subcapitate; leaves ovate-lanceolate, deeply serrate, and petiolate, bractes foliaceous, ovate, ciliate, and serrate, serratures conspicuously mucronate; upper segment of the calix much larger than the rest; flower scarcely longer than the calix. Hab. Around Fort Mandan, on the Missouri; on the borders of thickets. Obs. Biennial; nearly smooth, stem and petiole a little pubescent; flowers almost imbricated in a leafy capitulum, very small, and nearly white; calix awned, arid and membranaceous, semiquinquefid; bractes divaricately awned; upper lip of the corolla emarginate, arched; lower 3-lobed, central lobe subcrenate. Flowering time, July. Apparently allied to D. Moldavica, but the leaves are entirely destitute of punctures: scarcely a congener with D. virginianum, and closely allied to Melissa.

    Principally a Siberian genus.

     

    Other articles:
    • Golden Checklist Flora:  Lamium amplexicaule;  

    Lamium amplexicaule L. “Henbit”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 579) ...

    Original Text Translation and Comments
    3. LAMIUM foliis floralibus feffilibus amplexicaulibus obtufis. Hort. cliff. 314. Fl. fuec 495. Roy. lugdb. 319. Gron, virg. 66. Dalib. parif. 179. amplexicaule.  
    Lamium, folio caulem ambiente, minus. Bauh. pin. 231.    
    Morfus gallinae, folio hederulae, alternum. Lob. ic. 463.   “The bite of a hen is an alternative to the leaf of the ivy.”
    β Lamium, folio caulem ambiente, majus. Bauh. pin. 231.    
    Habitat in Europae cultis. ☉.    

     

    Other articles:
    • Golden Checklist Flora:  Lycopus americanus;  

    Lycopus americanus W. P. C. Barton “American Bugleweed”

     

    Literature Cited:
    - Muhlenberg, Henry, 1813.
    Full Size ImageMuhlenberg (1813) listing of Lycopus americanus.  

    Lycopus americanus Muhl., Cat. Pl. Amer. Sept. 3 (1813), nom. inval.

    Muhlenberg (1813) listed the name, but gave to way to distinguish from L. virginicus this rendering his name invalid.

    Literature Cited:
    - Barton, William P. C., 1815.
    Full Size ImageBarton (1815) publication of Lycopus americanus  

    Lycopus americanus Muhl. ex W.P.C.Barton, Fl. Philadelph. Prodr. 15 (1815).

    Barton (1815) again listed the name again, citing Muhlenberg (1813) this time with some characters for diagnosis, specifically “Very closely related to Mentha, but no odor; leaves sinuate-serrate, apendiculate.”

     

    Other articles:
    • Golden Checklist Flora:  Marrubium vulgare;  

    Marrubium vulgare L. “Horehound”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 583) ...

    Original Text
    5. MARRUBIUM dentibus calycinis fetaceis uncinatis. Hort. cliff. 342. Fl. fuec. 485. Mat. med. 305. Roy. lugdb. 315. Hall. helv. 648. Dalib. parif. 182. vulgare.
    Marrubium album vulgare. Bauh. pin. 230.  
    Marrubium vulgare. Cluf. hift. 2. p. 34.  
    Habitat in Europae borealioris ruderatis.  

     

    Other articles:
    • Golden Checklist Flora:  Mentha arvensis;  

    Mentha arvensis L. “Wild Mint”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 577) ...

    Original Text
    6. MENTHA floribus verticillatis, foliis ovatis acutis ferratis, ftaminibus corollam aequantibus. Hort. cliff. 307. Fl. fuec. 481. Roy. lugdb 326. Hall. helv. 658. Dalib. parif. 178. arvenfis.
    Mentha arvenfis verticillata procumbens. Morif. hift. 3. p. 369. f. 11. t. 7. f. 5.  
    Calamintha arvenfis verticillata.Bauh. pin. 229.  
    Habitat in Europae agris frequens poft meffem.  
    Caules diffufi; Verticilli laterales; Planta tota hirta; Stamina corollae limbo longitudine aequalia.  

     

    Other articles:
    • Golden Checklist Flora:  Mentha spicata;  

    Mentha spicata L. “Spearmint”

     

     

    Other articles:
    • Golden Checklist Flora:  Monarda fistulosa;  

    Monarda fistulosa L. “Wild Bergamot”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p. 22) ...

    Original Text
    fiftulofa. 1. MONARDA capitulis terminalibus, caule obtuf-angulo. Hort. upf. 12.
      Monarda floribus capitatis, caule obtufo. Vir. cliff. 3. Roy. lugdb. 313.
      Monarda. Hort. cliff. 11.
      Origanum fiftulofum canadenfe. Corn. canad. 13. t. 14.
      Habitat in Canada. ♃

     

    Other articles:
    • Golden Checklist Flora:  Monarda pectinata;  

    Monarda pectinata Nutt. “Plains Beebalm”

     
       

    Literature Cited:
    - Nuttall, Thomas, 1848b.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1848, publication details;  

    Nuttall (1848) described Monarda pectinata from a collection by William Gambel.

    Original Text
    MONARDA
    M. *pectinata. Biennial ? slightly pubescent ; leaves oblong-lanceolate, denticulate, shortly petiolate ; capituli proliferous, rather small, subtended by herbaceous, some of them purplish, ovate-acute, strongly ciliated, as well as the elongated, setaceous teeth of the calyx; corolla widely ringent, the tube scarcely exserted beyond the calyx.

    Hab. Near Santa Fe, New Mexico.

     

    Other articles:
    • Golden Checklist Flora:  Nepeta cataria;  

    Nepeta cataria L. “Catnip”

     

     

    Other articles:
    • Golden Checklist Flora:  Prunella vulgaris;  

    Prunella vulgaris L. “Common Selfheal”

     
     

    Linnaeus (1753, v. 2, p. 600) …

     

    Other articles:
    • Golden Checklist Flora:  Salvia pratensis;  

    Salvia pratensis L. “Meadow Clary”

    Linnaeus (1753, v. 1, p. 25) …

     

    Other articles:
    • Golden Checklist Flora:  Salvia reflexa;  

    Salvia reflexa Hornemann “Lanceleaf Sage”

    (Syn: S. lanceolata Rydb. [unpub.] , S. lanceolata Willd. [illeg.] , S. trichostemmoides Pursh )

     

    Other articles:
    • Golden Checklist Flora:  Scutellaria brittonii;  

    Scutellaria brittonii Porter “Britton's Skullcap”

     
      Isotype: NY415671, T. C. Porter, s.n., Colorado, Clear Creek Canyon, 9000 ft, June 15, 1873. The 9000 ft elevation in Clear Creek Canyon is just below Silver Plume.

    Literature Cited:
    - Porter, Thomas C., 1894.  

    Scutellaria resinosa Torr., Porter (1894, v. 21, p. 177).

    Original Text
    Scutellaria resinosa Torr. — If the description of the species given under this name in Dr. Gray's Synoptical Flora (p. 381) be compared with the references there cited, it will be found that they do not agree. The true S. resinosa of Torrey, collected during Long's First Expedition and published in the Annals of the New York Lyceum (2 ; 232) and by Bentham, in D.C. Prodr. (12 : 427), is beyond question nothing else than S. Wrightii K. Gray, published long after (Feb. 1872) in the Proceedings of the American Academy. This is clear both from the descriptions and the type specimen in the Torrey Herbarium, It differs from the Colorado plant in its strong woody root, branching stems, round leaves, smaller flowers and shorter corolla-tubes. Its range extends from Kansas to Texas and thence westward to the Huachuca Mountains of Arizona. The other species is most common in the Rocky Mountains of Colorado and being thus deprived of the name resinosa I bestow on it that of Scutellaria Brittonii
    Thos. C. Porter.

     

    Other articles:
    • Golden Checklist Flora:  Datura wrightii;  

    Datura wrightii Regel “Sacred Datura”

     

    Literature Cited:
    - Regel, Eduard, 1850.  

    Regel (1859, v. 8, pp. 193-194) ...

    Diese neue Datura ward aus Californien durch das Haus Vilmorin Andrieux et Comp, in Cultur gebracht und unter dem Namen D. meteloides D. C. verbreitet. Herr Ortgies, der die beistehende Tafel nach einer Pflanze anfertigen Hess, die im Botanischen Garten zu Zürich zur Blüthe kam, machte uns darauf aufmerksam, dass die von Vilmorin verbreitete Pflanze mit der Beschreibung , wie solche sich im 13. Bande von DeCandolle's Prodromus findet, nicht übereinstimmt. This new Datura was from California by the House of Vilmorin-Andrieux & Co., brought into culture and under known as D. meteloides D.C. Mr. Ortgies, who made a plate based on a plant Hess [grew] in the botanical garden [and] came to bloom in Zurich, made us [aware] that the common plant described by Vilmorin does not match that shown in the 13th volume of DeCandolle's Prodromus.

    Literature Cited:
    - Barkley, Arthus S., 1959.  

    Arthur S. Barclay. 1959. New Considerations in an Old Genus: Datura. Botanical Museum Leaflets, Harvard University. v. 18, No. 6, pp. 245- January 16, 1959. https://www.biodiversitylibrary.org/part/295184

    Barclay (1959, v. 18, n. 6., pp. 245-) ....

    In 1855, a new Datura was brought into cultivation by the French horticulturist M. Louis Vilmorin from seeds sent him bv Asa Gray who had obtained them from a collection made by Charles Wright (no. 526) in western Texas in 1849. This plant was cultivated and distributed under the name of Datura meteloides (Ewan, 1944). M. Ortgies, an employee of the Vilmorin Co. of France, noticed the discrepancy between de Candolle's description of D. meteloides and the plant cultivated under the same name and brought it to the attention of the German botanist, Dr. Eduard von Regel. As a result, a new species, D. Wrightii, was published by Regel (1859).
    Gray (1878) considered the southwestern perennial species of the United States to represent Datura meteloides and relegated D. Wrightii to synonymy handling the obvious lack of agreement between the two descriptions with the following comment : “ ... Dunal in DC. Prodr. l.c. 544 (the descr. and drawing of Mocino and Sesse wrong as to 10-dentate corolla).” This erroneous interpretation was adopted by Safford (1921) who apparently followed Gray.

     

    Other articles:
    • Golden Checklist Flora:  Lycium chinense;
    Full Size ImageColl. No. 2486, Lycium chinense
    Full Size ImageFlower of Coll. No. 2486, Lycium chinense  

    Lycium chinense Mill. “Chinese Desert-Thorn”

     

    Literature Cited:
    - Miller, Philip, 1768.  

    5. LYCIUM (Chinenfe) foliis ovato-lanceolatis, ramis difufis, floribus folitariis patentibus alaribus, ftylo longiori. Boxthorn with oval fpear-fhaped leaves, diffufed branches, and fingle fpreading flowers proceeding from the fides of the branches, with a longer ftyle.
    ....
    The fifth fort grows naturally in China, from whence the feeds were brought to England a few years paft, and the plants were raifed in feveral gardens, and by fome were thought to be the Thea. This rifes with weak, irregular, dilffufed branches to a great height, but require fupport, otherwife they will trail upon the ground : I have meafured fome of thefe branches, which in one year has been upward of twelve feet long : the lower leaves are more than four inches long, and three broad in the middle : they are of a light green and a thin confiftence, placed without order on every fide the branches ; as the fhoots advance in length, fo the fize of the leaves diminifh, and toward the upper part they are not more than an inch long, and a quarter of an inch broad ; they fit clofe to the ftalks on every fide. The flowers come out fingly at every joint toward the upper part of the branches, ftanding upon fhort flender foot-ftalks ; they are of a pale colour, with fhot tubes ; the brims are fpread open broader than either of the former forts, and the ftyle is confiderably longer than the tube of the flower. This fort flowers in Auguft, September, and October; the plant is very hardy, and retains its leaves till November before they dcay. It propagates faft enough by its creeping roots, which fend out fuckers at a great diftance, and the cuttings thruft into the ground will take root as freely as Willows.

     

    Other articles:
    • Golden Checklist Flora:  Physalis hederifolia var. comata;  

    Physalis hederifolia A. Gray var. comata (Rydb.) Waterfall “Ivy-Leaved Ground Cherry”

     

    Literature Cited:
    - Rydberg, Per Axel, 1895.  

    Rydberg (1895, v. 22, pp. 306-307) ...

    Physalis comata n. sp.
    Perennial, erect, about 0.5 metre high; pubescence fine and short, that on the calyx, pedicels and upper branches mixed with long white flat jointed hairs ; like P. heterophylla Nees (P. Virginiana Gray, not L.), but leaves smaller, blade not over 0.5 decimetre long, rounded, ovate, scarcely at all cordate at the base, about 5 centimetres long, thin, somewhat repand-dentate or nearly entire; petioles as long as the leaves; corolla greenish-yellow with brown centre; fruiting calyx as in P. Hederaefolia Gray, but of thinner texture.
    The validity of this as a species may be doubted. It is intermediate between P. Heterophylla and P. Hederaefolia and might be placed as a variety of either with about equal right. It, perhaps, most resembles the latter, but differs in the thinner texture of the leaves and the fruiting calyx, in the larger flower and in the long white hairs.
    The following localities are recorded :
    Nebraska: P. A. Rydberg, No. 269, 1891 (type).
    Kansas: E. Bartholomew, No. 2, 1892; E. A, Popenoe, 1875.
    Texes(sic) : G. Jermy, No. 618, 1890.

    Literature Cited:
    - Waterfall, Umaldy T., 1950.  

    Waterfall (1950, v. 52, n. 619, p. 171) ...

    Physalis hederaefolia Gray, var. comata (Rydb.), stat. nov. (P. comata Rydb. Bull. Torr. Bot. Cl. 22: 306. 1895). In the western part of the Oklahoma panhandle we have a Physalis that looks like P. hederaefolia, but has long (1-2 mm.) flat jointed hairs scattered on the stem, leaves and calyces among the abundant viscid capitate hairs. These plants the author is referring to var. comata. Typical P. hederaefolia may have a villous pubescence, of somewhat jointed hairs (up to half as long as in var. comata) mixed with the short capitate glandular hairs, but the pubescence seems to be denser and shorter. Referred to var. comata are: Waterfall 7867, 14 miles east of Hooker, Texas County, June 11, 1948; Waterfall 7445, plains north of the Black Mesa, north of Kenton, Cimarron County, July 9, 1947; Waterfall 9240, northeast slopes of the Black Mesa, 3 miles north and 1 east of Kenton, Cimarron County, August 10, 1949.

     

    Other articles:
    • Golden Checklist Flora:  Physalis hispida;  

    Physalis hispida (Waterf.) Cronquist “Prairie Ground Cherry”

     

     

    Other articles:
    • Golden Checklist Flora:  Physalis longifolia;  

    Physalis longifolia Nutt. “Longleaf Groundcherry”

     

    Literature Cited:
    - Nuttall, Thomas, 1834b.  

    Original Text
    3. *longifolia. Glaberrima, caule angulata erecto, foliis soltariis ovato-lanceolatis acuminatis sinuato-dentatis longe pedunculatis, floribus solitariis pendulis. — Obs. Herbaceous ; root perennial ; stem angular, about eighteen inches high, and branching above ; leaves smooth, four to five inches long, irregularly, sparingly and sinuously toothed ; flowers, as usual, yellowish, with five brown blotches towards the base; calix muxh larger than the berry. It bears much the aspect of Capiscum annuum, and, from the diagnosis, appears allied to P. chenopodifolia. — Hab. On the sandy banks of the Arkansas, near Belle Point. Flowering in June.

     

    Other articles:
    • Golden Checklist Flora:  Physalis virginiana;  

    Physalis virginiana Mill. “Virginia Groundcherry”

     

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015) comments on determinations of P. virginiana and P. longifolia.

    Original Text
    Most specimens originally identified as P. virginiana actually belong to P. longifolia. Physalis virginiana appears to be restricted to the base of the northern Front Range in Colorado.

     

    Other articles:
    • Golden Checklist Flora:  Solanum dulcamara;  

    Solanum dulcamara L. “Climbing Nightshade”

     

     

    Other articles:
    • Golden Checklist Flora:  Solanum physalifolium;  

    Solanum physalifolium Rusby “Hoe Nightshade”

     

     

    Other articles:
    • Golden Checklist Flora:  Solanum rostratum;  

    Solanum rostratum Dunal “Buffalo Bur Nightshade”

     

     

    Other articles:
    • Golden Checklist Flora:  Solanum triflorum;  

    Solanum triflorum Nutt. “Cutleaf Nightshade”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Nuttall (1818) described the Solanum of North America and included a review of Solanum and related taxa, their origins and uses.

    Original Text Comments
    6. *triflorum. Stem unarmed, herbaceous and procumbent; leaves dentately-pinnatifid, smooth, segments acute, somewhat undulated, with the margin more or less revolute; peduncles opposite the leaves, 2 or 3-flowered. — Flowers small and white, revolute; fruit about the size of a cherry, green when ripe. Stem a little hirsute, spreading and procumbent, about a foot long; leaves somewhat runcinate. This species, though very distinct, appears to have some affinity with the S. runcinatum of Peru and Chili (sic). — Hab. As a weed in and about the gardens of the Mandans and Minitarees, and in no other situations. Near Fort Mandan. Flowering from June to August. This is followed by a several paragraph review of Solanum and related species, their origins and uses.

     

    Other articles:
    • Golden Checklist Flora:  Limosella aquatica;  

    Limosella aquatica L. “Water Mudwort”

     

     

    Other articles:
    • Golden Checklist Flora:  Scrophularia lanceolata;  

    Scrophularia lanceolata Pursh “Lanceleaf Figwort”

     

     

    Other articles:
    • Golden Checklist Flora:  Verbascum blattaria;  

    Verbascum blattaria L. “Moth Mullein”

     

     

    Other articles:
    • Golden Checklist Flora:  Verbascum phlomoides;  

    Verbascum phlomoides L. “Orange Mullein”

     

     

    Other articles:
    • Golden Checklist Flora:  Castilleja integra;
    • Field Notes:  Coll. No. 1143, 15 Jun 2015;   Coll. No. 1379, 29 May 2016;   Coll. No. 1614, 15 May 2017;  Coll. No. 1865, 23 May 2018;  

    Castilleja integra A. Gray “Wholeleaf Indian Paintbrush”

    A very common paintbrush in Golden s.l., Castilleja integra A. Gray “Wholeleaf Indian Paintbrush” is found in all the open spaces. Jefferson County collections are mainly in the northern part of the county, on the plains and up in the foothills. Most of the Colorado state collections are in the Rocky Mountains and adjacent plains from Denver south.

    The name was applied by A. Gray (1859) in “Pacific Railroad Reports” while citing a number of contemporary collections in the southwest.

     

    In 1849, he joined an army expedition (with Gray's help) through Texas, botanising from Galveston to San Antonio and then on to El Paso. But he had to walk most of the 673 miles, (which took over 104 days effort). He collected seeds of Penstemon baccharifolius (Hook.), between Texas and El Paso, which were later given to William Hooker. Also, Castilleja lanata (found near the Rio Grande) and Castilleja integra (found in the Organ Mountains, near El Paso) (Pennell, 1935). In the spring of 1851, he joined the United States and Mexican Boundary Survey (also with Gray's help). (Wikipedia)

    Literature Cited:
    - Gray, Asa, 1849.

    Other articles:
    • Golden Checklist Flora:  Gray, 1849, publication details;  

    Castilleja integra is not listed in Gray (1849) Plantae Fendlerianae Novi-Mexicanae. Although, a good part of the report is "… to be continued."

    Literature Cited:
    - Gray, Asa, 1852-1853.  

    Gray (1952) does not mention any Wright collections in the Scrophulariaceae

    Literature Cited:
    - Torrey John, 1859.  

    Emory, William H., 1859. Report on the United States and Mexican Boundary Survey. Volume II. Torrey, John, 1859. Part I. Botany of the Boundary Washington, 1859. p. 119

    SCROPHULARIACEAE (by A. Gray)

    Castilleja integra (sp. nov.): perennis; caule stricto tomentoso; foliis linearibus integerrimis subtus tomentulosis, floralibus oblongis obocatisque integerrimis coloratis (paniceis);; spica conferta; calyce aequaliter vel postice profundius bifido, lobis bifidis lanceolatis obtusiusculis labium inferius galea multoties brevius adaequantibus. — Organ mountains, east of El Paso; Wright, (undistributed,) Bigelow. Guadaloupe cañon, Sonora; Capt. E. K. Smith. Also gathered in the Rocky Mountains further north by Dr. Kreuzfeldt, in Gunnison's expedition. Stem one or two feet high, mostly simple, rigid; leaves 1½ to 3 inches long, 2 to 3 lines wide, entire; most of the floral ones almost wholly petaloid, ample, shorted than the fully developed flowers. Calyx 8 or 12 lines long, red or reddish; “corolla reddish green;” glaea 6 to 8 lines long; the lower lip very short. Apparently a well marked new species of the section Callichroma. It is No. 584 of Fendler's New Mexican collection; and Dr. Bigelow gathered specimens in Whipple's expedition on the Llano Estacado.

     

    Pennell, Francis W. [Curator of Botany, Academy of Natural Sciences of Philadelphia] The Scrophulariaceae of Eastern Temperate North America The Academy of Natural Sciences Monogtaphs, Number 1 Philadelphia, 1935. p. 533

    9. Castilleja integra Gray

    Castilleja integra A. Gray, in Torrey, Bot. Mex. Bound. 119. 1859. “Organ mountains, east of El Paso; Wright ..., Bigelow. Guadaloupe canon, Conora; Capt. E. K. Smith."

    Bracts red. Flowering from May to August.

    Gravelly soil, plains and hills, Colorado to Texas, Chihuahua and Arizona. Known in our territory from a single record along the Rio grande.

    Texas. Valverde: bluffs of Devils R., Havard (U).

     

    Other articles:
    • Golden Checklist Flora:  Orobanche fasciculata;  

    Orobanche fasciculata Nutt. “Clustered Broomrape”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Described by Nuttall (1818, v. 2, p. 58-59) in a new section named Gymnocaulis as Oronanche fasciculata, with calyx and corolla almost equally 5-cleft.

    Original Text Comments
    448. OROBANCHE. L. (Broomrape.)  
    Calix 4 or 5-cleft, segments often unequal. Corolla ringent. Capsule ovate, acute, 1-celled, 2-valved; seeds numerous. A gland beneath the base of the germ.  
    [Description of the genus omitted.]
    [Description of O. americana. omitted.]
    * Gymnocaulis. Calix and corolla almost equally 5-cleft. Assumed to be a new section name.
    3. * fasciculata. Stem short and simple; peduncles many, naked, nearly terminal, and about the length of the stem; lobes of the corolla very short, rounded, and naked on the margin.  
    Hab. With the above. Flowering in June and July. Very nearly allied to the following. The above is O. ludoviciana, which was “… in sandy alluvial soils, around Fort Mandan …” The following is O. biflora. Handwritten note, “primarily Artemisia.”
    Obs. 4 or 5 inches high, densely and pulverulently pubescent. Stem angular, with few scales, terminating in 6 or more naked filiform peduncles, 2 to 2 1-2 inches in length. Calix campanulate, semiquinquefid, segments sublanceolate, acute, little more than one-third the length of the corolla. Corolla curved, tubular, dilute purple; segments of the border scarcely more than a line long; lower lip producing a grooved palate. Stamina smooth, lobes of the anthers acute below, opening marginally and longitudinally, the lobes being parallel. Capsule 1-celled, ovate, acuminate.  

     

    Other articles:
    • Golden Checklist Flora:  Orobanche uniflora;  

    Orobanche uniflora L. “Naked Broomrape”

    (Syn: Aphyllon uniflorum (L.) Torr. & A. Gray , Orobanche uniflora L. var. uniflora )

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 633) ...

    Original Text
    5. OROBANCHE caule unifloro. uniflora.
    Orobanche aut Helleborine affinis marilandica, caule nudo, unico in fummitate flore. Raj. fuppl. 595.  
    Dentariae f. Anblato cordi affinis, flore pallide caeruleo. Gron. virg. 70.  
    Gentiana minor aureo, flore fimplici amplo deflexo pallide flavefcente. Pluk. mant. 89. t. 348 f. 3.  
    Aphyllon. Mitch. 25.  
    Habitat in Virginia.  

     

    Other articles:
    • Golden Checklist Flora:  Orthocarpus luteus;  

    Orthocarpus luteus Nutt. “Yellow Owls Clover”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Nuttall (1818, v. 2, p. 57) described a new genus Orthocarpus and added O. luteus to it.

    Original Text Comments
    446. *ORTHOCARPUS. †  
    Calix tubular, semiquadrifid. Corolla bilabiate, closed; upper lip smaller, compressed, with the margin inflected; lower concave, obsoletely 3-toothed, unexpanded. Anthers unconnected; lobes unequal, divaricate. Capsule straight, elliptic-ovate, 2-celled, 2-valved, many-seeded, opening on both sides; dissepiment transverse. Seeds small, alated lunate margin.  
    Annual; stem simple; leaves alternate, entire; flowers axillary, alternate, sessile, and bracteate; bractes divaricately trifid.  
    Species. 1. O. luteus.  
    Descript. Root tortuous, perpendicular, and fibrous. Stem simple, hirsutely pilose, terete. Leaves increasing in size upwards, alternate, sessile, lanceolate-linear, acute, entire, opaque, and as will as the bractes and calix shortly and somewhat viscidly pubescent, margins scabrous; bractes cuneate, divaricately trifid, about an inch long and the same in width, 3-nerved. Calix much shorted than the bractes, partly compressed, striated, subcampanulate, segments linear-lanceolate, acute. Corolla yellow, smooth; tube slender, about the length of the calix; both of the lips concave, inflected, and closed, the lower somewhat plaited and terminated by 3 minute dentures, so inconspicuous as the give the corolla the appearance of being destitute of an under lip. Stamina 4, small; filaments capillary, ingrafted upon the upper lip a little below the orifice, approximating by pairs unde the same lip; anthers pale, unusually small, 2-celled, distinctly and separately 2-lobed, lobes pubescent, not parallel, one acute-angularly diverging from below the summit of the other, almost exactly similar to the small lamdba (sic) of the Greek alphabet (λ). Style filiform, stigma simple, minute. Capsule elliptic ovate, obtuse, staight (sic) and pubescent, included within the calix, 2-celled, 2-valved, many-seeded, margins of the valves partly inflected; dissepiment transverse, or origining from the middle of the valves and seminiferous. Seeds small, more than 10, having an alated interrupted margin. Cotyledones 2, very small, upon the growing plant, oval. Hab. In humid situations on the plains of the Missouri, near Fort Mandan; very local. Flowering in July and August. Height about 12 or 14 inches. Growing in quantities. Flowers of a bright and uniform yellow, almost of the size and form of the common species of Melampyrum at first sight. In point of affinity it cannot be compared with any other genus, notwithstanding its marked distinction.  

    † The straightness of the fruit, readily distinguishing this genus from Melampyrum.
    Melampyrum is a genus of about 20 species of herbaceous flowering plants in the family Orobanchaceae known commonly as cow wheat. They are native to temperate regions of the Northern Hemisphere, though not found in Colorado. They are hemiparasites on other plants, obtaining water and nutrients from host plants, though they are able to survive on their own without parasitising other plants.

     

    Other articles:
    • Golden Checklist Flora:  Erythranthe floribunda;  

    Erythranthe floribunda (Douglas ex Lindl.) G. L. Nesom “Many Flowered Monkey Flower”

    (Syn: Mimulus floribundus Lindl. )

    Literature Cited:
    - Lindley, John, 1827.  

    Lindley (1827, v. 13, t. 1125) ...

    A neat hardy annual, found by Mr. Douglas on moist rocks in the interior of the districts of the river Columbia.
       

     

    Other articles:
    • Golden Checklist Flora:  Erythranthe glabrata;  

    Erythranthe glabrata (Kunth) G. L. Nesom “Roundleaf Monkeyflower”

    (Syn: Mimulus glabratus Kunth)

     

    Other articles:
    • Golden Checklist Flora:  Collinsia;  

    Collinsia Nutt.

     

    Literature Cited:
    - Nuttall, Thomas, 1817.  

    Nuttall (1817, v. 1, pt. 1, p. 189) ...

    Description of Collinsia, a new genus of plants.
    By Thomas Nuttall.
    In the spring of 1810, during the course of an extensive journey into the north-western interior of the territories of the United States, I first became acquainted with the very singular and interesting plant which forms the subject of the present memoir. The specimens which I then obtained on the alluvial soils of the Alleghany and on the borders of lake Erie were finally lost. On arriving at St. Louis, near the confluence of the Missouri and Mississippi rivers, I found that Mr. John Bradbury, a botanist, had also detected this plant about the same time, on the banks of the Missouri and Mississippi, but I saw no specimen. In the spring of last year (1816), having undertaken a tour to the western states, I determined, if possible, again to collect this neglected plant, but after a journey of more than a hundred miles, for scarcely any other purpose, I arrived at Pittsburgh disappointed of my object. On descending the Ohio, however, nearly to Galiopolis, I at last recognised it on the more open alluvions of the river, withered and nearly past affording seed, accompanied by the Hesperis pinnatifida and the interesting Phalangium esculentum: from these seeds I have been at last fortunate enough to obtain the plant from which the accompanying drawing was taken by my friend, Mr. C. A. Le Sueur. This plant has for several years been known to Dr. Drake of Cincinati, in whose collection I saw specimens of it, and also with Dr. Short of Lexington, Kentucky, two gentlemen, who, amidst the avocations of an irksome profession, have still found leisure to cultivate some of the branches of natural science.
    I have dedicated this genus to the name of Zaccheus Collins, Esq. of Philadelphia, a gentleman, whose talents as a botanist and a mineralogist are deservedly acknowledged.

    The article goes on to a formal description of Collinsia verna and includes a black and white drawing.

     

    Other articles:
    • Golden Checklist Flora:  Collinsia parviflora;  

    Collinsia parviflora Lindl. “Maiden Blue-eyed Mary”

     

    Literature Cited:
    - Lindley, John, 1827.  

    “Received by the Horticultural Society from Mr. David Douglas, in 1827, by whome it was found in the vicinity of the river Columbia (Lindley, 1827).”

     

    Other articles:
    • Golden Checklist Flora:  Gratiola neglecta;  

    Gratiola neglecta Torr. “Clammy Hedge Hyssop”

     

    Literature Cited:
    - Torrey, John, 1819.  

    Torrey (1819, p. 10 & 89) ...

    Original Text
    9. Gratiola. L. Hedge Hyssop.
    1 aurea. Muhl. (Golden-Pert.) In sandy woods in moist places. New-Jersey. Long-Island, &c. ♃. Aug. yellow.
    2 Virginica. L.
    officinalis. Mx.
    B. Caroliniensis. Pers.
    In inundated and moist places (rare.) ♃. Aug.
    3. neglecta. Sp. nov. In similar situations. (common.) July-Aug. ♃. white and yellow.
    Page 10. Gratiola neglecta, nobis : G. caule pubescente assurgente tereti ; foliis glabris lanceolatis pauci-dentato-serratis basi attenuatis connatisque : calycis foliolis aequalibus ; filamenta sterilia nulla.

    Root fibrous, perenial. Stem about six inches, branched at the base, round ; leaves broad-lanceolate, narrowed at the base and a little connate, smooth, rather obtuse. Peduncles varying in length, some of them shorter, and some longer than the leaves, the upper ones opposite. Calyx with obtuse lanceolate segments a little more than one third the length of the corolla ; bracts longer than the calyx, beset with glanduliferous hairs. Corolla with divisions rounded, the upper segment longer, and all of them emarginate ; limb white ; tube yellow, bent obliquely, smooth ; throat hairy. Filaments shorter than the tube. Stigma oblique, compressed. Capsule ovate, acutish, and about the length of the calyx.

    This species approaches very near to G. Virginica, and may be the plant which Pursh has described as such. From the G. Virginica of Elliott it differs in several circumstances. Particularly in having a pubescent stem, and in the absence of abortive stamina. It has great affinity to Gratiola sphaerocarpa. Ell.

       

    Linaria canadensis (L.) Dum.-Cours. var. texana (Scheele) Pennell “Blue Toadflax, Texas Toadflax”

     

    Literature Cited:
    - Pennell, Francis W., 1921.

    Other articles:
    • Golden Checklist Flora:  Linaria canadensis texana;  

    Pennell (1921, p. 502) ...

    Original Text
    1a. Linaria canadensis texana (Scheele) Pennell, comb. nov.
    Linaria texana Scheele, in Linnaea 21: 4761. 1848. “Zwischen Houston and Austin [Texas] haufig:” Romer." Description sufficiently distinctive.
    Here distinguished primarily by roughness of seeds, which however varies from a few roughenings on the side to densely tuberculate. The wealth of Texan material at my disposal shows that each character of the contrasts previously outlined (in Contrib. U. S. Nat. Herb. 20: 323. 1920, and Proc. Acad. Nat. Sci. Phila. 71 : 258. 1920) between this and Linaria canadensis breaks down in numerous instances through the area where the two overlap. Consequently it seems more rational to consider texana a variety, its range and relation to the species roughly paralleling that of Veronica peregrina xalapensis (H. B. K.) Pennell. For an excellent illustration of Linaria candensis texana see the Botanical Magazine 63: pi. 3473. 1836.
    Open sand or sandy loam, frequently or usually a weed, common through the Coastal Plain, chiefly westward; occasional and probably introduced inland. Ranges widely through western North America, and into South America.
    List of specimens examined from Arkansas, Oklahoma, Louisiana, and Texas omitted.

     

    Other articles:
    • Golden Checklist Flora:  Linaria dalmatica;  

    Linaria dalmatica (L.) Mill. “Dalmatian ToadFlax”

    (Syn: Linaria dalmatica (L.) Mill. ssp. dalmatica, Linaria genistifolia (L.) Mill. ssp. dalmatica (L.) Maire & Petitm.)

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 616) ...

    Original Text
    dalmaticum. 18. ANTIRRHINUM foliis lanceolatis alternis, caule fruticofo.
      Linaria latifolia dalmatica, magno flore. Bauh. pin. 212. prodr. 106.
      Linaria maxima, folio lauri. Bauh. hift. 3. p. 458.
      Habitat in Creta. ♄.

     

    Other articles:
    • Golden Checklist Flora:  Penstemon secundiflorus;  

    Penstemon secundiflorus Benth. “Sidebells Penstemon”

    Found in every Golden s.l. open space. Jefferson County collections along the base of the Front Range and barely into the foothills, with just a few in the interior of southern county. Most collections along the Front Range and interior valleys.

    Name applied by Bentham (1846) to a specimen seen in Torrey's herbarium, possibly a Fremont collection.

    Literature Cited:
    - Bentham, George, 1846.  

    Bentham in DeCandolle (1846, v. 10, p.325) ...

    25. P. secundiflorus, erectus, foliis radicalibus petiolatis ellipticis lanceolatisve acutis, superioribus anguste lanceolatis, panicula angusta, pedunculis erectis plurifloris secundis, calycis segmentis ovali-oblongis acutis margine membranaceis, corollae tubo superne ampliato, antheris glabris, filamento sterili apice dilato barbato. ♃ in montibus scopulosis (Fremont !). Hinc P. nitido, illinc P. coeruleo affines. Caules 9-10-pollicares. Cymae laxiusculae, superiores 1-3-florae, inferiores sub-5-florae. (v. in herb. Torrey).

     

    Other articles:
    • Golden Checklist Flora:  Penstemon strictus;  

    Penstemon strictus Benth. “Rocky Mountain Penstemon”

     

    Literature Cited:
    - Bentham, George, 1846.  

    Bentham in DeCandolle (1846, v. 10, p. 324) ...

    Original Text Translation and Interpretation
    24. P. strictus, rigidus, subglaucus, foliis imis oblongo-spathulatis petiolatis, caulinis lineari-lancelatis, panicula elongata angusta virgata , pedunculis brevibus plurifloris , calycis segmentis ovali-oblongis obtusis margine membranaceis, antheris parce piloso-lanatis , filamento sterili superne dilatato subnudo. ♃ in montibus Scopulosis ad fontes fl. Sweetwater (Fremont !). P. micranthus Torr. in Frem. rep. p. 89, non Nutt. Specimen vidi unicum sesquipedale. Folia quam in P. coeruloa angustiosa. Corollae imperfectae iis P. coerulei subsimiles videntur. Antherarum pili pauci, longi, molles (v. in herb. Torrey.) 24. P. tight, rigid, subglaucous, the lower leaves oblong-spathulate, petiolate, of a linear-lanceolate cauline, elongated panicle, narrow striped peduncles calyx with short, numerous, oval-oblong segments, obtuse with membranous margin, anthers sparsely hairy, woolly, with a sterile filament enlarged above, I subdo. [forb] In the Rocky Mountains at the Sweetwater River Springs (Fremont!). P. micranthus Torr. in Frem. rep. p. 89, non Nutt. I saw only one foot and a half specimen. Leaves as in P. coeruloa angustiosa. The imperfect corolla appear to be similar to those of P. coerulei Anther hairs few, long, soft (seen in Torrey herbarium).

     

    Other articles:
    • Golden Checklist Flora:  Penstemon virens;  

    Penstemon virens Pennell ex Rydb. “Front Range Beardtongue”

    The name P. virens was applied by Rydberg (1917) apparently at the suggestion of Pennell who collected the type. The type locality is the general area of Red Rocks Amphitheater though the amphitheater did not exist at the time. As an entity it was known well before then, as early as Nuttall's 1834 trip to Oregon Territory, and by Parry's collection, and Hall & Harbour collections. However, it was treated as a form of P. pubescens until treated as P. humilis Gray (1862), of which part was segregated into P. virens Rydberg (1917) based upon Pennell's type.

    Literature Cited:
    - Gray, Asa, 1862.  

    Gray (1862, p. 69-70) ...

    43. P. humilis, Nutt. in herb. Acad. Philad. P. gracili Nutt. (P. pubescenti var. gracili supra) maxime affinis, sed vulgo humilior (3-9-pollicaris), foliis pallidioribus, thyrso etiam strictiore, corolla saturate caerulea breviore (haud ultra-semipollicari) fauce satis ampliata hiante plicis sub labio inferiore obsoletis. — Rocky Mountains, Nuttall (a very depauperate doubtless alpine specimen in herb. Acad. Philad.) ; common about Pike's Peak and vicinity, Parry, no. 257 (Enum. p. 27), Mr. Howard, and in taller specimens than before seen, Hall and Harbour, 1862, no. 387, on low mountains, “an early and very pretty species,” confirming its close relationship with P. gracilis. Yet this cannot be regarded as a variety of P. pubescens.

    Literature Cited:
    - Rydberg, Per Axel, 1917.  

    Rydberg (1917, p. 773 and 1066) ...

    60. P. virens Pennell. Stem 1-3 dm. high, glabrous up to the inflorescence; leaves glabrous, glaucescent, the lower petioled; blades 2—4 cm. long; upper stem leaves lanceolate or oblong, usually denticulate; calyx 4 mm. long; corolla deep blue, 15 mm. long, funnelform; lower lip slightly bearded; sterile stamen yellow bearded on the spatulate tip. P. humilis A. Gray, mainly, not Nutt. Plains and hills: Alta. — Colo. — Nev. Plain — Mont. Je-Au.
    ...
    New Genera and Species
    Pentstemon virens Pennell.
    Type: Stony hillsides, foothills north of Morrison, Colo., 1915, Pennell 5821 (N.Y.).
    P. humile A. Gray, Am. Jour. Sci. II, 34: 253. 1862. Not Nutt.

     

    Other articles:
    • Golden Checklist Flora:  Penstemon virgatus var. asa-grayi;  

    Penstemon virgatus A. Gray var. asa-grayi (Crosswh.) Dorn “Upright Blue Beardtongue”

     

    Literature Cited:
    - Crosswhite, Frank S., 1965.

    Locations: Empire.  

    Crosswhite (1965, p. 318) ...

    ...
    I have recently concluded that the plant called “P. unilateralis” by authors is nothing more than a well-marked subspecies of P. virgatus A. Gray (in N+bot. Mex. Bound. surv. 113. 1859; isotype seen). Dr. William T. Penland has provided the only description of the subspecies in H. D. Harrington's Manual of the Plants of Colorado (Sage Books, Denver, 1954). In recognition of the excellence of Gray's revisions of the genus, this previously nameless Penstemon is here designated Penstemon virgatus ssp. Asa-Grayii Crosswhite, ssp. nov. — foliis radicalis oblanceolatis; a ssp. virgatus caulibus glabrus differt: typus: H. N. Patterson 258 (PH, isotypes NY, US, WIS) Empire, Clear Creek Co., Colo., Jul. 13, 1892. Mounted with this flowering material, in all 4 herbaria cited, is fruiting material collected in Sept. of the same year. &Mdash; Frank S. Crosswhite, Herbarium, Universoty of Wisconsin, Madison, 53706.

     

    Other articles:
    • Golden Checklist Flora:  Plantago major;  

    Plantago major L. “Common Plantain”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1., p. 112-113) ...

    Original Text
    major. 1. PLANTAGO foliis ovatis glabris, nudo fcapo tereti, fpica flofculis imbricatis.
      Plantago foliis ovatis glabris. Hort. cliff. 43. Fl. fuec. 122. Mat. med. 49. Roy. lugdb. 401. Hall. helv. 470. Dalib. parif. 50.
      Plantago fcapo fpicato, foliis ovatis. Fl. lapp. 62.
      Plantago latifolia finuata. Bauh. pin. 189.
      Plantago major. Cam. epit. 261.
      β. Plantago latifolia glabra minor. Bauh. pin. 189.
      γ. Plantago latifolia rofea, floribus quafi in fpica difpofitis. Bauh. pin. 189. Hort. upf. 29.
      δ. Plantago latifolia rofea, flore expanfo. Bauh. pin. 189.
      ε. Plantago latifolia, fpica multiplici fparfa. Bauh. pin. 189.
      Habitat in Europa ad vias. ♃.

     

    Other articles:
    • Golden Checklist Flora:  Plantago patagonica;  

    Plantago patagonica Jacq. “Woolly Plantain”

     
     

    Jacquin (1793, v. 2., p. 9) short description with a reference to Jacq. coll. vol. 5. [Collectaneorum supplementum, where there is a longer description and a collecting locality.] and Jacquin (1793, v. 2, t.(plate) 306.

    Literature Cited:
    - Jacquin, Nikolaus Joseph, 1796.  

    Jacquin (1796, p. 35) ...

    Original Text Translation and Comments
    Crefcit ad Champion-River in Patagonia. Grows at Champion-River in Patagonia.

     

    Other articles:
    • Golden Checklist Flora:  Veronica americana;  

    Veronica americana Schwein. ex Benth. “American Speedwell”

     

    Literature Cited:
    - Bentham, George, 1846.  

    Bentham in DeCandolle (1846, v. 10, p. 468) ...

    Original Text
    47. V. Americana (Schweinitz! mss.), glabra, basi decumbens , superne erecta, foliis plerisque petiolatis ovatis oblongisve acutis vel obtusiusculis serratis crassiusculis basi truncato-subcordatis, capsula subrotunda turgida emarginata. ♃ in America boreali a Canada et Carolina usque ad flum. Oregon et in ins. Sitcha. V. Anagallis auct. plur. amer. non Linn. Species inter V. Anagallidem et V. Beccabungam fere media. Habitus potius prioris sed folia pleraque petiolata, breviora, latiora. Capsula et semina V. Beccabungae. Folia nunc vix semi-pollicaria, nunc 1-2poll. longa. (v. s. )

    Bentham wrote the section on Scrophulariaceae.

     

    Other articles:
    • Golden Checklist Flora:  Veronica anagallis-aquatica;  

    Veronica anagallis-aquatica L. “Water Speedwell”

     

    Literature Cited:
    - Linne´, Carl von, 1753.
    - Stearn, William T., 2004.  

    Linnaeus (1753, v. 1, p. 12) ...

    Anagall. 15. VERONICA racemis lateralibus, foliis lanceolatis ferratis, caule erecto. Fl. fuec. 10. Dalib. parif. 7.
      Veronica foliis oppofitis laevibus crenatis, floribus laxe fpicatis ex alis. Gron. virg. 4.
      Veronica aquatica major, folio oblongo. Bauh. pin. 252.
      Habitat in Europa ad foffas.

    The upside-down triangle (ᐁ) is a symbol for water, or aquatica, e.g., Anagall. ᐁ = Anagallis-aquatica (Stearn, 2004). Linneaus (1753) used symbols forming part of a specific epithet under Alisma, Campanula, Cotyledon, Scandix, Sysimbrium, and Veronica.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015) treats V. anagallis-aquatica as a native, and places V. catenata in synonomy.

    Literature Cited:
    - POWO, 2021 - 2023.  

    Kew POWO maps V. anagallis-aquatica as introduced to Colorado.

     

    Other articles:
    • Golden Checklist Flora:  Veronica arvensis;  

    Veronica arvensis L. “Corn Speedwell”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p. 13) ...

    Original Text
    21. VERONICA floribus folitariis, foliis cordatis incifis pedunculo longioribus. Fl. fuec. 16. Dalib. parif. 5. arvenfis
    Veronica foliis oppofitis cordatis crenatis, floribus folitariis feffilibus. Hort. cliff. 9. Gron. virg. 4. Roy. lugdb. 303.  
    Alfine veronicae foliis, flofculis cauliculis adhaerentibus. Bauh. pin. 250.  
    Habitat in Europae arvis, cultis. ☉.  

     

    Other articles:
    • Golden Checklist Flora:  Veronica biloba;  

    Veronica biloba L. “Twolobe Speedwell”

    (Syn: Pocilla biloba (L.) W. A. Weber)

    Literature Cited:
    - Linne´, Carl von, 1771.  

    Linnaeus (1771, p. 172) ...

    Original Text Translation and Interpretation
    Habitat inter Cappadociae fegetes. D. Schreber. ☉. She lives among the crops of Cappadocia [central Turkey]. D. Schreber. Annual.

     

    Other articles:
    • Golden Checklist Flora:  Veronica catenata;  

    Veronica catenata Pennell “Water Speedwell”

     

    Literature Cited:
    - Pennell, Francis W., 1921b.  

    Pennell (1921, v. 23, n. 266, p. 37) ...

    Original Text
    27. Veronica catenata Pennell, sp. nov.
    Flowering stem 1-3 dm. long, glabrous throughout. Leaves lanceolate, acute or acutish, crenate to nearly entire, 3-5 cm. long, 1 cm. wide, all clasping, when submersed elongating and reaching 12 cm. long and 2 cm. wide. Racemes axillary to the upper leaves, 6-12 cm. long, 15-25-flowered. Bracts narrowly lanceolate, 4-5 mm. long. Pedicels 3-5 mm. long, glabrous. Sepals 3-3.5 mm. long, lance-ovate, obtusish. Corolla-lobes pale-blue. Style 1.2-1.7 mm. long. Capsule 3 mm. long, 3.5 mm. wide, broad-globose, decidedly emarginate. Seeds 0.5 mm. long, yellow-brown.
    Type, Hot Springs, South Dakota, collected in flower and fruit June 16, 1892, P. A. Rydberg 926, in Herb. New York Botanical Garden. Named from the chain-like aspect of the long racemes of short-pedicelled flowers.
    Slow-flowing streams, plains, from North Dakota and Saskatchewan to Kansas and New Mexico, southward west to Nevada and southern California.

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Accepted by Weber & Wittmann (2012) in a paragraph that also mentions V. anagallis-aquatica noting that it is closely related.

    V. catenata is not identified as alien.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Treated by Ackerfield as a synonym of V. anagallis-aquatica.

    Literature Cited:
    - POWO, 2021 - 2023.  

    Kew Plants of the World Online accepts both V. anagallis-aquatica and V. catenata.

    According to Kew POWO, V. catenata does not occur in Colorado, and V. anagallis-aquatica is introduced to Colorado.

     

    Other articles:
    • Golden Checklist Flora:  Veronica peregrina ssp. xalapensis;  

    Veronica peregrina L. ssp. xalapensis (Kunth) Pennell “Neckweed”

    (Syn: Veronica xalapensis Kunth, V. peregrina L. var. xalapensis (Kunth) H.St.John )

    Literature Cited:
    - Kunth, Carol Sigismund, 1815-1825.  

    Kunth, Nov. Gen. Sp. [H.B.K.] 2(pt. 8): 389 (ed. qto.) (1818).

    2. VERONICA xalapensis. † 2. VERONICA xalapensis. nov. sp.
    V. caule erecto , pubescente ; foliis oblongis , cernulatis ; floribus axillaribus , solitariis , breviter pedunculatis , folio brevioribus ; laciniis calycinis oblongis , erectis. Veronica stems erect, pubescent leaves oblong, cerulate; flowers axillary, solitary, short pedunculate, shorter than leaves; calyx lobes oblong, erect.
    Crescit in Regno Mexicano prope Xalapa (alt. 630 hex.), in nemoribus Liquidambaris Styracifluae. ☉ Floret Febrruario. He grows in the Mexican kingdom near Xalapa (alt. 630 fathoms [=3780 ft.]), in the woods of the Liquidambar styracifluae [American sweetgum]. Annual. It blooms in February.
    Longer Latin description omitted.
    Differt a Veronica peregrina ; cui simillima, caule pubescente, foliis latioribus , floribus pedunculatis laciniisque calycinis latioribus. It differs from Veronica peregrina; which it most resembles, the stem is pubescent, the leaves wider, flowers pedunculate, with broad calyx lobes.

    Literature Cited:
    - Pennell, Francis W., 1919c.  

    Pennell (1919, v. 19, n. 9, p. 167) ...

    7a. Veronica peregrina xalapensis (H. B. K.) Pennell, comb. nov.
    Veronica xalapensis H. B. K., Nov. Gen. et Sp. 2: 389. 1817. “Crescit in Regno Mexicano prope Xalapa (alt. 630 hex.), in nemoribus Liquidambaris Styracifluae.”
    Occasional in cultivated soil. In the western half of the continent this glandular-pubescent plant completely replaces true peregrina. In the east it is only occasionally seen, and that probably as an introduction. Intergradation to the species seems to be complete.

     

    Other articles:
    • Golden Checklist Flora:  Galium aparine;  

    Galium aparine L. “Sticky Willy, Cleavers”

    Galium aparine L., Sp. Pl. 1: 108 (1753).

     

    Other articles:
    • Golden Checklist Flora:  Galium boreale;  

    Galium boreale S. Watson “Bedstraw”

     

     

    Other articles:
    • Golden Checklist Flora:  Galium triflorum;  

    Galium triflorum Michx. “Fragrant Bedstraw”

     

     

    Other articles:
    • Golden Checklist Flora:  Non-native Caprifoliaceae;  

    Caprifoliaceae in the Golden Flora

     

     

    Other articles:
    • Golden Checklist Flora:  Dipsacus fullonum;
    Full Size Image“Fuller's Teasel” at Tin Cup Ridge, Coll. No. 2677.
    Full Size ImageColl. No. 2677, Dipsacus fullonum  

    Dipsacus fullonum L. “Fuller's Teasel”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Original Text
    1. DIPSACUS foliis feffilibus ferratis. fullonum.
    Dipfacus foliis connato-perfoliatis, Hort. upf. 25. ariftis fructus rectis. Sauv. monfp. 156.  
    Dipfacus capitulis florum conicis. Hort. cliff. 29. Gron. virg. 15. Roy. lugdb. 188. Dalib. parif. 44.  
    Dipfacus fylveftris aut Virga Paftoris major. Bauh. pin. 385.  
    Dipfacus fylveftris. Dod. pempt. 735.  
    β Dipfacus fativus. Bauh. pin. 385. ariftis fructus hamatis. Sauv. monfp. 156.  
    Habitat in Gallia, Anglia, Italia. ♂  

     

    Other articles:
    • Golden Checklist Flora:  Dipsacus laciniatus;  

    Dipsacus laciniatus L. “Cutleaf Teasel”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Original Text
    2. DIPSACUS foliis connatis finuatis. laciniatus.
    Dipfacus folio laciniato. Hauh. pin. 385. Morif. hift. 3. p. 158. f. 7, t. 36. f. 4,  
    Habitat in Alfatia, Azow. ♂  

     

    Other articles:
    • Golden Checklist Flora:  Knautia macedonica;  

    Knautia macedonica Griesb. “Macedonian scabious”

     

     

    Other articles:
    • Golden Checklist Flora:  Lonicera morrowii;  

    Lonicera morrowii A. Gray “Morrow's honeysuckle”

     

    Literature Cited:
    - Gray, Asa, 1856.  

    Gray (1856, p. 305) in his preface to the list of plants collected in Japan ...

    Original Text
    This collection, hastily made as their small opportunities permitted, by Dr. Morrow and my early friend, Mr. Williams, of the American Mission at Macao, has proved a very interesting one, as might have been expected. No part of the world beyond his own country offers, as to its vegetation, a greater interest to the botanist of the United States than Japan. This is not the place to dwell upon the very remarkable relations which subsist between the Flora of Japan and that of the United States, which have been more or less noticed ever since Thunberg's Flora Japonica was published. As the present collection was principally made in the northern part of Japan, it was naturally anticipated that it would add to the Japanese Flora a considerable number of species identical with, or closely allied to, those already known to inhabit Kamtschatka and Northwest America. This proves to be the case ; and at the same time it has brought to view an equal number of Eastern United States forms, no insignificant portion of which are specifically identical. These interesting facts will arrest the botanist's attention as he turns over these pages. I refrain from commenting upon these relations until another and perhaps a larger collection of Japanese plants shall be made known, namely, that made by that excellent and most assiduous collector, Mr. Charles Wright, in the North Pacific Exploring Expedition, under Commodore Rodgers.
    The present collection makes known nearly forty new species and one new genus ; a very considerable number under the circumstances of the case. Thirteen of these are Carices ; but as most of these were gathered too early in the season, it is probable that the number will be somewhat reduced when they are better known. I have to thank my excellent friend, Dr. Boott, of London, the most learned Caricologist of the age, for having studied the specimens of this genus, and furnished the descriptions for this report. I have also to thank my promising young friend, Mr. Daniel C. Eaton, for having studied the Ferns of the collection, a department of botany to which he devotes special attention. Mr. Sullivant has obligingly determined the Mosses, and Prof. Harvey, of Dublin, the Algas of the collection.
    ASA GRAY.

    Gray (1856, p. 313) describes our plant ...

    Original Text
    Lonicera (Xylosteum) Morrowi (n. sp.) ; erecta, molliter tomentosa; foliis oblongis basi rotundata vel subcordata supra pubescentibus subtus canescenti-tomentosis ; pedunculis folio dimidio brevioribus flores longitudine asquantibus ; bracteis lanceolatis vel spathulatis ovariis plerumque discretis duplo quadruplove longioribus ; corolla ultra medium fissa basi hinc gibbosa ; filamentis basi styloque hirsutis. — Hakodadi, on hill-sides. Whole plant soft-downy, except the upper surface of the leaves. These are about an inch and a half long, by three-fourths of an inch in width, and on petioles a line and a half long. Corolla whitish ? somewhat pubescent ; the lobes narrowly oblong. Bracts variable in size, sometimes five lines long, foliaceous. The pubescence, and especially the long peduncles, must distinguish this from L. brachypoda, DC. The specific name is a deserved compliment to one of the collectors.

     

    Other articles:
    • Golden Checklist Flora:  Lonicera tatarica;  

    Lonicera tatarica L. “Tatarian Honeysuckle”

     

    Literature Cited:
    - Linne´, Carl von, 1753.
    - Stearn, William T., 2004.  

    Linnaeus (1753, v. 1, p. 173-174) ...

    Original Text
    5. LONICERA pedunculis bifloris, bassis diftinctis, foliis cordatis obtufis. Hort. npf. 42. tatarica.
    Chamaecerafus fructu gemino rubro, foliis glabris cordatis. Amm. ruth. 184.  
    Habitat in Tataria. ♄  

    In Linnaeus' time, “Tataria” referred to Central Asia and European Russia east of the river Don (Stearn, 1992).

     

    Other articles:
    • Golden Checklist Flora:  Symphoricarpos occidentalis;  

    Symphoricarpos occidentalis Hook. “Western Snowberry”

     

    Literature Cited:
    - Hooker, Sir William Jackson, 1829-1840.  

    Hooker (1840, v. 1, p. 285), volume 1 being concluded in 1834. IPNI shows the date of publication to be 1833.

    Original Text
    2. S. occidentalis ; spicis densis terminalibus axillaribusque nutantibus, corolla intus laciniisque dense barbatis, stylo staminibusque subexsertis. — Rich. in Frankl. 1st Journ. ed. 2. App. p. 6.
    Hab. Woody country, between lat. 54° and 64°, and known under tbe name of “Wolf-berry.” Dr. Richardson. Abundant about the Saskatchawan and Red River. Drummond. Douglas. Fort Vancouver, on tbe Columbia. Douglas. — Dr. Richardson has justly remarked of this plant “S. racemosi proximus:” and among the numerous specimens in the Herbarium, are some which appear almost as much allied to the one species as tbe other. But the majority of individuals of the two species are readily enough distinguished ; those belonging to the S. occidentalis, by their larger, less glaucous, more rigid, and denser foliage, (some of tbe leaves being two and a half inches long,) by the flowers arranged in dense drooping spikes, larger than in S. racemosus, and by the prominent style and stamens.

     

    Other articles:
    • Golden Checklist Flora:  Symphoricarpos rotundifolius;  

    Symphoricarpos rotundifolius A. Gray “Roundleaf Snowberry”

     

    Literature Cited:
    - Gray, Asa, 1852-1853.  

    Gray (1853, v. 5, p. 66) writing in Plantae Wrightianae described S. rotundifolius

    Symphoricarpos rotundifolius (sp. nov.) : foliis orbiculatis seu ovato-rotundis parvulis ramulisque molliter pubescentibus ; floribus axillaribus solitariis ; bracteolis ovario brevioribus dentibusque calycis cyathiformis ovatis obtusis glabris ; corolla infundibuliformi intus glabra, lobis stamina superantibus. — Sides of mountains around the copper mines, New Mexico ; Aug. (1388.) — Stems 3 or 4 feet high, much branched. Leaves 5 to 9 lines long and 4 to 9 wide, softly and finely pubescent above, and finely tomentose underneath, entire, or the larger often repand ; the petiole a line or less in length. Flowers almost sessile in the axils of the upper leaves. Calyx with the ovary a line and a half long, its lobes slightly if at all ciliate. Corolla 3 lines long, tubular-funnelform. Fruit not seen. — The leaves are rounded and more downy than in Nuttall's S. mollis, and the inflorescence as well as the shape of the corolla is entirely different. The flowers are more like those of S. glaucescens and S. montanus, but only half as large ; and they, as well as the leaves, are larger than in S. microphyllus (as described by Kunth), with which in other particulars its characters do not accord.

     

    Other articles:
    • Golden Checklist Flora:  Viburnum lantana;  

    Viburnum lantana L. “Wayfaringtree”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p. 268) ...

    Original Text
    Lantana. 5. VIBURNUM foliis ovatis ferratis venofis fubtus tomentofis Vir. cliff. 25. Hort. upf. 68 Roy. lugdb. 242. Sauv. monfp. 136.
      Viburnum. Hort. cliff. 107.
      Viburnum vulgo. Bauh. pin. 249.
      Lantana. Dod. pempt. 701.
      Habitat in Europae auftralioris fepibus afgillosis. [He lives on the hedges of the south of Europe.] ♄

     

    Other articles:
    • Golden Checklist Flora:  Sambucus canadensis;  

    Sambucus canadensis L. “American Black Elderberry”

     

     

    Literature Cited:
    - Linne´, Carl von, 1753.

    Other articles:
    • Golden Checklist Flora:  Campanula glomerata;

    Locations: Schweich Hill.  

    Campanula glomerata L. “Dane's Blood”

    There is one collection of “Dane's Blood” — Campanula glomerata L. — in Golden s.l. that was made on Schweich Hill. It is not really a garden escapee, since it was clearly thrown or dumped in the open space, i.e., it did not escape a garden on its own. Perhaps the correct term is that it was “yeeted” into the open space. It is the only collection from Jefferson County. There are three other collections scattered around Colorado, all of which appear to be garden escapees.

    It is one of four unrelated taxa known commonly as “Dane's Blood,” having in folklore sprung up in fields where Danes were slaughtered.

    The plant was described by Linnaeus (1753) who noted that it lived in England, France, and Sweden.

    Full Size Image
    Coll. No. 1982, Campanula glomerata

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, 166-167) ...

    Original Text
    glomerata. 19. CAMPANULA caule angulato finplici floribus feffilibus, capitulo terminali. Vir. cliff. 16. Fl.fuec. 188. it. gotl. 196. it. fcan. 306. Roy. lugdb. 245. Dalib. parif. 67.
      Campanula foliis lanceolato-ovatis crenatis, ramis capitulo florali terminatis. Hort. cliff. 64.
      Campanula pratenfis, flore conglomerato. Bauh. pin. 94.
      Trachelium alpinum, floribus conglomeratis, foliis afarinae rigidis & hirfutis. Herm. par. 235. t. 235.
      Habitat in Angliae, Galliae, Sueciae pratis aridis. ♃
      Folia ovato-oblinga, seffilia, obtufiufcula. Flores 3, fesfiles in alis foliorum; captiulum terminale.

     

    Other articles:
    • Golden Checklist Flora:  Campanula rapunculoides;  

    Campanula rapunculoides L. “Rampion Bellflower”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, 165) ...

    Original Text
    rapunculoides. 14. CAMPANULA foliis cordato-lanceolatis, cauleramofo, floribus fecundis fparfis. Vir. cliff. 17. Roy. lugdb. 246. Dalib. parif. 66.
      Campanula foliis ovatis crenatis, floribus per caulem uno verfu fparfis. Hort. cliff. 64.
      Campanula hortenfis, rapunculi radice. Bauh. pin. 94. Morif. hift. 2. p.f. 5. t. 3. f. 32.
      Habitat in Helvetia, Gallia.

     

    Other articles:
    • Golden Checklist Flora:  Campanula rotundifolia;  

    Campanula rotundifolia L. “Harebell”

     
     

    Linnaeus (1753, v. 1, p. 163) ...

    Original Text
    3. CAMPANULA foliis radicalibus reniformibus, caulinis linearibus. Fl. lapp. 83. Fl. fuec. 176 Hort. upf. 65. Roy. lugdb. 247. Dalib. parif. 66. rotundifolia.
    Campanula minor rotundifolia vulgaris. Bauh. pin. 93.  
    β Campanula minor rotundifolia alpina. Bauh. pin. 93. prodr. 34. t. 34.  
    γ Campanula alpina linifolia caerulea. Bauh. pin. 93. Fl. lapp. 84. it. gotl. 229.  
    Habitat in Europae pafcuis.  

     

    Literature Cited:
    - Herrando-Moraira, Sonia, et al., 2019.  

    Tribe Cardueae

    “Thistles” in the broad sense are in the tribe Cardueae Cassini, J. Phys. Chim. Hist. Nat. Arts. 88: 155. 1819 . Tribe Cynareae Lam. & DC. would seem to have precedence over Cardueae by nearly a decade. The North American representatives of Tribue Cardueae accepted by FNANM are as follows. Taxa found in Colorado are shown in bold.

    1. Leaves spiny
      1. Carduus L. Plumeless Thistle.
      2. Carlina L. Carline Thistle.
      3. Carthamus L. Distaff Thistle.
      4. Cirsium Mill. Thistle. All the native thistles in Colorado ae in the genus Cirsium Mill. In addition there are two non-native thistles that are also noxious weeds: C. arvense and C. vulgare.
      5. Echinops L. Globe Thistle.
      6. Onopordum L. Cotton Thistle.
      7. Silybum Adans. Milkthistle. Not listed for Colorado in Weber & Wittmann (2012) or Ackerfield (2015). There are two collections of Silybum marianum (L.) Gaertn. “Blessed Milk-thistle”, one from Larimer County and one from Saguache County.
    2. Leaves not spiny
      1. Acroptilon Cass. Hardheads.
      2. Amberboa (Pers.) Less. Sweet Sultan.
      3. Arctium L. Burdock.
      4. Centaurea L. Knapweed. Eight species of Centaurea are known from Colorado, all of which are non-native.
      5. Crupina (Pers.) DC. Crupina.
      6. Cynara L. Cynara. C. scolymus L. is the Globe Artichoke.
      7. Mantisalca Cass. Mantisalca.
      8. Plectocephalus D. Don in R. Sweet. Basketflower.
      9. Saussurea. Saw Wort.
      10. Volutaria Cass.

    There is a more recent paper (Herrando-Moraira, Sonia, et al. 2019) describing the phylogeny of Cardueae using Hyb-Seq data. It is, however, behind a paywall until August 1, 2020, and we eagerly await its unveiling.

     

    Other articles:
    • Golden Checklist Flora:  Achillea millefolium;  

    Achillea millefolium L. “Common Yarrow”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v.2, p. 899) published Achillea millefolium without mention of habitat in North America.

    Original Text Interpretation and Comments
    14. ACHILLEA foliis bipinnatis nudis: laciniis linearibus dentatis. Hort. cliff. 413. Fl. fuec. 705. Mat. med. 397. Roy. lugdb. 175. Gron. virg. 101. Dalib. parif. 263.
    Achillea foliis pinnato-pinnatis. Fl. lapp. 311.
    Millefolium vulgare album. Banh. pin. 140.
    β. Millefolium purpureum. Tabern hift. 130.

    Habitat in Europæ pascuis pratisque.

    Millefolium. Habitat in European meadows and pastures.

    Literature Cited:
    - Kalm, Pehr, 1762.
    Full Size ImageKalm (1762) mention of Achillea millefolium in Montreal, Canada.  

    Kalm (1762, v. III., p. 499) notes presence in Montreal, Canada, of A. millefolium, in addition to Prunella vulgaris, Oenothera biennis, Viola canadensis, and others. There might be additional information but I struggle to read Swedish in the old font used in the report.

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, p. 563) thought that Achillea millifolium may have been introduced from Europe.

    Original Text Interpretation and Comments
    3. A. foliis bipinnatifidis pilosis linearibus dentatis mucronatis, caulibus sulcatis. Smith fl. brit. 2. p. 908. Willd. sp. pl. 3. p. 2208.

    Icon. Fl. dan. 737. Engl. bot. 758.

    In fields an on road sides ; common. ♃. June-Aug. v. v. Probably introduced from Europe. Flowers white or rose-coloured.

    Millefolium. Probably need to see what Willdenow says about A. millefolium, i.e., does he mention North American occurrence? However, when I checked (11 May 2021), the Real Jardin Botanico (Madrid) server was not working.

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, v. 2, p. 171) also thought that A. millefolium was naturalized in North America.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1838-1843.  

    Torrey & Gray (1841-1843, v. 2, p. 409) recognized that A. millefolium occurred throughout North America, but also noted it was introduced from Europe into pastures, &c.

       

    Agoseris sp., “Goat-Chicory”

    [Greek agos, leader, and seris, chicory; allusion unclear]

    Agoseris has a New World, amphitropical distribution. All of the species are restricted to North America except A. coronopifolia (D’Urville) K. L. Chambers, which is found in temperate regions of southern South America. The South American disjunction appears to be the result of long-distance dispersal from North America (K. L. Chambers 1963).

    Literature Cited:
    - Schreber, Johann Christian Daniel van, 1791.

    Other articles:
    • Golden Checklist Flora:  Schreber, 1789-1791, publication details;  

    Schreber (1789-1791) published edition 8a of Linnaeus' Species Plantarum. In the second volume he described Krigia Schred. The volume is not available online, so I am unable to read the description. Publication of this volume preceeded the publication of Troximon by Gaertner. How much it preceeded Gaertner's publication, I don't know. But it was enough to give Krigia priority over Troximon.

    Literature Cited:
    - Robin, C. C. (Charles-Ce´sar), 1817.  

    C. C. Robin published an account of his travels in Louisiana, West Florida and the West Indies from 1802 to 1806. He included descriptions of plants observed in Louisiana. Rafinesque undertook to translate, revise, and improve (as he says!) the plant list, and to see it published in New York.

    Original Text English Translation Comments
    180. Troximon odoratum Raf. Hispidum, ramis fastigiatis, foliis sessilibus ovato-oblongis acutis integris ciliatis scabris, floribus racemosis nudis, perianthis pilosis 8 partitis, sub 12 floris Raf. — Chicoracee fenouillette Rob. P. 425. Stems four feet high, round milky; leaves a little thick, flowers pale yellow, sweet-scented, ligules five-toothed, anthers and stigmas very projecting, deep yellow ; seeds oblong compressed striated with a thick long down. This species together with Tr. Virginicum, Tr. Pallidum and Tr. Bulbosum will form the genus Troximon ; the other species which are acaules and with an embricated calyx, must form a peculiar genus which I shall call Agoseris. Raf. Found in woods.   The way I read this, Rafinesque has validly published Agoseris by saying that it is like Troximon that is acaulescent and has an imbricated calyx (or the involucral bracts are imbricated).
    Rafinesque gets/takes credit for the name Agoseris.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Troximon glaucum, Nuttall, 1841;  Agoseris parviflora, Nuttall, 1841;
    • Glossary:  rostrum;  

    Original Text Comments
    TROXIMON. (Nutt. Gen. Am., non Gaertner)  
    Capitulum many-flowered. Involucrum imbricate, subcampanulate, divisions lanceolate, distinct, or united at base. Receptacle naked, punctate. Achenium subterete, with ten obtuse ribs, attentuated above into a somewhat similarly striated, and rather short, thick rostrum. Pappus copious, setaceous, persistent, widest at base, longer than the achenium, and scarcely scabrous. — Stemless perennials, with fusiform roots, and mostly entire, linear, smooth, sublanceolate leaves. Scapes terete, exserted, one-flowered; flowers yellow or rose-coloured. OBS. The only species of this genus known to Gaertner, T. lanatum, is now referred to Scorzonera, the name thus unoccupied may, therefore, still be retained for the American species. The question is: was Troximon really available? Or, was Troximon illegitimate because Krigia had priority?
    Nuttall then goes on to propose Troximon glaucum (=Agoseris glauca) and T. parviflorum sp. Nov. (=A. parviflorum).

    Literature Cited:
    - Lee, Joongku, Bruce G. Baldwin, and L. D. Gottlieb, 2003.  

    Phylogenetic analysis of relationships in the Cichorieae place Agoseris in the Microseris clade along with Nothocalais, Uropappus, and Stebbinsoseris. Krigia (and Glyptopleura) was not placed in any major clade.

     

    Other articles:
    • Golden Checklist Flora:  Agoseris glauca;  

    Agoseris glauca (Pursh) Raf. “Pale Goat-Chicory”

     
     

    Troximon glaucum Pursh, Fl. Amer. Sept. (Pursh) 2: 505 (1813).

    Literature Cited:
    - Moulton, Gary E., 1999.
    - Pursh, Frederick, 1816.

    Other articles:
    • Golden Checklist Flora:  Pursh, 1814, publication details;  

    Pale Goat-Chicory was first published by Frederick Pursh in Volume 2 of his Flora Americae Septentrionalis.

    Original Text Translation and Interpretation Comments
    604. TROXIMON. Gaert. Carp. P. 360. Pers. Syn. 2 p. 360          
    1. T. scapo unifloro, calycinis foliolis imbricatis cuspidatis, foliis linearibus integerrimis utrinque glaucis. glaucum. Troximon with single-flowered scape, involucre with imbricate bracts with acute tips, green leaves are linear with entire margins. glaucum.    
    On the banks of the Missouri. ♂. v. s. ; v. v. in Hortis. Flowers bright yellow.   On the banks of the Missouri. Biennial. I have seen it dried. I have seen it live in gardens. Flowers bright Yellow.   Pursh doesn't tell us whose dried collection he saw. It could have been a Bradley collection. It probably was not a Lewis & Clark collection because Moulton (1999) does not list an Agoseris collection in the Lewis & Clark herbarium.  

    What we now consider to be a male symbol – ♂ – probably means the plant is biennial.

     

    Agoseris glauca (Pursh) Raf., Atlantic J. 6: 39 (1833).

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;  Troximon, Nuttall, 1841;
    • Glossary:  rostrum;  

    Original Text Comments
    Achenium terete, shortly rostrate, with obtuse ribs This is the first section of Nuttall's Troximon and will contain T. glaucum [=Agoseris glauca].

    T. parviflora [=A. parviflora] will be segregated into the second section that has a distinct rostrum (beak).

    Troximon glaucum. The involucrum is usually smooth, the divisions in about three series, the outer shorter, all of them lanceolate and acute.  
    HAB. On the plains of the Platte, and Missouri, about the Great Bend.  

    Literature Cited:
    - Dietrich, David Nathaniel Friedrich, 1847.

    Other articles:
    • Golden Checklist Flora:  Dietrich, 1847, publication details;  

    Though Rafinisque published Agoseris in 1817, it wasn't until 1847 that D. Dietrich placed Pursh's Troximon glaucum in Agoseris.

    3606. AGOSERIS Rafin. (1817). (Troximon Nutt.)
    2. A. cuspidata;
    2. A. glauca; glaucescens; fol. lineari-lanceolatis acutus integris scapoque glabriusculis; setis pappi capillaribus rigidis. Troximon Nutt. B. M. T, 1667. β dasycephalum Torr. Et Gray. T. glaucum α Hook. B. M. t 3462. Ammogeton scorzonerifolium Schrad. In Amer. Bor. ♃.
    3 A. parviflora (Troximon Nutt.):
    4. A. rosea
    5. A. taraxacifolia;

     

    Other articles:
    • Golden Checklist Flora:  Agoseris parviflora;
    Full Size ImageColl. No. 1101, Agoseris parviflora  

    Agoseris parviflora (Nutt.) D. Dietr. “Steppe Goat-Chicory”

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;  Troximon, Nuttall, 1841;  

    Nuttall (1841) described a new species of Troximon, T. parviflorum in describing the plants from his travels along the Oregon Trail in 1834 and 1835. It will become our current Agoseris parviflora.

    Original Text Comments
    †† Achenium compressed, with ten shallow, acute ribs, and attenuated into a distinct rostrum, shorter than the long and bristly pappus. Involucrum ovate, in about three series of unequal, lanceolate sepals. This is the second section of Nuttall's Troximon and it has a “distinct rostrum” or beak. The section starts off with Nuttall's new species T. parviflorum [=Agoseris parviflora].
    Troximon * parviflorum; leaves linear-lanceolate, acuminate, smooth or pubescent, often runcinately denticulate towards the base; scape pubescent at the summit, lanuginous; sepals nearly smooth, in three unequal series, lanceolate, acuminate; flowers yellow.
    Hab. On the plains of the Platte to the Rocky Mountains. About four or five inches high. The leaves about two lines wide, acuminated at each end. Pappus minutely scabrous; the outermost divisions of the involucrum only about half the length of the inner.

    Literature Cited:
    - Dietrich, David Nathaniel Friedrich, 1847.

    Other articles:
    • Golden Checklist Flora:  Dietrich, 1847, publication details;  

    Dietrich (1847) in his Synopsis Plantarum placed the North American Troximon into Agoseris Raf.

    3606. AGOSERIS Rafin. (1817). (Troximon Nutt.)
    . A. cuspidata;
    2. A. glauca;
    3 A. parviflora (Troximon Nutt.): glabriuscula; fol. Anguste lanceolato-linearibus acute asuminatis integris subretrorsp-denticulatis; pappo capillari rigido. In mont. Rocky. ♃.
    4. A. rosea
    5. A. taraxacifolia;

     

    Other articles:
    • Golden Checklist Flora:  Ambrosia acanthicarpa;  

    Ambrosia acanthicarpa Hook. “Flat-Spine Burr-Ragweed”

     

     

    Other articles:
    • Golden Checklist Flora:  Ambrosia artemisiifolia;  

    Ambrosia artemisiifolia L. “Annual Ragweed”

     

     

    Other articles:
    • Golden Checklist Flora:  Ambrosia psilostachya;  

    Ambrosia psilostachya DC. “Western Ragweed”

     

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1836.  

    DeCandolle (1836, v. 5, p. 526) published A. psilostachya from a collection by Berlander in Mexico.

    Original Text
    9. A. psilostachya, caule herbaceo erecto tereti paniculato-ramoso foliisque scabridis, foliis subsessilibus pinnatipartitis lobis lineari-lanceolatis acnminatis hinc inde pinnatim incisis, racemis plurimis gracilibus inter capitula mascula remota ebracteatis, fructibus infra apicem tuberculis obtusis 5-6 onustis apice conico elongato. — in Mexico inter San-Fernarido et Matamoros legit cl. Berlandier (pl. exs. n. 2280!). Fructus plurimi parvi in invol. foem. Capitula masc. pubero-scabrida parva. (v. s.)

     

    Other articles:
    • Golden Checklist Flora:  Ambrosia tomentosa;  

    Ambrosia tomentosa Nutt. “Ragweed”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    603. AMBROSIA. L. (Bitter-weed.)
    Monoicous (sic). — Masc. Calix 1-leaved. Anthers approximate, but not united. Receptacle naked. — Fem. Calix 1-leaved, entire or 5-toothed, 1-flowered. Corolla none. Nut formed from the indurated calix, 1-seeded.
    Tall herbaceous and mostly annual plants; leaves rough, the lower most opposite, the upper alternate, bipinnatifid, trifid, or rarely entire; flowers in long terminal and proximately axillar spikes, upper flowers masculine numerous, the lower fewer, feminine, glomerated, clusters 2 to 5-flowered, tribracteate.
    Species. 1. A. integrifolia. 2. bidentata. 3. trifida. 4. elatior. 5. artemisifolia. 6. paniculata. 7. heterophylla.
    8. * tomentosa. Perennial; stem low; leaves bipinnatifid, upper side white and tomentose; spikes solitary. Hab. In Upper Louisiana on the banks of the Missouri; rare. Only 1 or 2 feet high.
    A North American genus, with the exception of 1 species in Peru and another indigenous to the sea-coasts of the Levant.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;  

    FRANSERIA. (Cavan.)
    Franseria * bipinnatifida; ...
    Franseria * pumila; ...
    Franseria * discolor; ♃, root creeping; leaves ubterruptedly bipinnatifid, above nearly smooth, canescently and closely tomentose, segments subovate, acute, confluent in the wide rachis; stem short, with the lateral branched decumbent.
    Hab. In the Rocky Mountains, neat the Colorado of the West. A very remarkable and distinct, as well as elegant species. Stem about a span long, slightly pubescent; leaves on long petioles, with a lanceolate outline, acute, about six inches lone, white beneath, green above, the pinnatifid segments lanceolate, the rachis incisely toothed. Male florets rather numerous; receptacle with narrow, pubescent palea; involucrum about five or six-toothed; female flowers few, fruit spiny.
    Franseria * cuneifolia; ...

     

    Other articles:
    • Golden Checklist Flora:  Ambrosia trifida;  

    Ambrosia trifida L. “Giant Ragweed”

     

     

    Other articles:
    • Golden Checklist Flora:  Anaphalis margaritacea;  

    Anaphalis margaritacea (L.) Bentham & Hooker “Western Pearly Everlasting”

     
     

    Linnaeus (1753, v. 2, p. 850) was fairly clear that G. margaritaceum was native to North America.

    Original Text
    margaritace-
    um.
    3. GNAPHALIUM foliis lineari-lanceolatis acuminatis alternis, caule fuperne ramofo, corymbis faftigiatis. Hort. cliff. 401. Hort. upf. 255. Gmel. fib. 2. p. 107.

    Gnaphalium latifolium americanum. Bauh. pin. 263.

    Gnaphalium americanum. Clif. hift. 1. p. 327.

    Habitat in America feptentrionali, kamtfchatca.

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1836.  

    DeCandolle (1837, v. 6, p. 271) published Anaphalis, but he placed our margaritacea in Antennaria Sect. II. Margaripes.

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittman (2012) state that Anaphalis is an anagram of Gnaphalium.

     

    Other articles:
    • Golden Checklist Flora:  Antennaria parvifolia;
    Full Size ImageHabitat of Coll. No. 1128, Antennaria parvifolia
    Full Size ImageColl. No. 1909, Antennaria parvifolia  

    Antennaria parvifolia Nutt. “Small-Leaf Pussytoes”

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;
    • Glossary:  sarment;  

    Nuttall (1841) first described A. parvifolia collected on his trip on the Oregon Trail with the Wyeth expedition.

    Original Text
    Antennaria * parvifolia; subcaespitose, with procumbent sarments; stem simple; lower leaves spathulate, or spathulate-linear, the upper linear, all whitely tomentose; flowers conglomerate; scales of the involucrum oblong-ovate, eroded, yellow.
    Hab. On the Black Hills and plains of the upper part of the Platte. A dwarf species, spreading out in canescent tufts with very small leaves, which are about half or three quarters of an inch long, and about two or three lines wide; the flowers in an irregular, somewhat round mass, not a circular corymb, with the scales of the corynb sulphur yellow, and very conspicuous. Radical leaves somewhat rhomboidally spathulate. The pappus of the male flower is very conspicuously clavellate: the female flower has purple oblong-lanceolate scales to the involucrum, and a filiform pappus. A specimen of this sex from Altai has a near resemblance to our plant, but is larger in all its parts, and is the A. hyperborea of Don.

    Literature Cited:
    - Bayer, Randall J., 1990.  

    Randal J. Bayer, who wrote the treatment of Antennaria for the Flora of North America published a cladistic analysis (Bayer, 1990) of the sexually reproducing Antennaria. However, A. parvifolia is not one of them because some individuals are dioecious and others are apomictic. The article is behind a paywall. I thought this was interesting, “ … polyploidy and two asexual means of reproduction, agamospermy and horizontal stolons, have evolved in the more specialized groups, the Pulcherrimae and Catipes … ” This would imply to me that A. parvifolia would be a member of one of those two groups.

     

    Other articles:
    • Golden Checklist Flora:  Antennaria rosea;  

    Antennaria rosea Greene “Rosy Pussytoes”

     

    Literature Cited:
    - Greene, Edward L., 1898.  

    Greene (1898, p. 281) in his “Sect. 2. Some northern species of Antennaria” described a new Antennaria.

    Original Text
    A. rosea. A. dioica, var. rosea, Eaton, Bot. King Exp. 186. A. parvifolia, var. rosea, Greene, p. 175 supra. A. parvifolia, Nutt. in small part, but not of spec. char. Plant by no means small, often 12, sometimes 14 or 15 inches high, yet frequently only 6 or 8 inches: leaves comparatively small, of the thinnest, as to texture, only canescently tomentose, but permanently so on both faces, the quite gradually dilated upper portion acute; cauline long and narrow, acute or acuminate: heads small, closely compacted pound cymose rounded cluster: bracts of the involucre pluriserial, their basal part concealed by wool, the tips from broad and obtuse in the outer series to narrow and acute in the inner, all rose-red.
    Of this only the female plant is known to me; which is the more remarkable in view of the fact that no other north-western Antennaria so abounds in every large herbarium. It is a dry-ground species of subalpine habitat, with either short or elongated slender dry and subligneous stolons; in this quite unlike the true A. parvifolia. From A. hyperborea more difficult to distinguish it, except by the looser inflorescence, longer heads, less woolly involucre, and fewer paler narrower bracts of the last named. I give the following rather copious list of localities for A. rosea. North Park, Colorado, Chas. S. Sheldon, n. 128 ; Yellowstone Park, Frank Tweedy, n. 728; mountain meadows in Kootenai Co., Idaho, J. B. Leiberg, n. 646 ; Nez Perces Co., Idaho, Heller, n. 3441 ; Salmon River, British Columbia, Dawson (Can. Surv. n. 11281); Spence's Bridge, B. C, Macoun, n. 11282; summit of Mt. Arrowsmith, Vancouver Island, Macoun, n. 11279; pine woods near Spokane, Washington, C. V. Piper, n. 2273; Mt. Hood, Oregon, Thos. Howell; Crooked Creek, southeastern Oregon, and Warner Range, northeastern California, Mrs. Austin.

     

    Other articles:
    • Golden Checklist Flora:  Arnica cordifolia;  

    Arnica cordifolia Hook. “Heart-Leaf Leopardbane”

     

    Literature Cited:
    - Hooker, Sir William Jackson, 1829-1840.  

    Hooker (1834, v. 1, p. 334) described A. cordifolia from collections by Drummond in the Rocky Mountains and by Douglas along the Columbia River in Washington Territory.

    Original Text Comments and Interpretation
    4. A. cordifolia ; caule elato, foliis cordatis glabriusculis basi sinu distincto, inferioribus longe petiolatis pari supremo solummodo sessili, panicula 1-3-flora.  
    Hab. Alpine woods of the Rocky Mountains, on the east side, Drummond ; and on the west side, in mountain woods between the Kettle Falls and Spokan River, and in the Blue Mountains. Douglas. — This appears quite distinct from the preceding, and is very constant in the shape and petiolation of its leaves. The preceding was A. menziesii.

     

    Other articles:
    • Golden Checklist Flora:  Arnica fulgens;  

    Arnica fulgens Pursh “Shining Leopardbane”

     

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, p. 527) published A. fulgens from an unidentified collection on the banks of the Missouri. There is no Lewis & Clark collection of this taxon, so it may have been an unattributed Bradbury or Nuttall collection.

    Original Text Comments
    3. A. pubescens ; foliis radicalibus lanceolatis obtusiusculis basi attenuatis petiolatis trinervibus, caulinis oppositis remotis linearibus, caule unifloro.

    On the banks of the Missouri. ♃. v. s. About a foot or more high, very slender ; leaves on the stem generally two pairs ; flowers somewhat smaller than the preceding, of a very deep and beautiful yellow.

    fulgens. Pursh has seen a dried specimen, but does not say whose specimen or where he saw it.

    The preceding was A. plantaginea which Nuttall will reduce to a variety of A. fulgens.

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, v. 2, p. 164) treated A. fulgens as a variety of A. montana.

    Original Text
    563. ARNICA. L.
    Calix hemispherical, leaflets equal, mostly in a simple series? Radial florets often producing 5 filaments destitute of anthers. Receptacle naked. Pappus simple, scabrous.
    A polymorphous and divided genus? Some of the species caulescent and also shrubby; those of Europe and North America, with a few others, often scapigerous, scapes 1-flowered, sometimes producing 1 or 2 pair of opposite leaves; flowers mostly yellow.
    Species. 1. A. montana. β fulgens. A. fulgens. Ph. 2. p. 527. Scarcely dissimilar to specimens of the alpine variety in the Banksian herbarium. Hab. On the margins of marshy springs and in depressed situations, from the Arikarees to Fort Mandan, and probably as far as the Mountains. Flowering in July. Flowers bright yellow. Obs. Minutely pubescent. Scape about 12 inches high, mostly with 2 pair of leaves, terminated by 1 rarely 3 flowers. Leaves oblong-lanceolate, 3 to 5 nerved; summit of the caudex tomentose. Calix a simple series of leaflets, somewhat hirsute. Tube of the florets pilose; rays without filaments; seed hirsute, slender. A. plantaginea, of Pursh from the specimen which I have seen, appears also a mere variety of the above.
    2. Doronicum. 3. nudicaule. Doroonicum nudicaule. Mich. 2. p. 121. A. Claytoni. PH. 4. maritima.
    A genus principally indigenous to Europe and the Cape of Good Hope, there are also 2 species in South America, 2 in Japan, 1 in Arabia Felix, and a shrubby species, or something else, in New Zealand.

       

    Artemisia L.

    Artemisia L. is the largest genus of family Asteraceae, comprising about 350–500 diverse species. Artemisia has two basic chromosome numbers, with ploidy levels x=9 and x=8. Chromosome number in diploid is most often 2n=18 or 16. The genus is divided into five large groups Absinthium DC., Artemisia L., Dracunculus Besser, Seriphidium Besser and Tridantatae (Rydb.) McArthur. Its phylogeny was based on the two hypothesized evolutionary trends, loss of fertility in the disc florets and loss of ray florets.

       

    Artemisia Linnaeus subg. Absinthium DC.

     
     

    Four taxa of Artemisia within subgroup Absinthium are recognized by Ackerfield (2015).

    • A. absinthium, non-native
    • A. frigida Willd., native, E. Europe to Temp. Asia, N. America.
    • A. pattersonii A. Gray, native, =Delwiensia pattersonii (A. Gray) W. A. Weber & R. C. Wittmann, not universally accepted, native from Wyoming to New Mexico,
    • A. scopulorum, native, high altitude, e.g., Mt. Evans, west central United States.
    One taxon in Artemisia subgroup Absinthium is not recognized by Ackerfield (2015)
    • A. rupestris L., non-native, Europe to Siberia and Afghanistan.

       

    Artemisia Linnaeus subg. Dracunculus Besser, Bull. Soc. Imp. Naturalistes Moscou. 1: 223. 1829.

     

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Four species of Artemisia within subgroup dracunculus are recognized by Ackerfield (2015).

    • A. campestris, 493 collections, widely distributed except northwest Colorado, including 20 in Jefferson County.
    • A. dracunculus, 691 collections, including 27 in Jefferson County.
    • A. filifolia, 247 collections, mostly eastern Colorado, esp., out on the plains, not in Jefferson County.
    • A. pedatifida, 12 collections, mostly Moffatt County, none in Jefferson County.

    Five secies are not recognized by Ackerfield (2015):

    • A. aleutica, no collections.
    • A. borealis, 191 collections in Colorado, widely scattered in the mountains, treated by Ackerfield (2015) as A campestris var. purshii. The count of collections is not included in the count of A. campestris above.
    • A. porteri, no collections in Colorado.
    • A. pycnocephala, no collections in Colorado.

     

    Other articles:
    • Golden Checklist Flora:  Artemisia absinthium;  

    Artemisia absinthium L. “Common Wormwood”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Abfinthium 12. ARTEMISIA foliis conpofitis multifidis, floribus fubglobofis pendulis: receptaculo villofo. Hort. cliff. 404. Fl. fuec. 670, mat. med. 386. Roy. lugdb. 153.
      Abfinthium incanum, foliis compofitis latiufcule multifidis, floribus fubglobofis pendulis. Gmel. fib. 3. p. 129. t. 63.
      Abfinthium ponticum f. romanum, officinarum f. diofcoridis. Bauh. pin. 138.
      Abfinthium. Cam. epit. 452.
      Habitat in Europae ruderatis aridis.

     

    Other articles:
    • Golden Checklist Flora:  Artemisia campestris;  Notes on Artemisia dracunculus, Linnaeus, 1753;  

    Artemisia campestris L. “Field Sagewort”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 846) published A. campestris stating that its habitat was dry, sunny fields in Europe.

    Original Text Comments and Interpretation
    campeftris. 4. ARTEMISIA foliis multifidis linearibus, caulibus procumbentibus virgatis. Hort. cliff. 403. Fl. fuec. 668. Roy. lugdb. 154. Gmel. fib. 2. p. 117.

    Abrotanum campeftre. Bauh. pin. 136.

    Ambrofia altera. Cam. epit. 597.

    Habitat in Europæ campis apricis, aridis.

     

    Habitat in sunny, dry European fields.

    Literature Cited:
    - Flora of North America Editorial Committee, eds., 2006.  

    The Flora of North America (Flora of North America Editorial Committee, eds., 2006) was written by Leila M. Shultz. There is a local connection in that many collections at Rocky Flats were made by Ms. Shultz, who was working with G. Kunkel.

    In her description of A. campestris, Ms. Shultz states that the species varies, and that each morphological form grades into another. Taking a conservative approach, she recognizes three subspecies: subsp. pacifica (Nuttall) H. M. Hall & Clements, subsp. canadensis (Michaux) Scoggan, and subsp. caudata (Michaux) H. M. Hall & Clements. Subspecies caudata and pacifica are known to occur in Colorado.

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.

    Other articles:
    • Golden Checklist Flora:  Notes on Oligosporus, Cassini, 1817;  

    Weber & Wittmann (2012, p. 104) do not accept presence of A. campestris L. in Colorado. Instead, they accept Oligosporus caudatus (Michx.) Poljakov and O. pacificus (Nutt.) Poljakov.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015, p. 121) accepts three varieties of A. campestris L. in Colorado, variety purshii (Hook.) Cronquist, variety caudata (Michx.) Palmer & Steyerm., and variety pacifica (Nutt.) M. Peck.

     

    Other articles:
    • Golden Checklist Flora:  Artemisia dracunculus L.;  

    Artemisia dracunculus L. “Tarragon, Dragon Wort”

     

    Literature Cited:
    - Linne´, Carl von, 1753.

    Other articles:
    • Golden Checklist Flora:  Notes on Artemisia campestris;  

    Linnaeus (1753, v. 2, p. 849) ....

    Original Text
    16. ARTEMISIA foliis lanceolatis glabris integerrimis. Hort. cliff. 403. Hort. upf. 256. Gmel.fib. 2. p. 126. t. 59. & 60. f. 1.

    Abrotanum lini folio acriori & odorato. Tournef. inft. 459.

    Dracunculus hortenfis. Bauh. pin. 98.

    Draco herba. Dod. pempt. 709.

    Habitat in Sibiria, Tataria. ♃

    Dracunculus

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.

    Other articles:
    • Golden Checklist Flora:  Notes on Oligosporus, Cassini, 1817;  

    Weber & Wittman (2015, p. 103) recognize Oligosporus dracunculus (L.) Poljakov subsp. glaucus (Pallas) Löve & Löve for our Artemisia dracunculus.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015, p. 121) accepts A. dracunculus without infraspecific taxa.

     

    Other articles:
    • Golden Checklist Flora:  Artemisia filifolia;  

    Artemisia filifolia Torrey “Sand Sage, Old-Man Sagebrush”

     

    Literature Cited:
    - Torrey, John G., 1828.  

    In describing the collections of Edwin James, MD, John Torrey (1828, v. 2, p. 211) informally “published” A. filifolia.

    Original Text
    210. Artemisia ludoviciana, Nutt. gen. ii. p. 143. Arid plains of the Platte, with all the following of Nuttall : A. Serrata, columbiensis, longifolia, cernua, and canadensis.

    Obs. Besides the above six species, Dr. James found another, which appears to be new, but the specimen is without lower leaves, and cannot, therefore, be certainly determined.
    Artemisia filifolia, caule herbaceo ? ramoso ? foliis canescenti-pubescentibus ; superioribus simplicibus, filiformibus, et subpinnatifidis, semiteretibus ; floribus conglomeratis.

    Literature Cited:
    - Goodman, George J., and Cheryl A. Lawson, 1995.  

    Goodman & Lawson (1995, p. 159) note that James could have collected A. filifolia in any one of five states the expedition passed through, and that the species occurs abundantly in the Texas panhandle which is perhaps the most likely collecting site.

     

    Other articles:
    • Golden Checklist Flora:  Artemisia frigida;  

    Artemisia frigida Willd. “Prairie Sagewort”

     

    Literature Cited:
    - Stearn, William T., 2004.
    - Willdenow, Carl L., 1797-1830.  

    Willdenow (1803v. 3, pt. 3, p. 1838) published A. fridiga ...

    Original Text
    *51. ARTEMISIA frigida. W.
    A. foliis incanis pinnatis , pinnis tripartitis linearibus acutis, floralibus pinnatis tripartitisve, caule adfcendente, floribus globofis nutantibus. W.
    Kalter Beyfus. W.
    Habitat in aridis frigidis Dauuriae. ♃.
    Habitu praecedenti quoad Jiguram Gmelini aliquatenus affinis fed toto coelo diverfa. Caules adfceudentes femipedales vel pedales ramofi, inferne glabri, fuperne fubcanefcentes. Folia incana, caulina pinnata, foliolis tripartitis linearibus acutis, quadrantem pollicis longa fesfilia; radicalia petiolata ejusdem formae et magnitudinis , floralia triplo minora pinnata vel tantum tripartita fesflia. Panicula fimplex, ramis fimplicibus longis erectis. Flores globoji breve pedunculati nutantes maguitudine Abfinthii. Receptaculum villofum. W.

    I think the location of “Dauuriae” is likely equivalent to “Dauria,” “Dahuria,” or “Davuria,” which Stearn (2004) cites as a name for a region of southeast Siberia.

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Artemisia ludoviciana;  Nuttall, 1818, publication details;
    • Glossary:  connivent;
    Full Size ImageColl. No. 1281, Artemisia ludoviciana  

    Artemisia ludoviciana Nutt. “Silver Wormwood”

    553. ARTEMISIA. L. (Wormwood, Southern-wood, &c.)
    Calix imbricated, scales rounded, connivent. Rays of the corolla none. Receptacle subvillous, or nearly naked. Pappus none.
    Shrubby or herbaceous; leaves mostly multifid, flowers often racemose.
    § 1. Leaves simple.
    Species. 1. A * longifolia. ... 2. * serrata. ... 3. * columbiensis. ... 4. * Gnaphaloides. ...
    5. * ludoviciana. Stem simple and herbaceous; lower leaves incise, subpinnatifid, the upper lanceolate and en- entire(sic), on both sides pubescent, beneath tomentose; flowers ovate, erect and sessile; calix pubescent, panicle simple. — Hab. On the banks of the Missisippi, near St. Louis; also on the alluvial plains of the Missouri. Obs. Perennial. Stem about 2 feet high. Lower stem leaves lanceolate, irregularly and divaricately laciniate, segments entire, oblong-lanceolate and acute.
    6. * cernua. ... 7. chinensis ...
    § ii. Leaves compound; stem paniculate.
    8. Sontonia? Ph. 9. sericea.

    The specific epithet “ludoviciana” is a Latinization of “Louisiana.”

     

    Other articles:
    • Golden Checklist Flora:  Bahia dissecta;  

    Bahia dissecta (A. Gray) Britton “Ragleaf Bahia”

     

    Literature Cited:
    - Gray, Asa, 1849.

    Locations: Mora River.  

    Gray (1849, p. 104) was unsure what name to apply to Fendler's and Fremont's specimens because they were incomplete.

    † 425. Amauria ? dissecta (sp. nov.) : hierbacea, puberula ; caule adscendente apice corymbosi-polycephalo ; foliis alternis petiolatis biternatisectis segmentis cuneiformibus vel sublincaribus saepius 2-3-fidis, summis parvis ; pedunculis glandulosis ; involucri squamis oblongo-lanceolatis subtriseriatis, intimis subscariosis ; receptaculo convexo ; ligulis circiter 16; styli ramis fl. disci cono brevissimo truncati-capitatis ; acheniis ad angulos laevibus. — A few miles east of Mora River ; Aug. (537.) Also gathered in Fremont's third expedition, probably towards the head-waters of the Arkansas. — Stem 12 or 15 inches high, apparently from a perennial root. Leaves about an inch in diameter, cut into narrow divisions. Peduncles clothed both with viscous and capitate-glandular hairs. Involucre herbaceous, more or less viscous, a third of an inch in diameter. Receptacle entirely destitute of chaff. Flowers all yellow: rays linear-oblong, 2-3-toothed ; the tube very glandular. Disk-corollas with the slender tube extremely glandular, the expanded 5-cleft limb slightly so. Branches of the style short, flattish-semiterete, capitate with a very short and flattish obtuse cone. Achenia cuneate-linear, slender, compressed-quadrangular, smooth ; the ovary sprinkled with sparse and minute hairs. Pappus none. — The specimen of Fendler has not matured fruit ; and the stamens are abortive in all the disk-flowers. The specimen from Fremont's collection, communicated by Dr. Torrey, is very imperfect, but has ripe achenia. From the character of the Californian genus Amauria, Benth. in Bot. Voy. Sulph. p. 31, this plant differs very essentially in the styles, and in the convex receptacle. But I am unwilling to constitute it a distinct genus upon the present imperfect materials.*

    Literature Cited:
    - Britton, Nathaniel Lord, 1889.  

    Britton (1889, p. 68) published Bahia dissecta from collections by Dr. E. A. Mearns, in the Mongollon and San Francisco Mountains, Arizona.

    Original Text Comments
    Bahia dissecta (Gray). Amauria (?) dissecta, Gray. Mem. Amer. Acad. iv. 104 (1849); Villanova chrysanthemoides, Gray, Smithsonian Contr. v. 96 (1853); Bahia chrysanthemoides, Gray, Proc. Amer. Acad. xix. 28 (1883). Copper Cañon. A form with involucral scales not acuminate (183). It is unclear which Copper Cañon this might be. There is no Copper Cañon contained within the current named San Francisco Mountains or the Mongollon Rim. The closest might be Copper Canyon along US Interstate 17 near Camp Verde.

    Literature Cited:
    - Rydberg, Per Axel, 1915.

    Locations: Mora River.  

    Rydberg (1915, v. 34, pt. 1 [1914]) published Amauriopsis with A. dissecta being the type species.

    Original Text Comments
    28. AMAURIOPSIS Rydberg, gen. nov.  
    Glandular-pubescent annuals. Leaves alternate, twice or thrice ternately divided. Heads in leafy corymbs, radiate. Involucre hemispheric; bracts 16-20, herbaceous, oblanceolate in about 3 series. Receptacle fiat, alveolate. Ray-flowers 16-20, pistillate, fertile; ligules cuneate, 3-cleft. Disk-flowers numerous, hermaphrodite and fertile; corolla-tube densely glandular, longer than the funnelform throat; teeth lanceolate, longer than the throat. Achenes elongate and narrowly obpyramidal, 4-angled, striate, rounded at the apex. Pappus wanting.  
    Type species, Amauria dissecta A. Gray.  
    1. Amauriopsis dissecta (A. Gray) Rydberg.  

    Amauria dissecta A. Gray, Mem. Am. Acad. II. 4: 104. 1849.
    Villanova chrysanthemoides A. Gray, PI. Wright. 2: 96. 1853.
    Bahia chrysanthemoides A. Gray, Proc. Am. Acad. 19: 28. 1883.
    Bahia dissecta Britten, Trans. N. Y. Acad. Sci. 8: 68. 1888.
    Eriophyllum chrysanthemoides Kuntze, Rev. Gen. 307. 1891.
    Villanova dissecta Rydb. Bull. Torrey Club 37: 333. 1910.
     
    A tall annual; stems 3-6 dm. high, puberulent and glandular, especially above; leaves 1-3 times ternately divided into obovate, oblong, or oblanceolate toothed divisions, puberulent; involucre hemispheric, 6 mm. high, 10-13 mm. broad; bracts oblanceolate, acuminate. glandular-hirsute; ligules spatulate, 6-8 mm. long, 3-cleft with rounded teeth; disk-corollas 3 mm. long; achenes narrowly obpyramidal, 4 mm. long, 0.5 mm. thick, glandular-puberulent,  

    Type locality: A few miles east of Mora River [New Mexico].
    Distribution: Wyoming to New Mexico, Arizona, and Chihuahua.
    Illustration: Clements, Rocky Mt. Fl. pl. 41, f. 1.
     

    Literature Cited:
    - Baldwin, Bruce G., and Kenneth R. Wood, 2016.  

    Baldwin and Wood (2016) examined phyllotaxy within the Bahia alliance and, among other findings, showed that merger of Amauriopsis and Hymenothrix into a common, monophyletic genus within the Bahia alliance would simplify the taxonomy. Hymenothrix has priority for such a group, which is characterized in part by alternate leaves, obtuse to acute style-branch apices, often zygomorphic disc corollas (of outer florets), and pappus scales generally ≥ 10 or absent.

    Literature Cited:
    - SEINet, 2019+.  

    The SEINet taxon tree treats Amauriopsis dissecta (syn: Bahia dissecta) separatelty from Hymenothrix dissecta. Therefore, it is necessary to search for both when searching for either.

     

    Other articles:
    • Golden Checklist Flora:  Balsamorhiza sagittata;  

    Balsamorhiza sagittata (Pursh) Nutt. “Arrow-Leaf Balsam Root”

     

    Other articles:
    • Tin Cup Ridge (social trail):   at Coll. 1109;
    • Field Notes:   Coll. No. 1109, 14 May 2015;
    Full Size ImageColl. No. 1109, Balsamorhiza sagittata  

    How the heck did it get to Tin Cup Ridge?

    Literature Cited:
    - Pursh, Frederick, 1816.  

    Original Text
    649. BUPHTHALMUM. Gen. pl. 1231.
      ...
    sagittatum. 3. B. tomentosum ; foliis radicalibus longissime petiolatis oblongis sagittatis integerrimus subtrinervibus, caulinis oblongis in petiolum attenuatis, caule subtrifloro, calycinis foliolis exterioribus disco longioribus.
      On dry barren hills, in the Rocky-mountains. M. Lewis. ♃ June, July. v. s. in Herb. Lewis. Flowers large, yellow. The natives eat the young stems as they spring up, raw.

    Literature Cited:
    - Nuttall, Thomas, 1834a.  

    From Nuttall's (1834a) catalogue of plants collected by Nathaniel Wyeth.

    Original Text
    ESPELETIA, Humb. and Bonpl. — BALSAMORHIZA, Hooker.
    Calyx imbricatus, subsquarrosus, foliaceus. Colollulae radii femineae, vix dentatae. Receptaculum planum paleaceum. Pappus nullus. Semina compressa planiascula, subquadrangulata. Herba perennis, pumila, grandiflora. Helianthi facie ; radix resinosa.
    This genus has also been proposed by Dr. Hooker in his Flora Boreali-Americana, under the name of Balsamorhiza, for a species of Heliopsis there described.
    66. ESPELETIA sagittata. Tomentosa, incana, foliis radicalibus longe petiolatis cordato-hastatis integris acutis, caulinis paucis lineari-oblongis in periolum attenuatis, caule subtrifloro pumilo, calicibus foliosis squarrosis, pl. 4. BUPHTHALMUM sagittatum, Ph. 2, p. 564.
    ♃ With a large yellowish root like a dock. Radical leaves about the length of the stem, seven or eight inches, softly and copiously tomentose, hoary. The stem also downy, resembling a scape, with about two or three small leaves like bracts on its upper part. Calyx very white and softly tomentose, leafy, the inner leaves linear-lanceolate and somewhat acute. Rays bright yellow, very large, from sixteen to eighteen, bidentate at the tips. Seed flattish, elliptic, very smooth, and wholly devoid of any vestige of pappus.
    Hab. On the borders of Flat-Head river. Flowering in June.
    The root of this plant also, when fermented a day or two in the ground, in a hole made for the purpose, and heated with hot stones, is then eaten and possesses an agreeable saccharine taste. The stems are never employed for food.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.  

    Original Text
    § II. * ARTORHIZA. — Leaves entire, deltoid or cordate; involucrum very leafy at base; rays numerous. Receptacle flat.
    Balsamorhiza sagittata. Buphthalmum sagittatum? Pursh, Vol. II., p. 564. Espeltia sagittata; Nutt. in Journ, Acad. Nat. Sci. Philad., Vol. VII., p. 39. Canescently tomentose; stem low, about one to three flowered; radical leaves cordate-ovate, entire, somethat three-nerved at base; cauline leaves linear, attenuated below; external leaves of the involucrum longer than the inner, spreading, lanceolate, densely tomentose; rays numerous, (twenty to twenty-four.)
    Hab. In the Rocky Mountains, by Flat-Head River, towards the sources of the Oregon. Flower large and showy, about three to four inches in diameter, while the scapoid stem is not more than a span high. Stigmas very hirsute, filiform. Rays feminine, with infertile filaments.

     

    Literature Cited:
    - Elliott, Stephen, 1821-1824.

    Other articles:
    • Golden Checklist Flora:  Elliott, 1821-1824, publication details;  

    Brickellia Elliot

    Elliott (1823) published the name Brickellia in his Sketch of the Botany of South Carolina and Georgia.

    BRICKELLIA. E.
    Involucrum polyphyllum, inbricatum. Semina sub glabra, 10 striata. Pappus pilosus sive scaber. Receptaculum nudum, punctatum. Involucrum many leaved, imbricate. Seed nearly glabrous, 10 streaked. Pappus hairy or scabrous. Receptacle naked, dotted.
    1. Cordifolia. E. …
    This plant which in its artificial characters is closely allied to the Eupatorium, differing principally in size and number, in its general aspect, bear more resemblance to the Vernonia. I have named it in commemoration of Dr. John Brickell, of Savannah, who at one period of his life paid much attention to the botany of this country, and made known to Dr. Muhlenberg, Fraser and others, many of its undescribed plants.
    The name is conserved against Brickellia Raf., Med. Repos. Ser. 2, 5:353 (1808), nom. Illeg. Nom rej., which turns out to be a synonym of Gilia Ruiz & Pav.

    Literature Cited:
    - Schilling, Edward, Randall W. Scott, and Jose L. Panero, 2015.  

    Schilling, et al., 2015, infrageneric classification for Brickellia is behind a paywall.

    Literature Cited:
    - Schilling, Edward E., Jose L. Panero, Bonnie S. Crozier, Randall W. Scott, and Patricia Davila, 2015.  

    Schilling, et al., 2015a, also behind a paywall, results supported the hypothesis that Brickellia is monophyletic and showed Barroetea, Phanerostylis, and Kuhnia all embedded within the genus.

     

    Other articles:
    • Golden Checklist Flora:  Brickellia californica;  

    Brickellia californica (Torrey & A. Gray) A. Gray. “California Brickelbush”

     

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1836.
    - Gray, Asa, 1849.
    - Kunth, Karl Sigismund, 1833-1850.

    Other articles:
    • Golden Checklist Flora:  Gray, 1849, publication details;

    Locations: Mora River.  

    Gray (1849) describes B. californica in his Plantae Fendlerianae Novi-Mexicanae.

    Original Text Comments
    308. B. (Bulbostylis) Californica, Torr. & Gray, Fl. 2. p. 79 : var. foliis plerisque subcordatis. — Rocky hill-side on the Mora River, and eight miles eastward, in bottom land; Aug. (346.) — The cauline leaves are mostly cordate, and with rather longer petioles than in the Californian specimens collected by Douglas ; but the plant of Hartweg's recent collection is wholly intermediate. Mr. Bentham (in Bot. Voy. Sulph.) has very properly carried out the intimation given in the Flora of North America, and annexed Bulbostylis to Brickellia. The pappus is so strongly barbellate-denticulate in some species (as in the original B. cordifolia and especially in B. cylindracea, Gray & Engelm., from Texas, and an undescribed Mexican species found by Dr. Wislizenus *) that Clavigera is separated by a merely arbitary character.† I almost fell off my chair to see that Torrey & Gray applied Bulbostylis DC. to our Brickellia. DeCandolle (1836) did in fact propose it for a group in the Eupatorieae. That name is now considered illegitimate and the name in Cyperaceae, which would appear to be Bulbostylem Kunth, is a conserved name (Kunth, 1833-1850, v. 2, pg. 205).

    The undescribed Mexican species is Brickellia wislizeni A.Gray, an accepted species that occurs only in Mexico.

    The dagger (†) refers to an undescribed Stevia in Coulter's Mexican collection.

     

    Other articles:
    • Golden Checklist Flora:  Brickellia eupatorioides;  

    Brickellia eupatorioides (L.) Shinners. “False Boneset”

     

    Literature Cited:
    - Linne, Carl von, and Lars Salvius, 1763.  

    Linnaeus (1763) publication of Kuhnia eupatorioides from a collection in Pensylvania by Adam Kuhn.

    Original Text
    Eupatorioides. 289. KUHNIA. (Pentandra, Monogyna.)
      Eupatoria conyzoides odorata, folio crenato molli fubincano. Pluk. alm. 140. t. 87. f. 2?
      Habitat in Penfylvania , unde vivam attulit Adam Kuhn. ♃
      Caules fesquioedales, erecti, laeves, rigiduli. Folia alterna, petiolata, lato-lanceolata, nuda, fubrugefa, fubtus venofa, fusdentata ferraturis mediis majoribus. Rami alterni, e medio caule. Corymbi parvi, terminales & etiam fape ramorum lateralium. Corollae albe. Stamina albo-flavefcentia. Planta refert Eupatorium, fed Piftilla clavata, & Antherae diftinctae, cylindricaw, apice labio dehifcentes, absque exemplo in alio Compofits flore.

    Immediately above this entry is one for Ellisia nyctelea.

     

    Other articles:
    • Golden Checklist Flora:  Brickellia grandiflora;  

    Brickellia grandiflora (Hook.) Nutt. “Tasselflower Brickellbush”

     

    Literature Cited:
    - Hooker, Sir William Jackson, 1829-1840.  

    Hooker (1840) described a single spcimen collected by Douglas.

    Original Text
    COMPOSITAE Omissae
    Post Eupatorium occidentale, v. 1. p. 305, adde
    5.* Eupatorium? grandiflorum; glabrum, foliis alternis petiolatis cordato-triangularibus acuminatis grosse serratis venosis, paniculae floribus glomeratis, incolucri foliolis multiserialibus lineari-oblongis acutis striatis exterioribus e lata basi subulatis subsquarrosis, acheniis cylindraceis striatis scabridis.
    Hab. On the low hills betweeb the north and south branches of Lewis and Clarke's River, in stony places. Douglas. — Three to four feet high, herbaceous. Stem rounded. Flowers large, white, clustered. The leaves bear a considerable resemblance to those of E. occidentale, but the flowers are totally different, larger than is usual in this Genus, each of many florets, and with the stigmas less exserted than usual. I possess only a solitary specimen.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.  

    Nuttall (1841) described Brickellia grandiflora from collections on his trip with the Wyeth expedition, citing Eupatorium grandiflorum Hook. as a synonym.

    Original Text
    Brickellia grandiflora, leaves alternate, deltoid-cordate, acuminate, incisely dentate towards the base, entire at the point, smooth on both surfaces, and covered beneath with resinous atoms; flowers in fastigiate clusters, the upper part of the stem branching; inner scales of the involucrum linear-lanceolate, acute; pappus white, achenia smooth. — Eupatorium? grandiflorum. Hook. Flor. Am., Vol. II., p. 26.
    Hab. In the Rocky Mountain range, by streams, in gravelly places, and west, to the lower falls of the Columbia. — Perennial. Stems many from the same root, about twelve to fifteen inches high. The whole plant almost perfectly glabrous. Leaves alternate, sometimes almost opposite, approximate, on longish petioles, deltoid-cordate, acuminate, coarsely and deeply toothed towards the base, smooth and green, but shhining, aith a coating of yellow resinous atoms having a heavy aromatic scent; stem branching above; branches terminating in corymbulose clusters of subsessile flowers, about five capituli in each. Florets straw-yellow, inclining to white, cylindric and smooth, the border connivent. Stigmas exserted, smooth, thicker toward the extremity. Achenium cylindric, ten-striate. Pappus of a single series of twenty to twenty-four scabrous hairs. Receptacle naked, flat.

       

    Centaurea L “Knapweed”

     

    Literature Cited:
    - Hilpold, Andreas, et al., 2014.  

    The virtual absence of congruence between morphology and molecular data suggests that the morphological characters that have been used taxonomically are not reliable with respect to the true relationships.

     

    Other articles:
    • Golden Checklist Flora:  Centaurea diffusa;  

    Centaurea diffusa Lam. “White Knapweed”

     

    Literature Cited:
    - Lamarck, Jean-Baptiste, 1783.  

    Lamarck (1783, t. 1, p. 675-676) ...

    Original Text Translation and Comments
    70. Centauree diffufe , Centaurea diffufa. Centaurea calycibus coliato-fpinulofis exiguis, foliis radicalibus lyratis, caulinis anguftis fubintefris ; caule ramofiffimo paniculato diffufo. N. Carduus orientalis calcitrapae folio, flore monimo. Tourn. Cor. 31. 70. Centauree diffuse Carex diffused. Carex ealycibus coliato-spinulosis small; Radical leaves lyratis, caulinis tightened subintefris; Stem ramosissimo paniculato abroad. X. Eastern calcitrapae thistle leaf, flower Monimum. Tournefort Cor. 31.
    Cette plante reffemble beaucoup à la Centaurée panicuîee no. 36 , & pariot n'en différer efientîellement que par les pointes de fes écailles calîcinales , qui font très-aiguës , roides & un peu épineufes. La tige eft menue , anguleufe , légèrement cotonneule , haute de près d'un pied, extrêmeraent rameufe , panîculée , & diffure, Ses feuilles inférieures font en lyre; toutes les autres font petites , étroites , la plupart fimples , & linéaires. Les fleurs font très-petites , extremement nombreufes , & à écailles calicinales-lancéolces , ciliées , & terminées par une pointe un peu épijieufe. Cette plante croît dans le Levant, (v. f.)   This plant looks a lot like Knapweed panicuîee no. 36, & pariot actually differ than by the points of the calicinal scales, which are very sharp, stiff & a little thorns. The stem is slender, angular, slightly cotton, nearly a foot high, extremely branched, panicle, & diffure, Its lower leaves are lyre-shaped; all the others are small, narrow, mostly fimple, & linear. The flowers are very small, extremely numerous, with calicinal-lanceolate scales, ciliate, & terminated by a somewhat epijieuve point. This plant grows in the Levant. (Seen in the dried state.)

     

    Other articles:
    • Golden Checklist Flora:  Centaurea stoebe;  

    Centaurea stoebe L. “Spotted Knapweed”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 914) ...

    Original Text
    Stœbe. 23 CENTAUREA calycibus ciliatis oblongis, foliis pinnatifidis linearibus integerrimis. Roy. lugdb. 140.

    Stœbe incana, cyano fimilis, tenuifolia. Bauh. pin. 273.

    Stœbe auftriaca humilis. Cluf. hift. 2. p. 10.

    Habitat in Auftria.

    The genus name Stœbe now applies to a mainly south African Gnaphalieae. At the time this was published, Austria included all of Hungary, Transylvania, Galicia, Bohemia, and Lombardy.

    Literature Cited:
    - Lamarck, Jean-Baptiste, 1783.  

    Lamarck (1783, t. 1, p. 669) published C. maculosa ...

    Original Text Translation and Comments
    35. Centauree mouchetee , Centaurea maculofa. Centaurea calycibus ciliatis ovato-fubrotundis pulchre maculofis , foliis tenuibus bipinnatidifis , caule fubpaniculato. N. Centaurea , no. 79 & 80. Gmel. Sib. 2. p. 99. Tab. 44. f. 1. 2.  
    Cette Centauree fe diftingue aifement de la fuivante par fes feuilles decoupees tres-menu , & par fes fleurs , qui font au moins une fois plus groffes , & ont leur calice agreablement mouchete. Sa tige eft haute de huit a dix pouces , ftriee , blanchatre, & un peu paniculee vers fon fommet. Ses feuilles font blanchatres . a decoupures menues & pointues ; les inferieures font oblongues & bipinnatifides ; les fuperieures font plus petites , fimplement pinnees , & a decoupures lineaires. Les fleurs font purpurines & remarquables par leur calice gros, court, mouchete par les taches bbrunes qui font au fomment de fes ecailles. Nous avons obferve cette plante en Auvergne , aux environs de Clermont , fur le puits de Crouel. ( v.v. ) We have obtained this plant in Auvergne, near Clermont, on Puy de Crouel (a small local hill). Auvergne is a historical region in central France, and is now part of Auvergne-Rhône-Alpes. ( I have seen it live. )
     

    135. Ochsmann, Jorg. 2001. On the taxonomy of spotted knapweed (Centaurea stoebe L.). In: Smith, Lincoln, ed. Proceedings, 1st international knapweed symposium of the 21st century; 2001 March 15-16; Coeur d'Alene, ID. Albany, CA: U.S. Department of Agriculture, Agricultural Research Service: 33-41. [37829]

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Treated by Weber & Wittmann (2012) as Acosta maculosa (Lamarck) Holub.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Treated by Ackerfield (2015) as C. stoebe ssp. micranthos (S. G. Gmelin ex Gugler) Hayek.

     

    CABI.org (2021) says this ...

    Original Text
    Most of the North American plants called Centaurea maculosa are perennial, polycarpic and tetraploid (2n = 36). However, there have been multiple introductions, and some infestations might be diploid. Diploids can be identified by genetic analysis and growth habit, because they flower once and die after 2 years (Smith, 2001). Ochsmann (2001b) cites evidence of hybridization between spotted knapweed and diffuse knapweed (Centaurea diffusa) in at least seven US states. The hybrid is named Centaurea ×psammogena.
    I'd like to give credit and copyright information. However, any attempt to find out more about CABI.org fails with a 504 Gateway Time-out. Also it would be really helpful to find a copy of the Ochsmann (2001) paper because many sources of information refer to it.

    Literature Cited:
    - Flora of North America Editorial Committee, eds., 1993+.  

    FNANM (2021) says this ...

    Original Text
    Recent studies have shown that the American plants are identical with plants introduced to the whole of Europe (J. Ochsmann 2001). Subsp. micranthos, a tetraploid perennial, is clearly distinct from the diploid, biennial plants native to central Europe known as C. stoebe Linnaeus subsp. stoebe, C. rhenana Boreau, or C. maculosa Lamarck. In most American literature the name Centaurea maculosa Lamarck has been misapplied to C. stoebe subsp. micranthos. W. A. Weber (1987, 1990) treated this taxon as Acosta maculosa (Lamarck) Holub. The treatment of about 100 species of Centaurea sect. Acrolophus Cassini as the genus Acosta by J. Holub (1972) and others is supported by neither morphologic nor molecular characters and is not widely accepted in Europe.

    Literature Cited:
    - POWO, 2021 - 2023.  

    Plants of the World (2021)does not accept subsp. micranthos (Gugler) Hayek.

     

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Cirsium Mill. Thistle.

    Twenty species of Cirsium “Thistle” are found in Colorado (Ackerfield, 2015). Eighteen of them are native, while two are introduced: C. arvense and C. vulgare. There are two endemic thistles: C. osterhoutii and C. perplexans.

     

    Other articles:
    • Golden Checklist Flora:  Cirsium ochrocentrum;  

    Cirsium ochrocentrum A. Gray “Yellowspine Thistle”

     

     

    Literature Cited:
    - Flora of North America Editorial Committee, eds., 1993+.

    Other articles:
    • Golden Checklist Flora:  Cirsium undulatum;
    Full Size ImageDistribution of collections of Cirsium undulatum  

    Cirsium undulatum (Nutt.) Spreng. “Wavy Leaved Thistle”

    Cirsium undulatum is widely distributed in the wstern half of North America from the dry plains and plateaus of the Pacific Northwest eastward across the Great Plains to Manitoba and the Dakotas and south to Texas, New Mexico, and northwestern Mexico. It occurs in scattered localities in the Rocky Mountains and northeastern Great Basin region. At least some of the few widely scattered records from the eastern United States are probably introductions. Cirsium undulatum is both widespread and variable. Plants of the Great Plains region tend to be low-growing with a few large heads and elongate corollas. Plants of the Pacific Northwest are usually taller and produce smaller, more numerous heads with shorter corollas. A detailed study of this species might reveal races worthy of recognition as infraspecific taxa.

    Wavyleaf thistle is listed by California as a noxious weed. However, most reports of Cirsium undulatum in California are based upon misidentifications of C. canescens. Cirsium undulatum is known to hybridize with C. flodmanii, C. hookerianum, and C. scariosum var. coloradense. J. T. Howell (1960b) reported that C. undulatum was suspected to hybridize with C. brevifolium in the Pacific Northwest.

    — David J. Keil in Flora of North America (vol. 19, 20 and 21)

    Literature Cited:
    - Miller, Philip, 1754.  

    The genus Cirsium was first proposed by Miller (1754) in his Gardener's Dictionary.

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;
    • Glossary:  ventricose;  

    Nuttall (1818) published his Carduus undulatus in his Genera of North American Plants, in the section Cnicus because the pappus is plumose. He saw the plant on Lake Huron and in the upper Louisiana Territory. I have not found a type specimen, but saw a note that the material might be in the Gray Herbarium.

    538. CARDUUS. L. (Thistle.) Calix ventricose, imbricate, scales spiny. Receptacle villous. Pappus pilose or plumose, deciduous.
    § II. CNICUS. Pappus plimose
    11. * undulatus. Stem low and few-flowered; leaves amplexicaule, pinnatifidly sinuate, and plicately undulated, on both sides tomentose, but beneath white, lobes bifid and spiny; calix subglobose, scales lanceolate, eredt and mucronate. HAB. On the calcareous islands of lake Huron, and on the plains of Upper Louisiana. — Leaves almost like some species of Cynara, but not remarkably large, moreslenderly tomentose on the upper side; stem often 1,2, or few-flowered, and 1 to 2 feet high. Flowers large, reddish purple.

    Literature Cited:
    - Sprengel, Curt Polycarp Joachim, 1826.  

    Original Text Translation
    undulatum * 45. C. caule humili paucifloro, foliis amplexicaulibus finuato-pinnatifidis undulato-plicatus utrinque tomentofis, laciniis 2fidis fpinofis, fquamis anthodii lanceolatis erectis mucronatis. In infulis lacus Huronum et Louifiana fuper. (Cnicus Nutt.) undulatum * Cirsium [with] short few-flowered stems, leaves amplexicaul curved-pinnatifid undulate-plicate both sides hairy, lobes divided spiny scales anthodii laneeolatis putting pointed. The islands of Lake Huron and upper Louisiana.

    Literature Cited:
    - Frankton, C., and R. J. Moore, 1961.  

    Frankton and Moore (1961) compare and contrast of Cirsium undulatum and C. flodmanii, both of which are known to occur in Colorado is behind a paywall, so I have reproduced the abstract here.

    Abstract. The morphology and specific differences of Cirsium undulatum (Nutt.) Spreng. And of C. flodmanii (Rydb.) Arthur are described and their Canadian distributions are reported in detail. The chromosome numbers are C. undulatum f. undulatum and f. album Farwell, 2n = 26; C. flodmanii f. flodmanii and f. albiflorum D. Löve, 2n = 22. The origin of four North American species of Cirsium that do not follow the world-wide base number 17 is discussed; it is postulated that reduction in number has occurred by translocations. The chromosomes of species with reduced numbers are larger than those of the unreduced species but the total length of the chromosomes of both groups is approximately the same.

    Literature Cited:
    - Häffner, Eva, and Frank H. Hellwig, 1999.  

    Haffner and Hellwig (1999) investigated relationships within tribe Cardueae using ITS sequence data. The sole North American Cirsium in their study was C. texanum which does not occur in Colorado.

    Literature Cited:
    - Garcia-Jacas, Nuria, Teresa Garnatje, Alfoonso Susanna, and Rosier Vilatersana, 2002.  

    Garcia-Jacas, et al. (2002) worked on tribal and subtribal delimitation of the Cardueae, unfortunately without any North American specimens.

    Literature Cited:
    - Kelch, Dean G., and Bruce G. Baldwin, 2003.  

    Kelch and Baldwin (2003) studied historical biogeography and ecology of true thistles (Cirsium, Cardueae, Compositae) in the New World. Unfortunately, their sample of thistles did not include Cirsium undulatum. However, three other native thistles, C. muticum, C. discolor, and C. rhaphilepis, that commonly cluster with C. undulatum and did in fact cluster together in this study.

    Literature Cited:
    - Bodo Slotto, Tracey A., David P. Horvath, and Michael E. Foley, 2012.  

    Abstract. Weedy invasive Cirsium spp. are widespread in temperate regions of North America and some of their biological control agents have attacked native Cirsium spp. A phylogenetic tree was developed from DNA sequences for the internal transcribed spacer and external transcribed spacer regions from native and non-native Great Plains Cirsium spp. And other thistles to determine if host specificity follows phylogeny. The monophyly of Cirsium spp. And Carduus within the tribe Cardinae was confirmed with native North American and European lineages of the Cirsium spp. Examined. We did not detect interspecific hybridization between the introduced invasive and the native North American Cirsium spp. Selected host-biological control agent interactions were mapped onto the phylogenic tree derived by maximum likelihood analysis to examine the co-occurrence of known hosts with biological control agents. Within Cirsium-Cardueae, the insect biological control agents do not associate with host phylogenetic lines. Thus, more comprehensive testing of species in host-specificity trials, rather than relying on a single representative of a given clade may be necessary; because the assumption that host-specificity follows phylogeny does not necessarily hold. Since the assumption does not always hold, it will also be important to evaluate ecological factors to provide better cues for host specificity.

     

    Other articles:
    • Golden Checklist Flora:  Conyza canadensis;  

    Conyza canadensis L.

     

     

    Other articles:
    • Golden Checklist Flora:  Coreopsis tinctoria;  

    Coreopsis tinctoria Nutt. “Golden Tickseed”

     

    Literature Cited:
    - Nuttall, Thomas, 1821.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1821, publication details;  

    Nuttall (1821, v. 2, p. 114) described a plant he found in the Arkansa Territory and introduced into the gardens of Philadelphia.

    Original Text
    1. COREOPSIS * tinctoria, foliis radicalibus pseudobipinnatis, foliolis subovalibus integris glabris, superioribus pseudopinnatis laciniis linearibus; floribus binatis ternatisve; calcibus exterioribus brevissimis; radiis bicoloribus; seminibus nudis immarginatis.
    Habitat. Throughout the Arkansa territory to the banks of Red River, chiefly in the prairies which are subject to temporary inundation. — Flowering, from June to October.
    Description. Annual and biennial, stem erect, smooth, and much branched, extremely variable in magnitude, being from one to five feet high. The leaves, in common with the genus, are somewhat think and succulent, the primary ones simple, radical pseudobipinnate, the segments also occasionally pinnate, oblong-oval, commonly smooth, and entire, the ultimate divisions largest. Flowers often terminating the branchlets by pairs, with the peduncles unusually short. Exterior calix, minute, much shorter than the interior, and in common with it. And the number of rays mostly eight-leaved. Rays three-lobed at the extremity, of a bright orpiment yellow and brown towards the base; disk brown, and rather small. Receptable paleaceous, the leaflets deciduous. Seed small, blackish, immarginate, curved and naked at the summit.
    Economical Use. The flowers of this species afford a yellow dye, in common with those of the C. senifolia.
    As an ornamental plant, of easy culture and uncommon brilliance, it promises to become the favourite of every garden where it is introduced.

     

    Other articles:
    • Golden Checklist Flora:  Crepis occidentalis;  Nuttall, 1834a, publication details;  

    Crepis occidentalis Nutt. “Largeflower Hawksbeard”

     

    Literature Cited:
    - Nuttall, Thomas, 1834a.  

    Nuttall (1834a, v. 7, p. 29) described Crepis occidentalis from a collection by Nathaniel Wyeth in 1833 between the Falls of the Columbia and the first navigable waters of the Missouri.

    COMPOSITAE
    50. Crepis *Occidentalis. Canescente-pubescens, pumila, foliis sessilibus, runcinato-pinnatisentis, laciniis linearibus, acutis subdenticulatis, floribus paucis fastigiatis.
    ♃ About a span high, covered with a close very short whitish pubescence, The leaves runcinate and acute, about two on the spem, greatly resembling those of the common Shepard's purse ; above, beneath the ultimate flowers, diminishing into simple undivided bracts. Flowers (in the only specimen before me) three, axillary and terminal, all attaining nearly the same height on the stem. Calyx slightly caliculate, the larger leaves of it disposed in a single series, the divisions linear and rather obtuse. Flowers bright yellow, rather large, about the size and appearance of those of Apargia autumnalis ; liguli five-toothed ; the anthers simple, the style bifid, deeply and far exserted. Pappus pilose, somewhat scabrous through a lens, the hairs more than twenty, not dilated at base, or in any way distinguishable from those of Hieracium, nor are they all stipitate. Seed smooth, brown. Probably the Hymenonema laciniatum of Hooker, in Flor. Boreal. Amer. L. c.
    Hab. Common on the borders and in the vicinity of the river Columbia.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;
    • Glossary:  achenium;  testa;  

    In his Descriptions of new Species and Genera of Plants … Nuttall (1841) proposed Psilochenia occidentalis for his previously published Crepis occidentalis Nutt.

    Original Text
    * PSILOCHENIA
    Crepis, but with the achenium cylindric, curved, narrower above, and without any visible striae, the testa indurated, and, when mature, black ; an abortive outer series of florets, with the achenium empty. Pappus copious, slenderly pilose, scabrous, and yellowish white, about the length of the achenium. Receptable naked, alveolate, the alveoles minutely fringed. — A low perennial herb; stem dichotomous and corymbose. Leaves lanceolate, runcinately pinnatifid, and, as well as the somewhat hirsute involucrum, cinereously and closely lanuginous; flowers yellow, rather large.
    Psilochenia * occidentalis. Crepis occidentalis, Nutt. In Journ. Acad. Nat. Sci. Philad., Vol. VII., p. 29.
    Hab. On the plains of the Platte, towards the Rocky Mountains. The whole plant more or less canescently pubescent. Stem about six or seven inches high, forked and corymbose at the summit. Leaves about an inch wide, four or five inches long, deeply and runcinately pinnatifid, the segments linear-lanceolate and denticulate, uppermost leaves linear. Involucrum campanulate; sepals about twelve to fifteen in a single series, linear and somewhat acute; involucel or bractes four or five, small and subulate: there are blackish hairs mixed with the hoary pubescence of the sepals. Florets about twelve, yellow, exserted.

    Literature Cited:
    - Babcock, E. B., and G. Ledyard Stebbins, 1938.  

    Original Text
    TAXONOMIC HISTORY OF THE INDIGENOUS SPECIES
    The first record of the occurrence of Crepis in North America (outside of the arctic regions) was made by Hooker in his “Flora Boreali-Americani” (1834, vol. 1, p. 297), in which he identified as the European C. biennis James's Hieracium runcinatum. In the same year Nuttall published his C. occidentalis, and seven years later (1841), in describing the plants collected on a tour to the Pacific added another species, C. acuminata. In this publication he created a new genus, Psilochenia, for C. occidentalis, and another, Crepidium, for Hieracium runcinatum (Hooker's “C. biennis” of America), and placed Crepis acuminata in a new sub-genus, Leptotheca. Two new species, Crepidium glaucum and C. caulescens, were also described. Two years later Torrey and Gray (1843) reduced Nuttall's two genera to Crepis, and recognized, in addition to the arctic-alpine C. nana Richards., and C. elegans Hook., four species, C. runcinata, C. glauca, C. occidentalis, and C. acuminata. For the next fifty-three years the group received little attention, although a few species and varieties were described by Gray and others. In his “Syntopical Flora of North America” (1876) Gray recognized, in addition to the six species included in his earlier work, two more, C. Andersonii Gray and C. intermedia Gray, and listed two varieties under the latter and three under C.occidentalis. With the heading of the section including C. occidentalis, intermedia, and acuminata he made the terse comment “species difficult,” an opinion with which all later workers on the group, including the present writers, are inclined to agree.
    Coville (1896) was the first to make a monographic study of any of the American species. Omitting the arctic-alpine species and the group of C. runcinata, as well as C. acuminata and C. intermedia, which latter he considered to be not well enough understood, he included seven species, four of which, C. monticola, C. scopulorum, C. rostrata, and C. barbigera, were described as new. His treatment, although well worked out according to the knowledge then available, was based on a relatively meager series of specimens, and included little information on the interrelationships of the species, while their distribution could be stated only in relatively broad terms.
    Since the work of Coville, no particular attention has been paid to the genus in North America, except for the description of numerous new “species,” mostly of the group of C. runcinata, by Greene, Rydberg, and others. These were published without consideration of the group as a whole, were based on fickle characters such as leaf shape and pubescence, and none of them are considered valid by the present writers. Meanwhile in the affinity of C. occidentalis and C. acuminata the species were variously interpreted by the writers of the different floras, the keys to them varied considerably, while the identification of the ever increasing number of collections by various students of the western flora bore out Gray's opinion, “species difficult.”

    Literature Cited:
    - Weber, William A., 1983.  

    Original Text
    The American species of Crepis were treated in a now classic monograph by Babcock & Stebbins (Carnegie Inst. Wash. Publ. 504. 1938). The authors seem to have been preoccupied with the species alone, and unfortunately they did not discuss the significance of their cytological findings as having a bearing on the generic level, even though Nuttall (1841) had proposed the name Psilochenia for the American species.
    All of the native American species of Crepis, with the exception of two Old World species (C. elegans and C. nana), representing an ancient Tertiary extension of the genus onto western North America, have the chromosome base number x=11. “This is in striking contrast to the Old World species of Crepis, whose basic haploid numbers range from x=3 to x=7, 4 and 5 being much the most common" (Babcock & Stebbins, op. cit.). The authors went on to postulate that the American species may have arisen by amphidiploidy from a cross involving Crepis species with x=4 and x=7. Whether or not this can ever be substantiated, the fact remains that the American species of Crepis form an indisputably monophyletic line, spatially and genetically isolated from the Old World species.
    Recently Love (1982, p. 360) transferred Crepis runcinata to Nuttall's genus Psilochenia because of this evidence. but among the rest of the species, only the type, Psilochenia occidentalis Nuttall, has a name in that genus. The following combinations are needed.
    Weber then goes to to propose 23 new combinations in Psilochenia.

    Literature Cited:
    - Enke, Neela, 2009.

    Other articles:
    • Golden Checklist Flora:  Crepis occidentalis, Weber & Wittman, 2012;  

    Enke (2009) is one of several papers by Enke and others exploring the genomics of the genus Crepis. Unfortunately their data includes only one representative of the indigenous North American CrepisC. acuminata — which, I think, resulted in their results being somewhat unclear with regard to that group. Nevertheless, their data supports the maintenance of section Psilochenia within Crepis and does not rule out elevation of the section to the rank of genus as Weber (1983) has proposed.

    Original Text
    Crepis acuminata differs from all other species of clade V in morphology, karyology and geographic distribution. As all members of sect. Psilochaenia it occurs exclusively in North America, is polyploid and has a basic chromosome number of x=11. The singularity of these features within Crepis s.str. Support the maintenance of sect. Psilochaenia.

    Original Text
    Crepis sect. Psilochenia (Nutt.) Babc. (1947) ≡ Psilochenia Nutt. (1841). — Type: C. occidentalis Nutt.

    Note. — The North American species of Crepis sect. Psilochaenia are polyploid and their placement within the genus remains unlcear at present.

    *C. acuminata Nutt.
    ?C. atribarba A. Heller
    ?C. bakeri Greene
    ?C. barbigera Coville
    ?C. intermedia A. Gray
    ?C. modocensis Greene
    ?C. monticola Coville
    ?C. occidentalis Nutt.
    ?C. pleurocarpa A. Gray
    ?C. runcinata (E. James) Torr. & A. Gray

    Literature Cited:
    - Babcock, E. B., and G. Ledyard Stebbins, 1938.
    - Weber, William A., 1983.
    - Weber, William A., and Ronald C. Wittmann, 2012.

    Other articles:
    • Golden Checklist Flora:  Crepis occidentalis, Enke, 2009;  Solidago nana, Nuttall, 1841;  

    Weber & Wittmann (2012) place the indigenous North American Crepis in Psilochenia also spelled Psilochaenia saying,

    Original Text
    The genus Psilochenia encompasses the indigenous North American species of Crepis (with the exception of the Old World taxa C. elegans and C. nana, which represent a Tertiary extension). The chromosome base number x=11 is unknown in Eruasia. See Babcock & Stebbins 1938, Weber 1983. The FNA ignores Psilochenia altogether.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield retains our Largeflower Hawksbeard in Crepis as C. occidentalis Nutt.

     

    Other articles:
    • Golden Checklist Flora:  Cyclachaena xanthifolia;  

    Cyclachaena xanthifolia (Nutt.) Fresen. “Carelessweed”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Nuttall (1818, v. 2, p. 185) described his Iva xanthiifolia from a collection made near Fort Mandan. Fort Mandan was the name of the encampment which the Lewis and Clark Expedition built for wintering over in 1804-1805. The encampment was located on now-private land on the Missouri River approximately twelve miles from the site of present-day Washburn, North Dakota, which developed later. The exact location is unknown and may be partially submerged by the river.

    602. IVA. L.
    Calix about 5-leaved, or 5-parted. Feminine florets of the ray 5, naked. Receptacle setosely paleaceous. Seed obovate, naked.
    Herbaceous or shrubby; leaves 3-nerved, mostly carneous, opposite and alternate; flowers spiked or paniculated, axillar and terminal.
    Species. 1. ciliata. ... 2. * xanthiifolia. Annual; leaves opposite, petiolate, cordate-ovate, acuminated, douply serrate, softly villous, beneath canescent; spikes paniculated, naked; calix 5-cleft. Hab. In arid soils, near Fort Mandan, &c. on the banks of the Missouri. Obs. Plant very large, 5 or 6 feet high, with leaves nearly of the size and form of Xanthium Strumarium, but covered with a soft and almost velvet-like villus; upper leaves ovate; flowers extremely numerous, in a diffuse panicle. Calix 5-cleft, divisions ovate-lanceolate, acuminate. Style of the discal florets simple; stigma subcapitate. Fertile florets 5, naked. Receptacle subsetaceous. — Flowering in August.
    3. imbricata. ... 4. axillaris. ... &c.
    A North American genus with the exception of I. annua indigenous to the tropical regions of the same continent.

    There is a collection of Iva xanthiifolia attributed to Nuttall with a location of Upper Louisiana at the Harvard University Herbaria, though there is no image and the type status is not indicated in the data record.

    Literature Cited:
    - Fresenius, Georg, 1836.

    Other articles:
    • Golden Checklist Flora:  Fresenius, 1836, publication details;
    Full Size ImageFootnotes from Index Seminum in which Cyclachaena is proposed  

    Georg Fresenius is the apparent author of Index Seminum. Frankfort am Main, a list of seeds available from the Senckenberg Herbarium in Frankfort am Main. In the 1836 Index, seeds are available for Cyclachaena xanthiifolia Fresen. Footnote 4 in that document proposes Cyclachaena as a new genus. However, there is no reference to Iva xanthiifolia Nutt. As the basionym.

    Although no basionym reference was given Art. 41.4 (Melbourne Code) applies,

    41.4. If, for a name of a genus or taxon of lower rank published before 1 January 1953, no reference to a basionym is given but the conditions for its valid publication as the name of a new taxon or replacement name are fulfilled, that name is nevertheless treated as a new combination or name at new rank when this was the author’s presumed intent and a potential basionym (Art. 6.10) applying to the same taxon exists.

    — International Code of Nomenclature for algae, fungi, and plants, 2012 — Melbourne Code text: © 2012, IAPT — web-edition: © 2014, Paul van Rijckevorsel (all rights reserved)

    Basionym Iva xanthiifolia Nutt., Gen. N. Amer. Pl. [Nuttall]. 2: 185 (1818).

    Full Size Image
    Page 1 from Index Seminum Frankfort/Am Main

    Literature Cited:
    - Miao, Bomao, Billie L. Turner, and Tom Mabry, 1995.  

    Results support the dismemberment of Iva s.l. and recognition of Iva s. str., largely because Iva s. l. is shown to be paraphyletic. Most members of the section Cyclachaena were found to have relatively close interspecific relationships. Nevertheless cpDNA data strongly support two lineages within Cyclachaena. One lineage, including I. xanthiifolia, I. acerosa, I. nevadensis, and I. dealbata, has a close relationship with genus Euphrosyne.

    Literature Cited:
    - Flann, C (ed), 2009+.  

    The Global Composite Checklist treats Iva xanthiifolia as an accepted name, and Cyclachaena xanthiifolia (Nutt.) Fresen. as a synonym. Plants of the World (Kew) does not follow their lead and accepts Cyclachaena xanthiifolia (Nutt.) Fresen..

       

    Dieteria Nutt.

    Sometimes placed in Aster, Dieteria, or Macheranthera.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.  

    Nuttall(1840, p. 300) in describing his plants from trip to the Oregon Territory, described a new genus for a subset of Asters.

    Original Text
    * DIETERIA.
    Flower radiate, rays styliferous, fertile ? liguli one or two series, boradish, those of the disk hermaphrodite, fertile. Stigma filiform, hirsute and exserted. Sepals of the involucrum, for the most part, closely imbricated in two to four series, scariose and carinate, the tips usually reflected and herbaceous. Receptacle flat or convex, alveolate, the alveolæ deep, with toothed and lacerated margins. Achenium obovate, subcylindric, ten to fifteen striate, pubescent. Pappus of several series, scabrous and unequal, that of the ray shorter and less copious. — Annual or biennial, (in one anomalous species perennial,) divaricately branching herbs, more or less pubescent; leaves nearly entire, incisely serrate or pinnatifid, the points often pungently mucronulate. Flowers fastigiate. The disk yellow. Liguli red or purple. — Allied to Aster, but with the involucrum regular; the achenia convex, distinctly striate when ripe; the receptacle deeply alveolate; the pappus of the ray different from that of the disk; the leaves incise or pinnatifid, and the duration only to the first period of flowering. They are also allied to te first section of Heterotheca by the deficient pappus of the ray, but that of the disk is simple, and the rays are purple. The while olant bitter to the taste. — (So called from their biennial duration.)

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.  

    Dieteria Nuttall

    Nuttall (1841, p. 300) proposed a new genus Dieteria without identifying a type, the concept of which was lacking at the time. I suspect that D. canescens (syn: Aster canescens Pursh) is probably the closest thing we have to a type.

    *DIETERIA.
    Flower radiate, rays styliferous, fertile? liguli one or two series, broadish, those of the disk hermaphrodite, fertile. Stigma filiform, hirsute and exserted. Sepals of the involucrum, for the most part, closely imbricated in two to four series, scariose and carinate, the tips usually reflected and herbaceous. Receptacle flat or convex, alveolate, the alveolae deep, with toothed and lacerated margins. Achenium obovate, subcylindric, ten to fifteen striate, pubescent. Pappus of several series, scabrous and unequal, that of the ray shorter and less copious. — Annual or biennial, (in one anomalous species perennial,) divaricately branching herbs, more or less pubescent; leaves nearly entire, incisely serrate or pinnatifid, the points often pungently mucronulate. Flowers fastigiate. The disk yellow. Liguli red or purple. — Allied to Aster, but with the involucrum regular; the achenia convex, distinctly striate when ripe ; the receptacle deeply alveolate ; the pappus of the ray different from that of the disk; the leaves incise or pinnatifid, and the duration only to the first period of flowering. They are also allied to the first section of Heterotheca by the deficient pappus of the ray, but that of the disk is simple, and the rays are purple. The whole plant bitter to the taste. — (So called from their biennial duration.)

     

    Other articles:
    • Golden Checklist Flora:  Dieteria bigelovii;

    Locations: Sandia Mountains.  

    Dieteria bigelovii (A. Gray) D. R. Morgan & R. L. Hartman “Bigelow's Tansy Aster”

     

    Literature Cited:
    - Torrey, John, 1857.  

    Gray, in Torrey (1857, p. [97] 41)

    Original Text
    Aster Bigelovii (sp. nov,) : ramis viscido-hirsutis ad apicem usque foiiosis; ramulis corymbosis monocephalis ; foliis membranaceis oblongo-lanceolatis semiamplexicaulibus grosseserratis fenuiter triplinerviis hirto-puberulis glabratis ; capitulis magnis globosis ; involucri pluriserialis squamis attenuato-subulatis basi appressis superne longe caudato-appendiculatis squarroso-recurvis glanduloso-viscidis ; acheniis glaberrimis. Arroyos in the Sandia mountains ; October. A wholly new and most remarkable Aster, of the Grandiflori group ; but the apparently showy heads larger than those of A. grandiflorus, being an inch in diameter, and the numerous (blue and violet) rays an inch long. It is probably a tall plant ; but the base of the stem was not collected. Cauline leaves two or three inches long, coarsely dentate-serrate throughout ; the uppermost, and those of the short branchlets, smaller and less toothed. Scales of the imbricated involucre half an inch long when extended, very slender ; the long and almost filiform appendicular portion recurved, spreading and very glandular. Receptacle flat, alveolate ; the alveolae short and entire. Achenia perfectly glabrous, linear, compressed, three lines long. Pappus not abundant, nearly in a single series.

     

    Other articles:
    • Golden Checklist Flora:  Dieteria canescens;  

    Dieteria canescens (Pursh) Nutt. “Hoary Tansyaster”

     

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2., p. 547) ... Aster canescens ...

    Original Text
    636. ASTER. Gen. pl. 1291.  
    * Foliis integerrimis.  
    17. A. cano-pubescens ; foliis linearibus, panicula corymbosa ramisissima foliosa, calycibus imbricatis acutissimus disco longioribus. canescens.  
    On the banks of the Missouri. ♄. Aug.-Nov. v. s. Flowers the size of a daisy, rays pale purple.   Seen by Pursh in the dried state, but neither the collector nor the herbarium is identified. Probably was not Lewis & Clark, because their single specimen was collected on the Columbia River and first thought to be Aplopappus sp. More likely a Nuttall collection, though Bradbury cannot be ruled out.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.  

    Nuttall (1840, p. 300) published Dieteria and placed Aster canescens Pursh in it.

    Original Text
    Involucrum subovate, of three or four series of scales.
    Dieteria canescens; leaves entire, linear, sessile, radical spathulate; stem low and much branched, canescently villous, as well as the involucrum; flowers fastigiate; rays about eighteen to twenty; pappus very slender.
    Hab. On the denuded banks of the Missouri. Aster canescens, Pursh, Bor. Am., Vol. II., p. 547. Not in the least allied to Aster multiflorus. A. biennis, Nutt. Gen. Am., Vol.II., p. 155. I doubt if the leaves are always entire, a fact so contrary to all the rest of the genus to which it is, in all other respects, so intimately allied.

     

    Other articles:
    • Golden Checklist Flora:  Dyssodia papposa;  

    Dyssodia papposa (Vent.) Hitchc. “Fetid Marigold”

     

    Literature Cited:
    - Ventenat, Etienne P., Jacques M. Cels, and Henri J. Redoute, 1801.  

    Jacques Philippe Martin Cels (1740–1806) was a French botanist specializing in horticulture. He started a botanical garden in which he cultivated foreign plants for sale, contributing to the growing public appetite for exotic flowers. He received and acclimatized numerous North American plants brought back by André Michaux and Louis-Augustin Bosc d'Antic. He strove to introduce many exotic species into France. The species in his garden were described by the botanist Étienne Pierre Ventenat (1757–1808) and illustrated by Pierre-Joseph Redouté (1759–1840) in Description des plantes nouvelles et peu connues, cultivées dans le jardin de J.-M. Cels, published in Paris in 1799 (WikiPedia, 2021).

    Ventenat's (1801) description of Tagetes papposa states that it was discovered by A. Michaux in the countryside of Illinois.

    Literature Cited:
    - Hitchcock, A. S., 1891.  

    Hitchcock (1891, p. 503) moved our plant to Dyssodia papposa in a catalogue of plants in Ames, Iowa.

    Original Text

    Dysodia papposa, (Vent.) — Tagetes papposa, Vent. Hort. Cels. 1800. — D. chrysanthemoides, Lag. Nov. Gen. & Spec. 1816.

    Sterile soil; frequent.

    The pappus consists of “scales dissected into britles” instead of “capillary bristles,” as it may at first appear.

     

    Other articles:
    • How did rubber rabbitbrush get that long scientific name?:  Introduction;  

    Ericameria nauseosa (Pall. ex Pursh) G.L.Nesom & G.I.Baird var. graveolens (Nutt.) Reveal & Schuyler

    • How did we get to the name of Ericameria nauseosa var. graveolens?
      • What is the history of the genus name Chrysothamnus?
      • What is the history of the name Ericameria nauseosa?
      • What is the history of the name ____ graveolens? And how did it become a variety of Ericameria nauseosa?

    See my page about Ericameria nauseosa var. graveolens or “How did rubber rabbitbrush get that long scientific name?”

     

    Other articles:
    • Golden Checklist Flora:  Ericameria nauseosa nauseosa;
    Full Size ImageHabitat of Coll. No. 2440, Ericameria nauseosa var. nauseosa
    Full Size ImageColl. No. 2440, Ericameria nauseosa var. nauseosa  

    Ericameria nauseosa (Pall. ex Pursh) G. I. Nesom & G. I. Baird var. nauseosa. “Rubber Rabbitbush”

     

     

    Literature Cited:
    - Nesom, Guy L., 2008.  

    Erigeron L. “Fleabane; Daisy”

    The Erigerons that are known from collections in Golden s.l. are shown below, and are placed in taxonomic groups of Nesom (2008).

    • 20. Sect. Olygotrichum Nutt., Trans Amer. Philos. Soc. ser. 2, 7: 311. 1841. Type: Erigeron divergens Torr. & A. Gray

      • Erigeron divergens Torr. & A. Gray. Spreading Fleabane.
      • Erigeron flagellaris A. Gray. Trailing Fleabane.
      • Erigeron tracyi Greene. (Syn: Erigeron cinereus A. Gray, Erigeron colomexicanus A. Nelson, Erigeron divergens Torrey & A. Gray var. cinereus A. Gray) Running Fleabane.

    • 23. Sect. Phalacroloma (Cass.) Torr. & A. Gray, Fl. N. Amer. 2(1): 175. 1841. Phalacroloma Cass., Dict. Sci. Nat. 39: 404. 1826. Type: Erigeron strigosus Muhlenb. ex Willd.

      • Erigeron strigosus Muhl. ex Willd. (Syn: Stenactis strigosa (Muhl. ex Willd.) DC.) Prairie Fleabane.

    • 31. Sect. Stenactis Torr. & A. Gray, Fl. N. Amer. 2(1): 172. 1841. Lectotype: (Nesom 1989c): Erigeron pumilus Nutt.

      • Erigeron pumilus Nutt. Shaggy Fleabane.
      • Erigeron vetensis Rydb. Early Bluetop Fleabane.

    • 33. Sect. Tridactylia Nutt., Trans. Amer. Philos. Soc. ser. 2, 7: 310. 1841. Type: Erigeron compositus Pursh.

      • Erigeron compositus Pursh. (Syn: Erigeron compositus Pursh var. discoideus A. Gray, Erigeron compositus Pursh var. glabratus Macoun) Cutleaf Daisy.

     

    Other articles:
    • Golden Checklist Flora:  Erigeron compositus;  

    Erigeron compositus Pursh “Cutleaf Daisy”

    There are two collections of “Cutleaf Daisy” — Erigeron compositus Pursh — made on the higher edges of Golden s.l. of Lookout Mountain. This is consistent with other collections made in Jefferson County, and throughout Colorado, with a few collections made on the higher plains, such as the Palmer Divide, but generally in the foothills and higher mountain ranges.

    The species was described by Pursh (1814) from Lewis & Clark collections on the Kooskoosky (Clearwater) River.

    Literature Cited:
    - Moulton, Gary E., 1999.
    - Pursh, Frederick, 1814.

    Locations: Kooskoosky River.  

    Of the western Erigerons found in Golden s.l. the first to be described was E. compositus Pursh. The holotype was collected by Meriwether Lewis on the Kooskoosky (Clearwater) River, date unknown, and the voucher is now at ANS.

    Pursh (1814) described it as follows:

    Syngenesia Superflua. Erigeron 535
    14. E. pilosum, subacaule; foliis radicalibus longe petiolatis triplicato-3-partitis, laciniis linearibus divaricatis, caulinis linearibus plerumque indivisibus, caule superne nudo unifloro. compositum
      On the banks of the Kooskoosky. M. Lewis. ♃. July, Aug. v. s.; v. v. cultum. Not above a span high; flowers resembling a daisy exceedingly; they change during their flowering, from white to a lively pale red.  

    My collection of this taxon was made as Little Scraggy Peak, (Buffalo Creek Recreation Area), Jefferson County, Colorado.

    Full Size Image
    Coll. No. 1901, Erigeron compositus
     

    A. Gray (1862) proposed variety discoideus. The name is now treated as a synonym of the species.

     

    Macoun (1884) proposed variety glabratus, but the name is not now recognized. The name is now treated as a synonym of the species.

     

    Other articles:
    • Golden Checklist Flora:  Erigeron divergens;  

    Erigeron divergens Torr. & A. Gray. Spreading Fleabane.

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Glossary:   fastigiate;
    Full Size ImageColl. No. 2092, Erigeron divergens  

    The first publication of the entity we now call Erigeron divergens was by Nuttall (1840).

    Erigeron *divaricatum; ☉ hirsute, stem branching from base; branchlets one-flowered, rather naked, fastigiate; radical leaves spathulate, the rest linear, sessile, acute, attenuated below, all entire; inner pappus of about eight setæ rays nery numerous, narrow, white.
    Hab. In the Rocky Mountains and the plains of Oregon. About one foot high, at length very much branched, the leaves an inch or more long, about a line wide. Pappus double in ray and disk, the inner of remarkably few rays, very deciduous.

    It might be helpful to remember that the plains of Oregon would include a good part of southwest Wyoming.

    The name E. divericatum had been previously used by Michaux (1803) and was not available for use by Nuttall.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1838-1843.

    Other articles:
    • Field Notes:  Torrey & Gray, 1841;  

    Torrey & Gray (1841, p. 175) in Flora of North America proposed Erigeron divergens for the small fleabane that was previously illegimately published by Nuttall as E. divaricatuum.

    26. E. divergens: somewhat hoary with a minute hirsute pubescence, diffusely branched from the base; leaves small, entire, acute; the radical somewhat spatulate, narrowed into a short petiole; the cauline scattered, sessile, linear, narrowed at the base; heads (small) mostly solitary terminating the naked branchlets or pecuncles; rays very narrow and numerous, twice the length of the hirsute involucre; inner pappus of few (8-12) very slender and deciduous bristles. — Erigeron (Oligotrichum) divaricatum, Nutt. ! In trans. Amer. Phil. Soc. (n. ser.) 7. p. 311, not of Michx.
    Rocky Mountains, and plains of the interior of Oregon, Nuttall ! — ① Stems about a foot high, diffuse, ascending, slender, at length much branched. Leaves half an inch to an inch long, 1 to 2 lines wide, mucronate-acute. Heads rather smaller than in E. tenue; the rays (white, Nutt.) nearly similar; the exterior pappus shorter.

     

    Other articles:
    • Golden Checklist Flora:  Erigeron flagellaris;  

    Erigeron flagellaris A. Gray. Trailing Fleabane.

     

    Literature Cited:
    - Gray, Asa, 1849.

    Other articles:
    • Field Notes:  Gray, A., 1849;

    Locations: Santa Fe River.  

    Gray (1849, p. 68) described E. flagellare from a Fendler collection along Santa Fe Creek.

    334. E. flagellare (sp. nov.): bienne? striguloso-puberulum, pumilum; caulibus gracillimis e basi ramosis, floriferis seu primariis simplicibus superne aphyllis monocephalis, sterilibus patentibus flagelliformibus; foliis spathulatis mucronulatis inferioribus in petiolum attenuatis integris seu radicalibus parce inciso-lobatus, ramealibus parvulis sublinearibus sessilibus; ligulis numerosis gracilibus (albis purpureo tinctis) involucrum hirsutum duplo superantibus; pappo radii et discii conformiduplici, exteriore coroni-forme-squamellato conspicuo, interiore e setis sub-20 fragilibus. — Low, moist places, along Santa Fe Creek; May, June. (381.) — Root slender. Flowering stems 5 to 7 inches high, very slender, few-leaved below the middle, naked, and pedunculiform above; the head rather than the preceding species; the involucre, &c., similar. Lower leaves one to two inches long, including the slender petiole; those of the runner-like sterile branches decreasing to 2 or 3 lines in length. This species should rank next to the foregoing.*

    The “preceding species” and “foregoing” was Erigeron cinereum (sp. nov.), now treated as a synonym of E. tracyi Greene.

    The head was described as “… as large as those on Bellis perennis …” and the involucre was described as “… hirsutum duplo superantibus …” [… coarse erect or ascending hairs doubly … ??? ]

    The asterisk “*” refers to some collections sent by Mr. Spaulding from Oregon, which I don't think are particularly relevant to E. flagellaris.

    Other articles:
    • Plainview Road:   near coll loc;
    • Field Notes:  Coll. No. 1116, 28 May 2015;   Coll. No. 1829, 16 May 2018;
    Full Size ImageColl. No. 1116, Erigeron flagellaris
    Full Size ImageColl. No. 1829, Erigeron flagellaris  

     

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Erigeron pumilus Nutt. Shaggy Fleabane.

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Nuttall (1818) described E. pumilum found on the plains of the Missouri.

    Original Text Comments
    2. *pumilum. Hirsute; stems aggregated, 1-flowered, leaves obling-linear, entire and sessile; flower large and hemispherical, before flowering nutant; calix very hirsute. Hab. On the plains of the Missouri. Flowering in May. E. hisutum. Ph. 2. Suppl. p. 742. but this name has been previously employed for another species. Obs. Stems several from the same root, often, indeed, connected at the base, 4 or 5 inches high, 1-flowered. Leaves 2 inches, more or less, nearly linear, more or less, nearly linear, attenuated downwards, scarcely 2 lines wide. Flower naked, (or pedunculate) white, and large as a Daisy; rays as long as the calix, narrow and numerous. Pappus double, internal short, about 12-rayed.  

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Erigeron pumilis;  Nuttall, 1818, publication details;
    • Glossary:  nutant;  

    Syngenesia. Superflua. 147
    557. ERIGERON. L. (Flea-bane.)
    Calix imbricated, subhemispherical, in fruit often reflected. Rays of the corolla linear, very narrow, and numerous. Receptacle naked. Pappus double, exterior minute, interior pilose, of few rays. (12 to 25?)
    Herbaceous; stems simple or branched, flowers solitary, axillary and terminal, subcorymbose or fastigiately paniculate; radii white, or purplish, rarely destiture of pappus. Seed very small, rather smooth, oblong, and compressed; pappus simply pilose, deciduous, not much longer than the seed, exterior pappus minute and paleaceous.
    § I. Stem simple
    SPECIES. 1. E. alpinum. 2. * pumilum. Hirsute; stems aggregated, 1-flowered, leaves oblong-linear, entire and sessile; flower large and hemispherical, before flowering nutant; calyx very hirsute. HAB. On the plains of the Missouri. Flowering in May. E. hirsutum. Ph. 2. Suppl. p. 742. but this name has been previously employed for another species. OBS. Stems several from the same root, often, indeed, connected at the base, 4 or 5 inches high, 1-flowered. Leaves 2 inches, more or less, nearly linear, attenuated downwards, scarcely 2 lines wide. Flower naked, (or pedunculate) white, and large as a Daisy; rays as long as the calix, narrow and numerous. Pappus double, internal short, about 12-rayed.
    “nutant”, adj., drooping or nodding.

     

    Other articles:
    • Golden Checklist Flora:  Erigeron strigosus;  

    Erigeron strigosus Muhl. ex Willd. Prairie Fleabane.

     

    Literature Cited:
    - Willdenow, Carl L., 1797-1830.  

    The first of the Erigeron found in the Golden, Colorado area was described from a collection made in Pensylvania. Willdenow (1803) published the name from a description written by Mühlenberg.

    11. ERIGERON ʃtrigoʃum.
    E. foliis lanceolatis utrinque attenuatis ſubdentatis ſtrigoſo-piloſis, floribus corymboſo-paniculatis. W.
    Erigeron ſtrigoſum. Mühlenb. In litt.
    Behaartes Berufungskraut. W.
    Habitat in Penſylvania. (v. ʃ.)
    Caulis erectus ʃtriatus, pilis ʃparʃis albis obʃitus. Folia alterna lanceolata baʃ et apice attenuata, utrinque oilis copioʃis adpresʃis obʃita, integerrima, vel medio utrunque ʃerraturis 2. ʃ. 3 inʃtructa. Flores corymboʃo-paniculati. Radius copioʃus filiformis albus. W.

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1836.  

    DeCandolle (1836) proposed Stenactis strigosa DC.

    5. S. strigosa, caule erecto sparse et patentim piloso, foliis lanceolatis utrinque stteniatis dentatis strigoso-hispodid, capitulis laxe corymboso-paniculatis, invol. Glaberrimo, ligulis numerosis filiformibus. ② ? In Pensylvania (Muhl.), Noveboraco (Torrey!). Erigeron strigosum Muhl. In Willd. Sp. 3. p. 1956. Erig. Australe Horn. Ex Spreng. An Erig. Strigosun Ell. Sketch 2. p. 394 et Doronicum ramosum Walt. Fl. Car. 206 in Carolina cresc. Ad hanc speciem referenda? (v. s. comm. A cl. Torr.)

    Literature Cited:
    - Nesom, Guy L., 2008.  

    Stenactis is currently treated as a section of Erigeron (Nesom, 2008).

     

    Other articles:
    • Golden Checklist Flora:  Erigeron tracyi;  

    Erigeron tracyi Greene. Running Fleabane.

     
     

    Some of the current synonyms for E. tracyi are:

    • Erigeron cinereus A. Gray
    • Erigeron cinereus var. cinereus
    • Erigeron cinereus var. aridus M. E. Jones
    • Erigeron colomexicanus A. Nelson
    • Erigeron commixtus Greene
    • Erigeron dicladus Greene
    • Erigeron divergens Torrey & A. Gray var. cinereus A. Gray
    • Erigeron tephroides Greene
    • Erigeron williamsii Phil. The Plant List and the Global Composite Checklist each give E. williamsii Phil as a synonym of E. tracyi. IPNI.org does not. I suspect that E. williamsii is a valid name for a South American species, and does not apply to E. tracyi.

    Literature Cited:
    - Gray, Asa, 1849.

    Other articles:
    • Golden Checklist Flora:  Gray, 1849, publication details;  

    The first known description of the taxon we now call E. tracyi was by A. Gray (1849) in Plantae Fendlerianae Novi-Mexicanae, a description of plants collected by Augustus Fendler in New Mexico in 1841

    333. E. cinereum (sp. Nov.): bienne? Undique molliter cinereo-pilosum: caule e basi ramoso; ramis adsurgentibus apice longe nudis monocephalis; foliis spathulatis vel lineari-oblongis basi attenuatis integerrimis seu radicalibus paucidentatis incisisve; ligulus numerosissimis gracilibus (albis nunc purpureo tinctis) involucrum hirsutum duplo superantibus; pappo radii et disci conformi duplici, exteriore coroniformi-squamellato, interiore e setis sub-20 fragilibus deciduis. — Var. a. Is a dwarf, vernal form, only a span high, quite hoary, the primary flowering stems erect and almost scapiform (no. 374 of the distribution). Dry, exposed places around Santa Fe; May. Var. ß has taller and more diffuse stems (10 inches high), the leaves almost lanceolate, entire, the lower tapering into slender petioles. Low, sandy banks of the Rio del Norte and of Santa Fe Creek; May to June. (380.) Var. ? is a larger, coarser, and much more leafy state; from the valley of Santa Fe Creek, near irrigating ditches; May to July. (385.) — The heads are as large as those of Bellis perennis, solitary on peduncles, or the naked summit of the stems, of from 2 to 4 inches in length. The species belongs to the first division of the section Phalacroloma, Torr. & Gray, l. c. † Some forms of this, or of an allied species (possibly E. affine, DC.), with rather less numerous and white rays, and either entire or incised leaves, were gathered at Buena Vista and Encantada by Dr. Gregg.

    Unfortunately, the name was unavailable, having been previously used by Hooker and Arnold for their E. cinereus, a native of Chile.

    Literature Cited:
    - Gray, Asa, and Charles Wright, 1852.  

    Gray (1852, pt. 1, p. 91) places his E. cinereum as a variety of E. divergens in Plantae Wrightinae. He make no comments that describe his thinking.

    268. E. divergens, var. cinereum. E. cinereum, Gray, Pl. Fendl. p. 68. New Mexico; the locality not recorded.

    Gray (1852, pt. 2 p. 77) also places his E. cinereum as a variety of E. divergens in Plantae Wrightinae, part 2.

    E. divergens, var. cinereum, Gray, Pl. Wright, l. c. Moutains, around the copper mines; Oct: a late, much-branched state. (1169) Hills near El Paso; March, April: the early normal form (1398.)

    This placement is still accepted by Harrington one hundred years later (Harrington, 1954, 1964 2nd ed.).

    Literature Cited:
    - Buckley, S. B., 1861.  

    These plants were collected by the author while engaged in the State Geological Survey of Texas, during 1860 and '61. Specimens of them are in the herbarium of the Academy of Natural Sciences at Philadelphia, and also in the herbarium of Elias Durand, Esq.

    Erigeron (Eurigera) nudiflorum, s. n. — Hirsutum pumilum, ramosum; foliis lineari-oblongis, integris, sentis, confertissimis; ramis apice longe nudis, monocephalis; ligulis albis, plurimis, subuniseriatis, involucrum duplo excidentibus, acheniis oblongis glabris vel parum pilosis; pappo radii et disci conformi, duplici; exteriore breve setacea.

    Northern Texas. May.
    Very much branched from the root. Stems erect, 4-6 inches high; leaves numerous near the root and upwards on the stems about 3 inches; the upper stems, two inches below the flowers, naked; whole plant strigose hirsute, the white hairs not appressed; flowers about 1/4 of an inch in diameter; leaves 1/2 - 1 inch long and a little more than a line wide.

     

    Literature Cited:
    - Greene, Edward Lee, 1902a.

    Other articles:
    • Field Notes:   Greene, E. L., 1902;  

    E. L. Greene (1902) then published two names for what we now treat as a single species. The names he published were: Erigeron commixtus and E. tracyi.

    Erigeron commixtus. With the habit of E. flagellaris, smaller, less stoloniferous, the leaves relativelt broader, some entire, others with one or more conspicuous lobes at base of the blade, the whole plant almost hoary with stiffly hirsute pubescence, this spreading on the leaves and petioles, retrorse on the stems and peduncles: heads, rays, achenes, etc., much as in E. flagellaris.
    Cañon of the Limpia, mountains of western Texas, 26 April 1902, S. M. Tracy and F. S. Earle; also collected by the present writer, in the same general region, namely in the mountains near Silver City, New Mexico, 18 May 1880, and distributed for E. flagellaris. The species last named has a rather obscure, fine closely appressed hairiness. That of the new one is so extremely different, that were the plants the size of a Sunflower or Goldenrod, no botanist would confuse them as one species, were the pubescence the only character. I may remark that true E. flagellaris reaches the mountains of even southern New Mexico, where, however, it occurs only in a more elevated biological zone

    Literature Cited:
    - Greene, Edward Lee, 1902a.
    - Nesom, Guy L., 2004.  

    Erigeron Tracyi. Allied to the last (Erigeron commixtus — Ed.) but dwarf, only 2 or 3 inches high, densely leafy at base and with no stolons (at least at flowering time): petioles of the spatulate-lanceolate entire leaves shorter than the blade or obsolete; the whole herbage silvery-hoary with a fine dense strigulose pubescence, or this more sparse and spreading on the solitary scapiform peduncle: involucre hispidulous: outer pappus very conspicuous though consisting of only shorter and setiform hairs, the inner of a few very delicate capillary ones.
    Davis Mountains, western Texas, Tracy and Earle, 28 April 1902.

    E. commixtus was published before E. tracyi, quite literally a matter of a few inches. It would therefore seem to have priority over E. tracyi.

    Samuel Mills Tracy (1847-1920) was the first Director of the Mississippi Experiment Station. Tracy is well known for his work in the taxonomy of various grasses, plant breeds, and for adapting numerous forage plants to the biomes of the Southern United States. He is perhaps best known for his work, Flora of Missouri, which was published in 1886.

    Franklin Sumner Earle (1856-1929), served as the superintendent of the Mississippi Agriculture Experiment Station (1892–1895). Soon after that Earle worked as a biologist and horticulturist of the Alabama Agriculture Experiment Station (1895–1900).

    Tracy and Earle to not appear to be part of any particular expedition. Perhaps, they were just two old botanists out for some fun field work.

    Literature Cited:
    - Coulter, John M., and Aven Nelson, 1909.

    Other articles:
    • Glossary:  cinereous;  

    33. Erigeron colo-mexicanus A. Nels. Stems few to several from s slender annual taproot, ascending, naked pedunculiform above the middle, very leafy, flowering when very short, the later heads on stems 7-15 cm. long: leaves mostly linear-spatulate to linear, the basal sometimes oblanceolate and 3-lobed at apex, all softly cinereous-pubescent: heads musc as in E. divergens. (E cinereus Gray, Pl. Fendl. 68. 1848; not E. cinereus H. & A. Comp. Bot. Mag. 2: 50. 1836) — Southern Colorado and New Mexico.

    It is interesting to me that A. Nelson does not mention Greene's (1902) publication of Erigeron tracyi, whereas other names of Greene are acknowledged.

    Literature Cited:
    - Rydberg, Per Axel, 1910.  

    Erigeron commixtus Greene, Pittonia 5: 58. 1902
    E. cinereus A. Gray, Mem. Am. Acad. 4: 68. 1849. Not E.cinereus H. & A. 1836.
    E. colo-mexicanus A. Nels,; Coult. & Nels. New Man. Cent. Rocky Mts. 529. 1909.
    Another specific name proposed by Professor Nelson, Erigeron colo-mexicanus, to replace the untenable E. cinereus A. Gray, is in my opinion rather distasteful. Fortunately I do not need to use the name, as the same species has been described by Dr. Greene under the name E. commixtus.

    Literature Cited:
    - Tidestrom, Ivar, 1925.  

    Tidestrom (1925) recognizes Buckley's E. nudiflorus.

    7. Erigeron nudiflorus Buckley, Proc. Acad. Phil. 1861. 456. 1862.
    Erigeron cinereus A. Gray, Mem. Amer. Acad. n. ser. 4: 68. 1849. Not E. cinereus Hook. & Arn. 1836
    Erigeron divergens cinereus A. Gray, Pl. Wright. 1: 91. 1852.
    E. commixtus Greene, Pittonia 5: 58.1902.
    Erigeron colo-mexicanus A. Nels. in Coulter, New Man. Rocky Mount. 529. 1909.
    Artemisia, pinyon and yellow pine belts; Utah and Colorado to Mexico.

    The author also recognizes E. divergens Torr. & Gray, and E. flagellaris Gray.

    Literature Cited:
    - Tidestrom, Ivar, and Sister Teresita Kittel, 1941.  

    Tidestrom and Kittell (1941) recognize Buckley's E. nudiflorus.

    9. Erigeron nudiflorus Buckley, Proc. Acad. Phil. 1861. 456. 1862.
    E. commixtus Greene, Pittonia 5: 58.1902.
    Artemisia, Pinyon and Yellow Pine belts; Utah and Colorado southward to Mexico.

    The authors also recognize E. divergens Torr. & Gray, and E. flagellaris Gray.

    Literature Cited:
    - Cronquist, Arthur, 1947.
    - Gray, Asa, and Charles Wright, 1852.  

    Cronquist (1947) accepts Grays (1852) treatment of E. tracyi as a variety of E. divergens.

    Along the way his key uses arrangement of hairs on stems as as a character to distinguish between E. divergens and E. flagelaris:

    6. Some or all of the hairs of the stem appressed or closely ascending, or the stem glabrous.
    which leads to E. flagellaris. The other half of the couplet:
    6. Hairs of the stem all spreading.
    leads to E. divergens where E. tracyi is treated as a variety.

    Key to the varieties of Erigeron divergens
    1. Earliest heads on leafy peduncles; plant without long stoloniform branches. A. var. typicus.
    1. Earliest heads on long naked peduncles; plant later producing long leafy stolons or stoloniform branches. B. var. cinereus.

    112B. Erigeron divergens var. cinereus A. Gray, Pl. Wright. 1: 91. 1852.
     
    E. cinereus A. Gray, Mem. Am. Acad. II 4:68. 1849. Not H. & A. 1836.
    E. nudiflorus Buckl. Prec. Acad. Phil. 1861: 456. 1862.
    E. commixtus Greene, Pittonia 5:58. 1902.
    E. tracyi Greene. Pittonia 5: 59. 1902.
    E. divergens nudiflorus A. Nels. Man. Bot. Rocky Mts. 529. 1909.
    E. colo-mexicanus A. Nels. Loc. Cit.
    ? E. dicladus Greene. Leafl. 2: 214. 1912.
    Earliest heads on long naked peduncles; plant later producing long loafy stolons or stoliform branches; basal leaves more often persistent than in var. typicus. Southern Nevada and Utah, through Arizona and New Mexico to northern Mexico, central Texas, western Oklahome, and southwestern Kansas; reported from Colorado and western Arkansas.
    Type: Fendler 374, near Santa Fe, New Mexico, 1847 (NY).
    … Texas: Tracy & Earle 279, Limpia Canyon, April 26, 1902 (type of E. commixtus Greene) (MI, NY); Tracy & Earle 320, Davis Mountains, April 28, 1902 (type of E. tracyi Greene) (MI, NY); … Colorado: Rydberg & Vreeland 5438, 5 miles southwest of La Veta, Huerfano Counto, May 22, 1900 (NY); …
    The variety cinereus has usually been treated as a distinct species under the name E. nudiflorus Buckl., but there are two many intermediates for it to stand. There are occasional intermediates with E. modestus, having the pubescence of the latter, and habit of E. divergens var. cinereus. These plants are more likely to have conspicuously lobed basal leaves than is typical var. cinereus.

    The variety cinereus is what we are calling E. tracyi today.

    Literature Cited:
    - Harrington, H. D., 1954.
    - Harrington, H. D., 1964, 2nd ed..  

    Harrington (2nd ed., 1964, p. 565) placed E. tracyi as a variety of E. divergens. This repeats his treatment of the taxon in his first edition (Harrington, 1954).

    8B. Erigeron divergens cinereus A. Gray, (var.) Pl. Wright. 1:91. 1852.
    E. cinereus A. Gray; E. nudiflorus Buckl.; E. commixtus Greene; E. colo-mexicanus A. Nels.; E. divergens nudiflorus (Buckl.) A. Nels. -- Earliest heads on long naked or nearly naked peduncles; plants producing long leafy stolons later in the season. --- Dry often sandy soil. Kansas ro Nevada, south to Texas and Arizona. Our records scattered over Colorado, mostly in the southern part, at 4000-8000 feet.

    Of note is Harrington's acceptance of E. commixtus but not E. tracyi.

    Literature Cited:
    - Nesom, Guy L., 1989.  

    Nesom (1989c) accepted E. tracyi as having priority over E. colomexicanus.

    Original Text Comments
    Erigeron tracyi an earlier name for Erigeron colomexicanus  
    I have been using Erigeron colomexicanus as the name for this species, but both E. tracyi and E. commixtus were published seven years earlier, simultaneously (immediate succession in the same paper: Greene 1902). Plants of the type collections of E. tracyi, as well as those of E. commixtus, are early season forms (essentially a basal rosette with a single, subscapiform, monocephalous stem) that had not yet produced runners characteristic of the species.  
    Erigeron tracyi Greene, Pittonia 5:59. 1902. Type: U.S.A. Texas. [Jeff Davis Co.:] Davis Mts., 28 Apr 1902, S. M. Tracy and F. S. Earle 320 (Holotype: US!; Isotypes: GH!, NY!, OC!).  
    Erigeron commixtus Greene, Pittonia 5:58. 1902. Type: U.S.A. Texas. [Jeff Davis Co.:] Cañon of the Limpia, Mountains of west Texas, 26 Apr 1902, S. M. Tracy and F. S. Earle 279 (Holotype: US!; isotypes: GH!, NY!, TAES!, TEX!).  
    Erigeron cinereus A, Gray, Mem. Amer. Acad. Arts n.s., 4 [Pl. Fendler.]; 68. 1849 (not Hook. & Arn. 1836). Erigeron divergens Torrey & A. Gray var. cinereus (A. Gray) A. Gray, Smithsonian Contr. Knowl. 3, Art. 5 [Pl. Wright.]:91. 1852. Erigeron colomexicanus A. Nels. [nom. Nov.], Man. Bot. Rocky Mts., 529. 1909. Type: U. S. A. NEW MEXICO. [Santa Fe Co.:] near Santa Fe, 1847, A. Fendler 374 (Holotype: GH!; isotypes: GH, NY! UC-2 sheets!, US!).  

    Literature Cited:
    - Flora of North America Editorial Committee, eds., 1993+.  

    Volume 20 of Flora of North America North of Mexico was published in 2006. The treatment of Erigeron was written by Guy L. Nesom.

    160. Erigeron tracyi Greene, Pittonia. 5: 59. 1902.
    Running fleabane
    Erigeron cinereus A. Gray 1849, not Hooker & Arnott 1836; E. colomexicanus A. Nelson; E. commixtus Greene; E. divergens Torrey & A. Gray var. cinereus A. Gray
    Annuals, biennials, or short-lived perennials, 2.5–8(–12, 18) cm; usually taprooted, sometimes fibrous-rooted, caudices simple or branched. Stems first erect (greenish proximally), then producing herbaceous, leafy, prostrate runners (stoloniform branches, sometimes with rooting plantlets at tips), densely hirsutulous (hairs spreading-deflexed, of relatively even lengths and orientations), sparsely minutely glandular. Leaves mostly basal (persistent in early season); blades oblanceolate to spatulate (obovate-elliptic laminae), 10–30(–60) × 3–6(–12) mm, cauline abruptly reduced distally, margins entire, dentate, or lobed, faces densely hirsute, eglandular. Heads 1(–3 rarely, from midstem or proximal branches). Involucres 3.5–4.5(–6) × 6–9(–12) mm. Phyllaries in 3–4 series, sparsely to moderately hirsute, minutely glandular. Ray florets 60–130; corollas white, often purplish abaxially, sometimes with an abaxial midstripe, 5–9 mm, laminae not coiling or reflexing. Disc corollas 2–3 mm (throats indurate and slightly inflated). Cypselae 0.7–1.3 mm, 2-nerved, faces sparsely strigose; pappi: outer of setae, inner of 12–16 bristles. 2n = 27.
    Flowering Mar–Oct. Desert scrub, grassy slopes, oak chaparral, pinyon-juniper woodlands, Douglas fir-ponderosa pine; 700–2300(–2400) m; Ariz., Colo., Kans., Nev., N.Mex., Okla., Tex., Utah; Mexico (Baja California, Chihuahua, Coahuila, Durango, Sonora, Zacatecas).
    In March through June, plants of Erigeron tracyi produce leaves in a basal rosette usually with a single, erect, monocephalous, stem. Stoloniform branches are soon formed (often recognized on pressed specimens by the leaves mostly on one side of the branches), and by the end of the season (August through October), prostrate runners are usually evident, sometimes forming terminal, rooting plantlets.
    Erigeron tracyi is similar in habit to E. flagellaris, particularly in the herbaceous stolons or stoloniform branches; the stem pubescence of E. tracyi is different, the stolons much less commonly produce rooting plantlets at the tips, and the plants tend to be perennial with woody or lignescent caudices, although they are variable both in habit and duration. Apparent hybrids with E. modestus and E. flagellaris are occasionally encountered, and the most common form of E. tracyi is perhaps (speculative) a stabilized, apomictic hybrid between the latter and E. divergens. All chromosome counts thus far have shown E. tracyi to be triploid and asynaptic.

    Literature Cited:
    - Snow, Neil, 2009.  

    Erigeron tracyi Greene - FNA
    [Erigeron cinereus A. Gray 1849 {not Hook. & Arnott 1836} - FNA
    [Erigeron colomexicanus A. Nelson] - KTZ, E
    [Erigeron colo-mexicanus A. Nelson] - SFE, W&W
    [Erigeron commixtus Greene] - FNA
    [Erigeron divergens Torr. & A. Gray var. cinereus A. Gray] - HDH, UTF
    [Erigeron divergens Torr. & A. Gray var. cinereus (A. Gray) A. Gray] - IMF
    [Erigeron modestus A. Gray] - GPF

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    11a. [10] Heads solitary on elongate, mostly leafless stems from the basal leaf clusters; plants developing long, nonrooting, stolon-like, spreading shoots. E. tracyi Greene [for Samuel Mills Tracy, 1847-1920]. Dry, gravelly floodplains and meadows, mimicking E. flagellaris but with spreading stem hairs. Flowering very early in the spring. The FNA suggests that this is a triploid apomictic hybrid between E. divergens and E. flagellaris. (E. colomexicanus is a later name.)

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield's (2015) description of E. tracyi does not describe the stem hairs, or the direction in which they point. Also, I think the stated length of the stoloniferous branches is quite short. Perhaps, the length should be expressed in decimeters, such as “... stoloniferous branches, 1-3(6) dm. Long.” Finally, the only synonym identified is E. colomexicanus perhaps because it has long been used in Colorado and New Mexico, but E. commixtus and E. nudiflorus are not noted.

    Erigeron tracyi Greene, Running Daisy. [E. colomexicanus A. Nelson]. Annuals, biennials, or short-lived perennials, 1-4 dm.; leaves oblanceolate to linear, entire or few-toothed, the stems forming stoloniferous branches, 1-3(6) cm. long; involucre 3-5 mm. hih, hirsute with curved hairs; disk flowers 2-2.5 mm. long; ray flowers 4-6 mm. long, white or pink; pappus double. Common on the eastern slope in open meadows and on dry slopes, with scattered occurrences on the estern slope, 4000-9000 ft. April-July. E/W

    Couplet 24 of Ackerfield's key to Erigeron addresses the question of stem hairs:

    24a Stems with spreading or tangled (pointing in all directions) hairs or mostly glandular … 25
    24b. Stems with appressed or ascending hairs (at above the middle), not glandular … 34

    Couplet 25 eventually ends up at Couplet 29 which differentiates between E. tracyi and E. divergens.

    Couplet 34 eventually leads to E. flagellaris.

     

    Other articles:
    • Golden Checklist Flora:  Erigeron vetensis;  

    Erigeron vetensis Rydb. Early Bluetop Fleabane.

     

    Literature Cited:
    - Rydberg, Per Axel, 1905.  

    Erigeron vetensis sp. nov.
    Densely cespitose-puvinate perennial ; stems 5-8 cm. high, hirsute, few-leaved ; leaves linear or linear-oblanceolate, hirsute, 2-4 cm. long; heads solitary, about 7 mm, high ; bracts hnear, acuminate, hirsute as w^ell as slightly glandular-puberulent ; rays purple, 8-10 mm. long, over i mm. wide; achenes strigose; pappus more or less double.
    In dry places on high mountains of southern Colorado at an altitude of 2400-3000 m. It is intermediate between E. radicatus and E. glandulosus, resembling the former most in pubescence, and the latter in habit.
    Colorado: Mountains near Veta Pass, 1900, Rydberg & Vreeland 5427 (type); Veta Mountain and Ojo, 5421, 5422; West Spanish Peak, 5424.

     

    Other articles:
    • Golden Checklist Flora:  Gaillardia aristata;  

    Gaillardia aristata Pursh “Blanketflower”

     

    Literature Cited:
    - Pursh, Frederick, 1814.
    - Schreber, Johann Christian Daniel van, 1791.  

    Pursh (1814, v. 2, p. 573) ...

    Original Text Comments
    656. GALARDIA. Gen. pl. 1323. FromPursh's Index Auctorum: “Gen. pl. — Car. a Linne, Genera Plantarum, Ed. 8vo. curante J. C. D. Schreber. Vol. 2. Francof. 1789. 1791. 8vo.”
     
    3. G. hirsutissima, scabra ; caule ramoso, foliis lineari-lanceolatis sessilibus, paleis pappi (5—7) ovato-lanceolatis longissime aristatis. aristata.  
    On dry hills on the Rocky-mountains. M. Lewis. ♂. v. s. in Herb. Lewis. Flowers orange-coloured.    
    Planta hirsutissima. Caulis erectus, teres, ramosus. Rami uniflori. Folia caulina alterna, longe-lineari-lanceolata, sessilia, scabra, hirsutissima. Calyx sub-imbricatus, polyphyllus : foliolis lineari-oblongis, acutissimis, disco duplo longioribus, margine coloratis. Corollulae : Radii circiter 12. obcuneatae, 3-fidae : laciniis oblongis, subaequalibus, obtusiusculis; Disci tubulosae, 5-fidae : laciniis extus purpurascentibus hirsutissimis. Receptaculum in ambitu subpaleaceum, centro setosum. Pappus : paleis 5—7. scariosis, ovatis, longissime-setaceo-aristatis.    

     

    Other articles:
    • Golden Checklist Flora:  Galinsoga parviflora;  

    Galinsoga parviflora Cav. “Gallant Soldier”

     

    Literature Cited:
    - Cavanilles, Antonio Jose, 1797.  

    Cavanilles (1791, v. 3, p. 41),

    Original Text Comments
    GALINSOGA PARVIFLORA. tab. 281.  
    308. GALINSOGA caule herbáceo: foliis oppositis, ovato-lanceolatis, subserratis: pedunculis elongatis uni, bifloris.  
    (a) Le réceptacle (loquitur de Amello tenuifolio) est chargé de paillettes , ainsi que chaque semenee , dont l'aigrette n'est pas formée par des poils: ce caractère indique un rapport marqué entre les deux premières espèces de ce genre, et les Verbesines.  
    Verbesina biflora. Hort. Reg. Parisiens.  
    Caulis herbaceus, glaber, bipedalis, ramis oppositis, iterum ramosis.  
    Folia opposita, ovato-lanceolata, subserrata, trinervia, glabra, nervis exceptis: petioli breves, subconnati.  
    Flores axillares et terminales, pedunculis elongatis uni aut bifloris.  
    Calix glaber, hemisphaericus, imbricatus squamulis ovatis dupli serie: harum quinque exteriores carinatae, apice scariosae; interiores vero planiusculae coloratae.  
    Radii fere semper quinque, aliquando septem, quorum corolla brevis alba: hermaphroditorum corolla lutea, quemadmodum anthera et stigmata.  
    Receptaculi paleae apice bi-trifidae, nonnumquam integrae; papi vero brevissimae, sublineares, ciliatae.  
    Habitat in Peruvia. O Floret Septembri.  
    Explìc. tab. a Flos integer. b Calix. c Radius auctus. d Flosculus disci auctus. e Semen. f Idem auctum. g Papi palea. h Receptaculi paleae.  
    Obs. Hanc plantam vidi in Regio horto Parisiensi anno 1785 nomine Verbesinae biflorae, ibi enata ex seminibus e Peruvia missis a D. Dombeyo : in patriam redux vidi eam iterum in Regio horto Matritense, cuius nomen tandem mutatum fuit in debitum Galinsogae. Obs. I saw it in the Royal Garden of Paris, in the year 1785 given the name of Verbesina biflorae, arisen from the seeds of Peru, which were sent by D. Dombeyo from there. I also saw it in my own country in the Royal Garden of Madrid, under a changed name, that of Galinsoga.

       

    Grindelia Willd.

     

     

    Other articles:
    • Golden Checklist Flora:  Grindelia squarrosa;  Notes on Grindelia, Willdenow, 1807;  

    Grindelia squarrosa (Pursh) Dunal “Curlycup Gumweed”

     

    Other articles:
    • Golden Checklist Flora:  Notes on Grindelia squarrosa;  

    First published in Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 1: 259 (1807)

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, p. 559) ...

    Original Text Comments
    637. DONIA. Ait. kew. ed. 2. t. 4. p. 82.    
    1. D. herbacea ; foliis oblongis amplexicaulibus serratis, squarrosa . calycinis squamis apice filiformibus revoluto-squarrosis. squarrosa.  
    In open prairies, on the banks of the Missouri. M. Lewis. ♃ Aug. Sept. v. s. in Herb. Lewis. ; v. v. cult. Flowers yellow, resembling those of Inula. The whole plant is viscous, and has a strong resiniferous or balsamic scent.    
    Caulis erectus, teres, albidus, corymboso-ramosus. Rami alterni, angulati, apice uni- aut triflori. Folia alterna, basi dilatata, amplexicauiia, oblonga, acuta aut obtusa, serrata, glabra, utrinque squamulis punctiformibus resiniferis obsita. Flores 1 — 3. in summitate ramulorum, corymbosi, flavi, ante explicationem copiose resiniferi. Calyx hemisphaericus, arcte imbricatus : Squamis apice filiformibus, revolutis et quasi caput Medusae referentibus. Flosculi radii lineari-lanceolatae, acutae. Semina oblongo-obovata. Pappus: aristis 2 — 4. deciduis. Receptaculum planum, nudum, favulosum.    
    It approaches near to Aster glutinosus Cav. ic. 2. p. 53. t. 168. which is the Donia glutinosa Ait. kew. ed. 2. t. 4. p. 82. which I distinguish by the following diagnosis : Donia glutinosa : frutescens ; foliis oblongo-obcuneatis sessilibus serratis, calycinis squamis linearibus erectis.    

    Literature Cited:
    - Dunal, Felix, 1819.  

    Dunal (1819, t. 2, p. 50) …

    Original Text
    2. Grindelia squarrosa.
    G. caule herbaceo , foliis oblongis amplexicaulibus serratis, involucri squamis apice filiformibus revoluto-squarrosis.
    Donia squarrosa. Pursh. fl. bor. am. 2. p. 559*. — Sims. bot . mag. t. 1706. — Desfont. Tab. ed. 2. p. 269.
    Hab. in pratis apricis secùs rivos fluminis Missouri. ( Pursh. ) ♃ Fl. aug. sept.
    Caulis erectus teres albidus corymboso-ramosus. Rami altérai an gulati apice 1-3 flori ex Pursh; omnes 1-flori in speciminibus mihi obviis. Folia alterna sessilia basi dilatata amplexicaulia oblonga acuta (interdum obtusa ex Pursh.), serrata , glabra , utrinque squamulis punctiformibus glutiniferis obsita. Authodia flava ante explicationem copiosè glutinifera erecta corymbosim disposita G. glutinosæ magnitudine. Involucrum hemisphæricum, squamis basi arctè imbricatis, apice filiformibus patulis révolutisve et ideo capitulum squarrosum constituentibus. Flosculi Iigulati patentes oblongo-lineares acuti integri. Semina oblongo-obovata , aristis 2-4 deciduis coronata. Receptaculum planum nudum favulosum.
    Etiamsi herbacea propiùs ad G. glutinosam quam ad alias accedit species.

     

    Other articles:
    • Golden Checklist Flora:  Grindelia subalpina;  

    Grindelia subalpina Greene “Subalpine Gumweed”

     

    Literature Cited:
    - Greene, Edward L., 1898.  

    Greene (1898, v. 3, n. 18, p. 297) in Sect. 6. Miscellaneous New Species describes G. subalpina.

    Original Text
    Grindelia subalpina. Low perennial, the stoutish corymbose-panicled stems seldom a foot high, only sparingly leafy except at base; lowest leaves oblanceolate, petiolate, acute, coarsely and remotely incised, usually scabrous- puberulent; the few cauline obloug-spatulate, rather remotely and sharply serrate, glabrous: depressed globose heads rather large ; bracts of the involucre numerous, with filiform squarrose-spreading tips, the whole very glutinous: rays numerous, narrow: bristles of the pappus 2 to 4, slender, barbellulate.
    High plains of southern Wyoming, and at subalpine elevations on the mountains of northern Colorado. Hitherto confused with G. squarrosa, which is wholly of the plains, strictly biennial, branching and very leafy, perfectly glabrous, and with stouter smooth pappus-bristles.

     

    Other articles:
    • Golden Checklist Flora:  Gutierrezia sarothrae;  

    Gutierrezia sarothrae (Pursh) Britton & Rusby “Broom Snakeweed”

     

    Literature Cited:
    - Pursh, Frederick, 1814.
    Full Size ImageHabitat of Coll. No. 1289, Gutierrezia sarothrae  

    Pursh (1814, v. 2, p. 540) described Solidago Sarothrae from a Lewis and Clark collection.

    Original Text Comments
    Sarothrae. 32. S. caule angulato scabro inferne nudo, superne corymboso, ramis summitate paucifloris, foliis linearibus stricte-appressis obsolete 3-nervibus utrinque scabris, axillis nudis, ligulis disco duplo longioribus.  
      On the plains of the Missouri. M. Lewis. ♃ Sept. v. s . in Herb. Lewis. About a span high; resembling Hypericum Sarothra in general habit very much. This and the two preceding have a great affinity to one another ; but I consider them good and permanent species, as there are no intermediate varieties, which might lead to the suspicion that all belong to one genuine species. The two preceding were Solidago lanceolata and Solidago tenuifolia, which are now treated as synonyms of Euthamnia graminifolia (L.) Nutt. and Euthamia caroliniana (L.) Greene ex Porter & Britton, respectively.

    Literature Cited:
    - Lagasca y Segura, Mariano, 1816.
    Full Size ImageColl. No. 1289, Gutierrezia sarothrae  

    Lagasca (1816, p. 30) published Gutierrezia as a place to put something that had no place.

    Original Text Comments
    G. N. GUTIERREZIA.  
    Ord. nat. Corymbiferae. Juss. Juxta Collumeliam Willdenowì.

    Char. gener. differ.

    Anthodium imbricatum , squamis apice reflexis : Corolla radiata. Receptaculum favosum, dentato-paleaceum. Pappus polyphyllus, paleaceus.

     
    372. GUTIÉRREZ. linearifolia : foliis linearibus integerrimis sparsis: floribus corymbosis.

    Planta suffruticosa, glabra, resinosa. Folia linearía, acuta, integerrima, vix lineam lata , pollicaria et amplius, superiora breviorà, sessilia. Corymbus pauciflorús,terminalis. Anthodium oblongum, 8-florum. Radius triflorus ligulatus. Hab. in N. H. ♃

    “Hab. in N. H.” — Habitat in Nova Hispania, i.e., Mexico.

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, v. 2, p. 163) published a new genus Brachyris and the specific name B. Euthamiae placing Pursh's (1814) in synonomy. As it turns out, though, Lagasca's genus name Gutierrezia and Pursh's specific name sarothrae will have priority.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1838-1843.  

    Torrey & Gray (1838-1843, v. 2, p. 193) recognized that our plant belonged in Gutierrezia. However, they adopted Nuttall's (1818) specific name Euthamiae when Pursh's (1814) Sarothrae had prioirty.

    Literature Cited:
    - Britton, Nathaniel Lord, and H. H. Rusby, 1887.  

    Britton & Rusby (1887, p. 10) placed our plant in Gutierrezia and applied Pursh's (1814) specific name sarothrae.

    Original Text Comments
    55. Gutierrezia Sarothrae (Pursh.). (Solidago Sarothrae, Pursh. Flor. Amer. Sept., ii., 540, 1814 : Bachyris Euthamniae, Nutt., Genera, ii. 163, 1818 : G. Euthamiae, T. & G.)  

    Literature Cited:
    - Lane, Meredith A., 1982.  

    As it happens, Lane (1982) designated a neotype for Gutierrezia linearifolia: MEXICO BAJA CALIFORNIA [Norte]: 1 mi. S Las Juntas (Sierra Juarez), Moran #16563, 6 Sep 1969 (Neotype: LL; isotype: SD). Lane also placed G. linearifolia Lag. in synonomy under G. sarothrae (Pursh) Britton & Rusby (Lane, 1980).

     

    Other articles:
    • Golden Checklist Flora:  Helianthus annuus;  

    Helianthus annuus L. “Common Sunflower”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 904) ...

    Original Text
      POLYGAMIA FRUSTRANEA
      HELIANTHUS.
    annuus. 1. HELIANTHUS foliis omnibus cordatis : nervis pone bafin unitis : extrorsum denudatis.
      Helianthus radice annua. Vir. cliff. 88. Hort. cliff. 419. Hort. upf. 268. Roy. lugdb. 180.
      Helenium indicum maximum. Bauh. pin. 276.
      Herba maxima. Dod. pempt. 264.
      Chryfis. Reneal. fpec. 84. t. 83.
      Habitat in Peru, Mexico. (.)

     

    Other articles:
    • Golden Checklist Flora:  Helianthus nuttallii;  

    Helianthus nuttallii Torr. & A. Gray “Nuttall's Sunflower”

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.
    - Torrey, John, and Asa Gray, 1838-1843.  

    Torrey & Gray (1842, v. 2, pp. 324-5) published H. nuttallii though apparently Nuttall, himself, had placed the name H. californicus on specimens in the herbarium of the Academy of Arts and Sciences in Philadelphia. Thomas Nuttall collected the type on the “plains of the Lewis River” – the Columbia River – and intended to give it the name of Helianthus californicus. However, the name was unavailable having been previously used by DeCandolle (1836) for a collection in California by David Douglas. Torrey & Gray (1842) applied the name H. nuttallii giving credit to Nuttall for the collection. An image of the type may be available at JSTOR (https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.ph00014119). However, it is securely locked away behind a paywall. Nuttall (1840-1841) does not mention this in his description of Compositae from his 1834-5 expedition across the continent.

    Original Text
    * * * * * Perennial : heads middle-sized : rays 8-24 : involucre irregularly imbricated ; the scales loose, or with squarrose-spreading often foliaceous summits, as long as the yellow disk (achenia glabrous). — Corona-solis.
    † Leaves commonly alternate or scattered, the lower often opposite, feather-verined, sometimes obscurely triplinerved.
    16. H. Nuttallii : stem smooth ; leaves alternate, the lower opposite, narrowly lanceolate-linear, acute, mostly entire, scarcely petioled, both sides scabrous ; scales of the involucre lanceolate-subulate, hirsute-ciliate towards the base; pappus of 2 linear-lanceolate chaffy awns or scales. — H. californicus, Nutt. ! in herb. acad. Philad. &c., not of DC.
    Plains of Lewis River, Nuttall ! — Stem apparently strict and simple. Leaves 4-6 inches long, 3-5 lines wide, feather veined, obscurely triplinerved near the base, somewhat cinereous beneath ; the lower remotely and slightly serrate. Heads nearly as large as in H. giganteus. Involucre clothed with whitish hairs, or often smoothish. The disk-corolla is 5-nerved, of with intermediate nerves corresponding with the axis of 2 only of the laciniae, but not reaching the apex.

     

    Other articles:
    • Golden Checklist Flora:  Helianthus pauciflorus ssp. subrhomboideus;  

    Helianthus pauciflorus ssp. subrhomboideus (Rydb.) O. Spring & E. Schilling “Stiff Sunflower”

    The synonyms (POWO, 2022) of H. pauciflorus Nutt. ssp. pauciflorus, which does not occur in Colorado, in order of date published, are:

    • Helianthus atrorubens Michx. In Fl. Bor.-Amer. 2: 140 (1803), nom. illeg. Described by Michaux as native to Pennsylvania to Carolina, whereas to day the range along the eastern seaboard is limited to Virginia.
    • Helianthus diffusus Sims in Bot. Mag. 45: t. 2020 (1818)
    • Helianthus pedunculatus Link in Enum. Hort. Berol. Alt. 2: 352 (1822)
    • Helianthus rigidus Desf. In Tabl. École Bot., ed. 3: 184 (1829)
    • Helianthus asper Bosse in Vollst. Handb. Bl.-Gärtn. 2: 564 (1829)
    • Helianthus crassifolius Nutt. In Trans. Amer. Philos. Soc., ser. 2, 7: 367 (1841)
    • Helianthus missouriensis Schwein. Ex Nutt. In Trans. Amer. Philos. Soc., ser. 2, 7: 368 (1841)

    Synonyms (POWO, 2022) of H. pauciflorus Nutt. Ssp. subrhomboideus (Rydb.) O. Spring & E. Schilling, which does occur in Colorado, again in order of pub;ication date, are:

    • Helianthus subrhomboideus Rydb. In Mem. New York Bot. Gard. 1: 419 (1900)
    • Helianthus × scaberrimus var. subrhomboideus (Rydb.) Farw. In Amer. Midl. Naturalist 8: 278 (1923)
    • Helianthus × laetiflorus var. subrhomboideus (Rydb.) Fernald in Rhodora 48: 79 (1946)
    • Helianthus rigidus subsp. subrhomboideus (Rydb.) Heiser in Mem. Torrey Bot. Club 22(3): 136 (1969)
    • Helianthus rigidus var. subrhomboideus (Rydb.) Cronquist in Fl. Pacif. Northw.: 528 (1973)
    • Helianthus pauciflorus var. subrhomboideus (Rydb.) Cronquist in Man. Vasc. Pl. North E. U.S. Canad., ed. 2: 864 (1991)
    • Helianthus rigidus var. subrhomboideus (Rydb.) B.L.Turner in Sida, Bot. Misc. 24: 6 (2003)

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    On his 1811 trip up the Missouri River, Thomas Nuttall collected a sunflower that has become our Helianthus pauciflorus. The location is given as Lower Louisiana. Since the current range of our plant is Missouri and northward, one would assume the collection was not made south of Missouri. He apparently brought seeds of this plant back to England. I have seen references to plants growing in Lambert's garden, the Chelsea garden, and the royal garden of Paris. These garden-grown plants can be found in the text below and would gave rise to multiple names for Nuttall's plant. I have not seen any reference to collected plant material, either in US herbaria, or the British Museum.

    Nuttall (1818, v. 2, p. 177) published this plant on April 3, 1818, as Helianthus pauciflorus. The actual date of publication is important because there were two more names published later in 1818; those by Cassini (1818) and Sims (1818)

    589. HELIANTHUS. L. (Sunflower.)
    Calix imbricated, subsquarrose, foliaceous. Receptacle paleaceous, flat. Pappus paleaceous, 2-leaved, caducous.
    9. * pauciflorus. Leaves opposite, linear-lanceolate, acuminate, serrate, nearly smooth; stem naked, trichotomous, few-flowered; calix closely imnricated; leaflets ovate. HAB. In Lower Louisiana. — Leaves sometimes ternately verticillate, very long, paler beneath and somewhat pubescent; ray and disk nearly the same colour. 4 or 5 feet high.

    Literature Cited:
    - Cassini, Alexandre Henri Gabriel, 1818.
    - Kartesz, John T., and Kancheepuram N. Gandhi, 1990a.  

    Cassini (1818) published the same plant grown from seeds in the Jardin du Roi of Paris. This name was published in September, 1818, per Kartesz & Gandhi (1990), below.

    Harpalium rigidum, H. Cass. Tige herbacée, haute d'environ cinq pieds, dressée, rameuse, cyliudrique, garnie de poils roides. Feuilles opposées, presque sessiles, lancéolées, pas sensiblement dentées, d'une substance ferme et roide, d'un vert glauque ou cendré, munies sur les deux faces de poils courts et roides. Calalhides grandes, solitaires au sommet des rameaux nus et pédonculiformes ; fleurs jaunes. (Cultivé au Jardin du Roi.) Harpalium rigidum, H. Cass. Stem herbaceous, about five high feet, erect, branched, cylindrical, furnished with stiff hairs. Leaves opposite, almost sessile, lanceolate, not noticeably toothed, of a firm and stiff substance, of a glaucous or ashen green, provided on both sides of short, straight hair. Calalhids large, solitary at the top of bare and stalked branches; yellow flowers. (Cultivated in the King's Garden.)

    Literature Cited:
    - Kartesz, John T., and Kancheepuram N. Gandhi, 1990a.
    - Sims, John, M.D., 1821.
    Full Size ImageSims (1818) Plate 2020 of Helianthus diffusus.  

    Meanwhile Sims (1818) Curtis's Botanical Magazine, Plate 2020 and following discussion, published a third name for our plant using an image from Lambert's garden and a description from the Botanic Garden at Chelsea. This name was published in October, 1818, per Kartesz & Gandhi (1990), below.

    Helianthus diffusus. Missouri Sun-Flower.
    ********************
    Class and Order.
    Syngenesia Polygamia Frustranea.
    Generic Character.
    Recept. paleaceum, planum. Pappus diphyllus. Cal. imbricatus, subsquarrosus.
    Specific Character.
    Helianthus diffusus ; caule hispido divaricato, foliis ovatis rigidis scabris serrulatis alternis oppositisque, pedunculis longissimis unifloris.

    Descr. Stem angular, purple, hispid : branches distant, rambling, bearing one terminal flower on a very long peduncle. The whole plant is clothed with stiff, stinging hairs, scarcely exceeds two feet in height, but spreads wide. Leaves both opposite and alternate, oblong-ovate, rigid, very rough. Calyx imbricated, scales ovate, in four series, smooth, with villous edges. Flowers large : rays about twenty, oblong-ovate, plicate, three-toothed, of a very full yellow colour. Floscules of the disk yellow: tube filiform pedicle-like : border cylindrical. Anthers dark purple. Stigmas revolute, golden yellow. Chaff of the receptacle linear-lanceolate, concave, green. Germens three-cornered, crowned with a two-leaved, awl-shaped pappus.

    From its rambling manner of growth, this plant does not appear very sightly in the garden ; but is one of the most desirable flowers imaginable for ornamenting rooms, as it lives long in water, and from its spreading branches, the flowers dispose well, and make a very brilliant appearance.

    A hardy perennial. Blooms in August and September. Our drawing was made from a specimen communicated by Aylmer Bourke Lambert, Esq. three years ago, out of his collection at Boyton, where it was raised from seeds collected on the borders of the Missouri, by Mr. Nuttall ; but our description was taken at the Botanic Garden at Chelsea in August last.

    Literature Cited:
    - Nuttall, Thomas, 1834b.
    - Nuttall, Thomas, 1840-1841.  

    Nuttall's expedition to Arkansas spanned 1818 to 1820, though he was in that state mostly in 1819.

    There is a voucher in the British Museum (BM001025610) that appears to be from Nuttall's trip to Arkansas. It is currently filed as Helianthus rigidus (Cass.) Desf. subsp. rigidus (Cass.) Desf.

    Nuttall (1834b) does not mention any Asteraceae in his "Collections towards a flora of the territory of Arkansas." He will though in Nuttall (1841) "Descriptions of new Species and Genera of Plants. "

    Literature Cited:
    - Cuvier, Frederic, 1821.  

    Cuvier (1821, v. 20, p. 300) in his dictionary of natural sciences repeats Cassini's (1818) diagnosis and name, while noting that Rene Dessfontaine (at the Jardin du Roi) thinks Sim's (1818) H. diffusus may be the same plant.

    Harpalion roide ; Harpalium rigidum, H. Cass., Bull. Des Sc., septembre 1818. La tige est herbacee, haute d'environ cinq pieds, dressee, rameuse, cyiindrique, garnie de poils roides. Les feuilles sont opposees, presque sessiles, lanceolees, pas sensiblement dentees, d'une substance ferme et roide, d'un vert glauque ou cendre, rnunies sur les deux faces de poils courts et roides.Les calathides sont grandes, solitaires au sommet des rameaux nus et pedonculiformes; les fleurs sont jaunes. Cette plante est cultivee au Jardin du Roi , ou nous avons observe, sur des individus vivans , les caracteres generiques et specifiques que nous venons de decrire. M. Desfontaines pense que c'est l'helianthus diffusus, plante vivace, de l'Amerique septentrionale, decrite dans le Botanical Magazine. (H.Cass.) Stiff Harpalion; Harpalium rigidum, H. Cass., Bull. Of Scs, September 1818. The stem is herbaceous, about five feet, erect, branched, cylindrical, furnished with stiff hairs. The leaves are opposite, almost sessile, lanceolate, not noticeably toothed, of a firm and stiff substance, of a glaucous green or ash, with hairs on both sides short and stiff. Calathids are large, solitary in tips of bare and stalked branches; the flowers are yellow. This plant is grown in the Jardin du Roi, where we observed, on living individuals, the generic characters and specific that we have just described. Mr. Desfontaines thinks that it is the Helianthus diffusus, perennial plant, from northern America, described in the Botanical Magazine. (H.Cass.)

    “H. Cass.” = Count Alexandre Henri Gabriel de Cassini (9 May 1781 – 23 April 1832) was a French botanist and naturalist, who specialised in the sunflower family (Asteraceae) (then known as family Compositae).

    Literature Cited:
    - Desfontaines, Rene Louiche, 1829, 3rd edition.  

    Desfontaines (1829) in his 3rd edition of the catalog, in English, “Catalog of the Plants of the Royal Garden of Paris with annotations on new or less known plants” published Helianthus rigidus, placing H. diffusus and Harpalium rigidum in synonomy.

    Helianthus.
    ...
    { rigidus Am. S. ♃
    diffusus? Bot Mag.
    Harpalium rigidum. H. Cas. Dict. V. 20 p. 3000.
    Desfontaines make no mention of Nuttall's Gen. Am. or his H. pauciflorus.

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.  

    Nuttall's (1841) "Descriptions of new species and genera of plants in the natural order of the Compositae" was mostly about his 1834 travels across the continent, though he also published plants from the Arkansas trip and some of those collected by William Gambel. He also published Helianthus crassifolius, now treated as a synonym of Helianthus pauciflorus ssp. pauciflorus.

    Helianthus * crassifolius; ♃, stem simple, herbaceous, subscabrous; leaves nearly all opposite, lanceolate, acuminate at either end, serrate, above smooth, beneath very scabrous, and, as it were, shagreened; stem naked, one to three-flowered, with a few small, linear leaves; involucrum closely imbricated, the scales ovate, slenderly and finely ciliate; achenium subquadrangular, with sometimes four scales, the two central ones smaller.
    Hab. Plains of Arkansa. About two feet high, the lower part of the stem very leafy, above nearly naked from the sudden diminution in the size of the leaves, the plant here and there scattered with drops of resin. Leaves half a foot long, or more, about an inch wide, very thiick and coriaceous. Rays about twenty, bidentate. Achenium pubescent above; with small scales from the inner angles; discal florets as usual, enlarged and pubescent towards the base. Allied to H. pauciflorus.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1841.  

    Torrey & Gray (1841, v. 2, p. 323-331) Flora of North America accepts H. rigidus Desf. and treats H. pauciflorus Nutt. as an obscure or little-known species. I would speculate this was the result of no actual plant material being available in America.

    11. H. rigidus (Desf.) : stem simple or sparingly branched, rough; cauline leaves opposite, very thick and rigid, lanceolate or oblong-lanceolate, acute or acuminate at each end, subsessile, obscurely serrate or entire, somewhat triplinerved, extremely hispid-scabrous on both sides ; the uppermost sometimes alternate; the radical oval, obtuse, strongly triplinerved, petioled ; scales of the involucre ovate, mostly obtuse, finely ciliate, numerous, closely imbricated; chaff of the receptacle obtuse; achenia somewhat hairy; pappus of 2 concave lanceolate scales, rather than awns, and frequently with one or several intermediate small scales. — Desf. Cat. Hort. Par. Ed. 3. p. 184. H. atrorubens, Michx.! Herb, in part; Hort. Berol. ! 1839 ; Bot. mag. T. 2668 ; Hook. ! Compan. To bot. mag. 1. p. 98 (var. foliis acutioribus). H. scaberrimus. Ell. ! Sk. 2. p. 423 ; DC. I. c. p. 588. H. crassifolius, Nutt. ! In trans. Amer. Phil. Soc. I. c. Harpalium rigidum, Cass, in dict. Sci. nat. 20. p. 300 ; DC. ! Prodr. 5. p. 583.

    β branches or peduncles simple, elongated; scales of the involucre ovate-lanceolate or ovate, more acute. — H. diffusus, Sims, bot. mag. T. 2020 (poor.) H. Missuricus, "Spreng. Pug. P. 21 ;" Link. Enum. 2. p. 352.?' H. Missouriensis, Schwein. ! Herb. ; Nutt. ! In trans. Amer. Phil. Soc. I. c. H. atrorubens, Bot. reg. t. 508 ; Hook. ! Fl. Bor.-Am. 1. p. 312 ; DC. Prodr. I. c. ex char. ; not of Linn.

    Plains and prairies, western part of Georgia, Elliott ! Illinois, Mr. Buckley ! And St. Louis, Drummond! To the Upper Missouri, Nuttall! Mr. Nicollet! Arkansas, Nuttall! Louisiana, Dr. Leavenworth! And Texas, Drummond! β Missouri, Nuttall! (spec, cult.) and Saskatchawan, Drummond! Aug.-Sept. — Plant stout, 1-3 feet high, rather naked above ; the rigid stem hispid with short papillose hairs, often smoothish above : the coriaceous thick leaves (cauline 3-5 inches long, an inch or less wide) very rough with short bristles arising from papillas, which give the surface a cinereous hue, and often a whitish blistered appearance. Heads few, but showy ; the disk about an inch in diameter; the 20-24 rays an inch long. Scales of the involucre regularly and closely imbricated in 3-4 series. Receptacle convex : the chaff entire, ciliate on the back towards the summit. Lobes of the disk-corolla purple : style yellow. Achenia somewhat lenticular, hirsute when young, but more smooth when mature, except the edges and summit. The pappus is variable, even in different flowers from the same individual ; consisting either of the two scale-like awns without intermediate squamellae, or of 1-4 of the latter on each side, or with the squamaellae confluent with the lateral scales, which become dilated and auriculate or lobed at the base, &c. &c. ; whence we conclude that Harpalium, Cass. Is founded upon insufficient and very inconstant characters. The var. β differs but slightly, and passes completely into the other forms of this well-marked species, so that it hardly merits to be distinguished. In some of the Texan specimens, and in those described by Nuttall as H. crassifolius, the leaves are more attenuated to each end, more serrated, and the upper surface comparatively smooth. — The stem sometimes exudes resin in small quantity.

    ‡ Obscure or little-known species.
    32. H. paucifiorus (Nutt.) : leaves opposite, linear-lanceolate, acuminate, serrate, nearly smooth ; stem naked, trichotomous, few-flowered ; scales of the involucre closely imbricated, ovate. Nutt. Gen. 2. p. 177.

    Lower Louisiana. — Plant 4-5 feet high. Leaves sometimes ternately verticillate, very long, paler beneath and somewhat pubescent. Ray and disk nearly the same color. Nutt. — This species is unknown to us ; we find no specimen in the herbarium of the Academy of Natural Sciences.

    Literature Cited:
    - Porter, Thomas C., and John M. Coulter, 1874.  

    Porter & Coulter (1874, p. 71) accepted Helianthis rigidus Desf.

    Helianthus rigidus, Desf.Hall & Harbour ; Dr. Smith. Cañon City, Brandegee. Platte River, Coulter.

    I don't see any vouchers in SEINet determined Helianthus pauciflorus or H. rigidus from Colorado that pre-dates 1874 publication date. Nor do I see any Hall & Harbour or Brandegee collections of either taxon. These searches in SEINet would include the Gray Herbarium and New York Botanic Garden by default.

    Literature Cited:
    - Gray, Asa, 1884.  

    Gray (1884, v. 1, pt. 2, p. 274-175) accepts H. rigidus Desf. as H. pauciflorus Nutt. slips farther into obscurity.

    “§ 2. Perennials : receptacle convex, or in some at length low-conical : lower leaves aalmost opposite. ”

    “ *** Stem and (mostly opposite and triplinerves) leaves more or less hispid, hirsute, or scabrous ... : chaff of receptacle enture or some 3-toothed at the apex, pointless : rays numerous and conspicuous”

    “ ** Disk of the head dark purple”

    H. rigidus, Desf. A foot or two (rarely 6 to 8 feet) high, rigid, sparingly branched : leaves very firm-coriaceous and thick, both sides hispidulous-scabrous, shagreen-like, entire or serrate, lightly triplinerved but indistinctly and sparingly veined ; lower oblong and ovate- lanceolate, attenuate at base into short winged petioles ; upper mostly lanceolate : heads comparatively large, showy (disk three-fourths inch high) : involucre pluriserially imbricated ; its bracts mainly ovate, obtuse or acutish, rigid, appressed, densely and minutely ciliate: rays numerous, generally inch and a half long : akenes oblong-obovate, 3 lines long : pappus of two large ovate-lanceolate paleae, and sometimes two or four rather stout intermediate paleae ! more commonly none. — Cat. Hort. Par. ed. 3, 184; Torr. & Gray, Fl. ii. 322. H. atrorubens, Michx. l. c, in part ; Bot. Reg. t. 508 ; Bot. Mag. t. 2668 ; DC. Prodr. v. 586. H. diffusus, Sims, Bot. Mag. t. 2020. H. Missuricus, Spreng. Syst. iii. 618, name in place of diffusus. H. scaberrimus, Ell. Sk. ii. 423. H. Missouriensis (Schweinitz) & H. crassifolius, Nutt. Trans. Am. Phil. Soc. l. c. Harpalium rigidum, Cass. Dict. Sci. Nat. xx. 200 ; DC. Prodr. v. 583, founded on the form with intermediate paleae to the pappus. Plains and prairies, Saskatchewan and Michigan to W. Georgia, Texas, and eastern part of Colorado. Sometimes the disk-corollas are at first yellow !

    Literature Cited:
    - Coulter, John Merle, 1885.  

    Coulter (1885, p. 186) also accepted Helianthis rigidus Desf. but gave no sources.

    4. H. rigidus, Desf. A foot or two (rarely 6 to 8 feet) high, rigid, sparingly branched : leaves very firm-coriaceous and thick, both sides hispidulous-scabrous, shagreen-like, entire or serrate ; lower oblong and ovate-lanceolate, attenuate at base into short winged petioles ; upper mostly lanceolate : heads comparatively large, showy; disk ¾inch high, dark purple or brownish: involucre pluriserially imbricated ; its bracts mainly ovate, obtuse or acutish, rigid, appressed, densely and minutely ciliate. Plains and prairies from Michigan to Texas and west to E. Colorado.

    Literature Cited:
    - Rydberg, Per Axel, 1900a.  

    Rydberg (1900, p. 419) in his Catalogue of the Flora of Montana and Yellowstone National Park publishes H. subrhomboideus Rydb.

    * Helianthus subrhomboideus.
    Perennial ; stem 3-6 dm. high, terete, striate, tinged with red, sparingly hirsute, simple ; leaves opposite, firm, very scabrous, triple-veined, generally not strongly serrate, the basal broadly ovate or obovate-spatulate ; stem-leaves rhomboid-ovate or rhomboid-lanceolate, short-petioled, 5-10 cm. long, the uppermost diminutive, lanceolate ; heads mostly solitary, sometimes 2 or 3, 1-1.2 cm. high and 1.5-2 cm. in diameter; bracts in 4—5 rows, oblong, acutish, densely white-ciliate on the margins ; disk dark brown or purplish ; rays about 1.5 cm. long.
    Nearest related to H. scaberrimus, but differs in the broader and shorter leaves, the less acute bracts, the smaller heads, and generally longer peduncles. It grows on prairies, up to an altitude of about 1000 m.
    Montana: Sand Coulee, 1885, R. S. Williams, 249; Columbia Falls, Mrs. Kennedy, 6 ; Judith Mts., 1882, R. W. Springer, XXXV.
    Nebraska: Keya Paha River, 1893, Clements, 2866 ; Long Pine, 1890, G. D. Sweezey, 70; Whitman, 1893, Rydberg, 1627 (type).
    Dakota: Mouth of Big Sioux River, 1853, Hayden Survey; Upper Missouri, Nicollet; Custer, 1892, Rydberg, 805.
    Assiniboia: Cypress Hills, 1880, John Macoum ; Souris Plains, 1883, J. M. Macoiun.
    Saskatchewan: 1857-8, E. Bourgeau.

    Literature Cited:
    - Rydberg, Per Axel, 1906.  

    Rydberg (1906, p. 373) accepted H. subrhomboideus replacing H. rigidus in part. I don’t think ne intended to say that A. Gray was the author of H. rigidus. Instead I think he meant that some of the plants that A. Gray would describe as H. rigidus should be segregated into H. subrhomboideus.

    5. Helianthus subrhomboideus Rydb. (H. rigidus A. Gray, in part) On plains from Man., Sask. And Mont. To Colo. And Neb. — Alt. 4000-8000 ft. — La Pagosa ; mountains, Larimer Co. ; near Pagosa Peak ; Cheyenne Mountain; Bosworth's ranch, Stove Prairie; Boulder; canon west of Palmer Lake; Horsetooth Mountain.

    Literature Cited:
    - Coulter, John M., and Aven Nelson, 1909.  

    Coulter & Nelson (1909, p. 548) accept H. scaberrimus Ell., placing H. rigidus Desf. and H. subrhomboideus Rydb. In synonomy.

    4. Helianthus scaberrimus Ell. Bot. S. C. & Ga. 2: 423. 1824. Stem 3-12 dm. high, rigid, sparingly branched: leaves very firm-coriaceous and thick, both sides hispidulous-scabrous, shagreen-like, entire or serrate; the lower ovate and ovate-lanceolate, attenuate at base into short- winged petioles; the upper mostly lanceolate: heads comparatively large, showy: disk 15-18 mm. high, dark purple or brownish ; involucre pluriserially imbricated ; the bracts mainly ovate, obtuse or acutish, rigid, appressed, densely and minutely ciliate. H. rigidus. (H. subrhomboideus Rydb. Mem. N. Y. Bot. Gard. 1: 419. 1900.) Plains and prairies; Montana to Colorado and Georgia.

    Literature Cited:
    - Kartesz, John T., and Kancheepuram N. Gandhi, 1990a.  

    Kartesz & Ganghi (1990) accept Helianthus pauciflorus Nutt.

    Helianthus.
    Helianthus rigidus (Cass.) Desf. (Cat. Pl., ed. 3, 184. 1829) was based on Harpalium rigidum Cass. (Bull. Sci. Soc. Philom. Paris 141. Sep 1818). For Helianthus rigidus ssp. rigidus, Heiser (Mem. Torrey Bot. Club 22[3]:131-138. 1969) cited several synonyms, including Helianthus diffusus Sims (Bot. Mag. 45:2020. Oct 1818) and Helianthus paucifiorus Nutt. (Gen. 2:177. 3 Apr 1818). Cleariy, Helianthus paucifiorus has priority over Harpalium ngidum and Helianthus diffusus; hence, the latter two cannot serve as basionyms. For the North American flora, we accept Helianthus paucifiorus Nutt. As the correct name.
     

    Kartesz & Gandhi (1990b) accept Helianthus pauciflorus Nutt. ssp. subrhomboideus (Rydb.) Spring & Schilling.

    Helianthus.
    The new combination Helianthus pauciflorus Nutt. ssp. subrhomboideus (Rydb.) Spring & E. Schilling (Biochem. Syst. Ecol. 18(1):22. Mar 1990) predates H. pauciflorus ssp. subrhomboideus (Rydb.) Kartesz & Gandhi (Phytologia 68(6):423. Jun 1990) and renders the latter to be superfluous. The March issue of Biochemical Systematics & Ecology was received in May 1990 at UNC-Botany Library, and by this time, our article in Phytologia was already in press. Nevertheless, we regret the oversight and correct the subspecies authorship to: (Rydb.) Spring & E. Schilling.

    Literature Cited:
    - Spring, Otmas, and Edward E. Schilling, 1990.  

    Spring & Schiling (1990) ...

    H. pauciflorus ssp. subrhomboideus. The following nomenclatural change is necessitated by the decision [6] that H. rigidus is not the correct epithet for this taxon: Helianthus pauciflorus Nutt. ssp. subrhomboideus (Rydb.) Spring and E. Schilling, comb. nov. Basionym: Helianthus subrhomboideus Rydb. (1900) Mem. N. Y. Bot. Gard. 1, 419. Synonym: Helianthus rigidus ssp. subrhomboideus (Rydb.) Heiser (1969) Mem. Torrey Bot. Club 22, 136. Oklahoma, Kay Co., Taylor 28348; Indiana, Heiser 439 (IU); Iowa, Ames Co., Stolzfus 7259 (IU).

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Helianthus pumilus;  Nuttall, 1841, publication details;  

    Helianthus pumilus Nutt. “Little Sunflower”

     
     

    Nuttall (1841, p. 366) described the Little Sunflower from plants he collected on plains of the Platte enroute to Oregon Territory.

    Original Text
    Helianthus * pumilus; ♃ hirsutely pilose and scabrous; leaves ovate-lanceolate, opposite, attenuated below, subpetiolate, nearly entire, and three-nerved, upper leaves lanceolate, alternate; involucrum hoary, hispid; sepals imbricated, lanceolate, acute, as well as the receptacular paleae; achenia smooth.
    Hab. Rocky Mountains and plains of the Platte. A low, perennial, simple stemmed species, about a foot high, leaves two to three inches long, about an inch wide. Capituli about three to five (apparently) sessile. Rays about sixteen, longer than the disk, paleae somewhat obtuse, hirsute at the summit, scales of the achenium rather large and wide.
    There are no online data records of the Nuttall collection of his Little Sunflower.

     

    Other articles:
    • Golden Checklist Flora:  Heliomeris multiflora;  

    Heliomeris multiflora Nutt. “Showy Golden Eye”

    The International Plant Names Index continues to use H. multiflorus, whereas nearly all other floras and checklists use the corrected Latin gender of H. multiflora.

    Literature Cited:
    - Nuttall, Thomas, 1848b.  

    Nuttall (1848, p. 171) described the genus Heliomeris and the species H. multiflorus from a collection by William Gambel in Upper California.

    Original Text
    * HELIOMERIS. †
    Capitulum many-flowered, heterogamous; rays ligulate, in a single series, neuter; discal florets tubular, hermaphrodite. Involucrum irregularly imbricated and leafy, in about two series, and rather spreading. Receptacle conic, the palea embracing the florets, lanceolate and acute. Corolla, rays ligulate, (10-12,) those of the disk tubular, the tube short, throat wide and cylindric, border five-toothed. Stigmata with obling tips. Achenia laterally compressed, somewhat tetragonous, smooth, and without any pappus.
    A perennial tall herb, exactly resembling an Helianthus, with narrow, entire, somewhat scabrous leaves, the lower ones opposite ; flowers yellow, terminal, numerous.
    H. *multiflorus. A stoutish perennial, three or four feet high, much branched ; stems terete and striated ; leaves narrow, lanceolate, entire, nearly sessile, above somewhat scabrous, opposite below, on the branchlets alternate ; flowers terminal, numerous and showy, bright yellow, with to to twelve rays, sometimes with deeper discoloured blotches towards the base or lower half, entire and oblong ; leaflets of the calyx oblong-lanceolate, somewhat spreading, in nearly a simple series ; achenia black and smooth. Scarcely distinguishable from Helianthus, except by a short conic receptacle, and naked seeds.
    Hab. Mountains of Upper California, (Dr. Gambel,) and was also collected in the Rocky Mountains by Mr. Gordon.
    ‡ In allusion to its close affinity to Helianthus.

    Literature Cited:
    - Harrington, H. D., 1964, 2nd ed..  

     

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Heterotheca foliosa;
    Full Size ImageColl. No. 1258, Heterotheca foliosa  

    Heterotheca foliosa (Nutt.) Shinners. “Hairy False Goldenaster”

    Full Size Image
    Inflorescence of Coll. No. 2177, Heterotheca foliosa
     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;  

    Nuttall (1840) described this from the Rocky Mountain plains, near the banks of the Platte.

    Original Text Comments
    Chrysopsis * foliosa; ♃, sericeously villous, and more or less canescent, the margin and lower surface of the leaves scabrous; flowers fastigiate, corymbose; leaves entire, oblong or oblong-ovate, subamplexicaule, crowded, acute, ciliate below; scales of the involucrum linear, acute, villous; achenium silky; pappus scarcely scabrous, outer pappus slender, dimidiate.
    Hab. In the Rocky Mountain plains, near the banks of the Platte. Flowering in August. About a foot high, sending up many hairy stems from the same root. Nearly allied to C. villosa, but far more pubescent and hoary, with the leaves widest at the base. In some specimens quite hoary, the hairs feel as soft as silk, but on removing this clothing, the under surface is covered with numerous scabrous elevations. A very showy species.  
     

    Heterotheca foliosa (Nutt.) Shinners was published in a journal called Field & Laboratory, contributions from the Science Departments of Southern Methodist University (Field & Lab. 19:71 (1951).)

     

    Other articles:
    • Golden Checklist Flora:  Heterotheca villosa;  

    Heterotheca villosa (Pursh) Shinners “Hairy False Goldenaster”

     

    Literature Cited:
    - Nuttall, Thomas, 1813.  

    Nuttall (1813) published Sideranthus integrifolius which is a nomen nudum.

    Original Text Comments
    80*Sideranthus integrifolius. ‡ M. ‡ = Perennial
    M. = from the Missourie.

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, p. 564) ...

    Original Text
      650. AMELLUS. Gen. pi. 131 5.
    villosus. 1. A. villosissimus ; foliis sessilibus oblongis acuminatis integerrimis, floribus axillaribus brevi-petiolatis, radiis integris, paleis setaceis.
      On the Missouri. ♃. v. s. Rays yellow.

    Literature Cited:
    - Shinners, Lloyd Herbert, 1951.  

    Publication of Heterotheca villosa by Shinners; not available online.

       

    Hymenopappus filifolius Hook. “Fine Leaved Wooly White”

     

    Literature Cited:
    - Hooker, Sir William Jackson, 1829-1840.

    Other articles:
    • Golden Checklist Flora:  Notes on Hymenopappus filifolius Hook. var. polycephalus;  

    Hooker (1833, 1(6), p. 317) published Hymenopappus filifolius

    Original Text
    2. H. filifolius ; lana alba decidua tomentosus, ramis paniculatis sparsis, foliis bipinnatifidis segmentis filiformibus elongatis integerrimis, floribus solitariis, involucri foliolis sub-12 ellipticis pubescentibus capitulo subbrevioribus, corollae tubo snperne campanulato, pappo brevissimo. — H. tenuifolius. Dougl. MSS. apud. Herb. Hort. Soc. Lond. (non Pursh.)
    Hab. On the undulating arid grounds of the Columbia, near the Wallawallah, and on the hanks of the Spokan and Flat-head Rivers. — 18 inches to two feet high, the stem loosely panicled above. This is quite different from the H. tenuifolius of Pursh, of Mhich I possess a specimen from Mr. Nuttall, gathered on the Arkansa. That has a remarkably dense corymb of flowers, whereas here the branches form a lax terminal panicle, each elongated extremity bearing a solitary flower. The pappus is extremely minute.

     

    Other articles:
    • Golden Checklist Flora:  Hymenopappus filifolius Hook. var. polycephalus;  Notes on Hymenopappus filifolius, Hooker, 1833;  

    Hymenopappus filifolius Hook. var. polycephalus (Osterh.) B.L. Turner "Many-Headed Fine-Leaved Wooly-White"

     

    Literature Cited:
    - Osterhout, George E., 1918.  

    Osterhout (1918, p. 90) ...

    Original Text Comments
    A NEW HYMENOPAPPUS FROM COLORADO  
    By Geo. E. Osterhout  
    Hymenopappus polycephalus sp. nov.  
    Perennial from a tap-root; stems one to several, 4-6 dm. high, tomentose, becoming glabrate, leafy at the base and less so to the middle, the stem having 6-8 leaves; much alternately branched, the branches beginning near the base; leaves 10-12 cm. long, pinnately or bipinnately parted into very narrowly linear and rather distant divisions, the upper smaller; inflorescence paniculate, the heads long peduncled and single at the ends of the branches; the bracts narrowly obovate, 5-6 mm. long, tomentose, somewhat thickened in the middle, with a rather broad scarious margin; flowers yellow, the throat of the corolla broadly campanulate, scarcely 1.5 mm. long, the reflexed lobes about a third as much, the tube of the corolla about as long as the throat, glandular, the pappus about half the length of the tube; the achenes narrowly obpyramidal 4 mm. long, long villous, but the pappus not covered by their villosity.  
    Another perennial, yellow-flowered, Hymenopappus has been described from Colorado, H. cinereus Rydb., but that is a smaller plant, with fewer heads of flowers, and is less leafy. Dr. Rydberg's description says: "Stem about 2 dm. high, branched, with 2-4 leaves." The close relationship of Hymenopappus polycephalus, however, is not with H. cinereus but with H. tenuifolius Pursh. It is as high and as leafy as H. tenuifolius, but the flowers are yellow, not dull white, and it is a perennial, not a biennial. The achenes are very similar to those of H. tenuifolius. H. cinereus is a synonym of H. filifolius var. cinereus I.M.Johnst. H. tenuifolius is an accepted name, for a mostly plains species, with just one collection in Jefferson County.
    H. polycephalus is found on the foothills of the eastern side of the mountains of northern Colorado at an altitude of 6,000 to 7,500 feet. It is plentiful in the open country about Livermore, Larimer Co., Colo., and northward to Dale Creek, and on into Wyoming, I think. It blossoms from the last of June to early in September. I have collected it a number of times, and for a time thought it belonged with H. tenuifolius. The type specimens were collected in the vicinity of Livermore, Larimer Co., Colo., Aug. 10-11, 1917, No. 5680.  
    Windsor Colo.  

    Literature Cited:
    - Turner, Billie L., 1956.  

    Turner (1956, p. 237) ...

    Original Text
    Osterhout proposed the specific name polycephalus in 1918, but it was not taken up by other workers. He apparently was very familiar with the plant in the field, and his numerous collections preserved at the Rocky Mountain Herbarium, with their annotated comments, indicate that he was dissatisfied with placing it either under H. cinereus Rydb. or H. tenuifolius. Nevertheless, Osterhout thought polycephalus was more closely aligned with H. tenuifolius since, in describing the species, he stated, “The close relationship of Hymenopappus polycephalus, however, is not with H. cinereus but with H. tenuifolius Pursh. It is as high and as leafy as H. tenuifolius, but the flowers are yellow, not dull white, and it is a perennial, not a biennial.” Johnson treated the entity as a questionable synonym of H. cinereus stating, “Hymenoppapus polycephalus is a puzzling form which suggests a perennial phase of H. tenuifolius.”
    In the present treatment, polycephalus has been treated as a variety of H. filifolius since its total characteristics indicate a much closer relationship to the members of this species. In spite of this closer affinity, var. polycephalus does have several characters which possibly link it to H. tenuifolius; indeed, there seems to be a considerable degree of intergradation between the two entities with respect to leaf dissection, amount of tomentum on the stem, and throat/lobe ratio.
    It is interesting to note that var. polycephalus, so far as known, is tetraploid with n = 34, while both var. cinereus and H. tenuifolius are diploid with n = 17. Var. polycephalus, in Colorado, occupies a geographical position between these two taxa. From these facts it might be speculated that at some past time the Rocky Mountain var. cinereus came in contact with H. tenuifolius so that extensive hybridization occurred on the diploid level and concomitantly amphiploidy occurred. (Evidence of a once farther eastward extension of H. filifolius var. cinereus is found in the disjunct races of this variety in the panhandle region of Texas, a considerable distance from the normal range now occupied by the variety.) With recession of this peripheral contact at a later time, the amphiploids survived in the habitats in which they were produced and have since spread to the area they now occupy. In the few meiotic slides prepared of var. polycephalus pairing appeared normal, there being 34 bivalents. Unfortunately, seed of this variety was germinated too late in the study of this group to provide material for genetical experiments to test this hypothesis.

     

    Other articles:
    • Golden Checklist Flora:  Liatris ligulistylis;  

    Liatris ligulistylis (A. Nelson) K. Schum. “Northern Plains Gayfeather”

    There is a single collection of Liatris ligulistylis (A. Nelson) K. Schum. “Northern Plains Gayfeather” in Golden s.l. and indeed in Jefferson County. State-wide, collections seem to be concentrated around Colorado Springs, and then scattered in odd locations in the interior mountains.

    First described as Lacinaria ligulistylis A. Nelson Bot. Gaz. 31: 405. 1901, it was placed in Liatris ligulistylis (A. Nelson) K. Schumann Just’s Bot. Jahresber. 29(1): 569. 1903.

     

    Other articles:
    • Golden Checklist Flora:  Liatris punctata;  

    Liatris punctata Hook. “Dotted Blazing Star”

     

     

    Other articles:
    • Golden Checklist Flora:  Lygodesmia juncea;  Notes on Stephanomeria pauciflora;  

    Lygodesmia juncea (Pursh) D. Don ex Hooker “Rush Skeletonplant”

    Superficially Lygodesmia juncea is very similar to Stephanomeria pauciflora. However, it turns out they are quite distinct, being separated by consistent differences in cotyledon, achene, and pollen morphology and base chromosome numbers (x = 9 in Lygodesmia; x = 8 in Stephanomeria).

    Both Weber & Wittmann (2012) and Ackerfield (2015) separate them on morphology of the pappus, with Lygodesmia having a pappus of capillary brinstles and Stephanomeria a pappus of plumose bristles.

     

    Other articles:
    • Golden Checklist Flora:  Mulgedium pulchellum;  

    Mulgedium pulchellum (Pursh) G. Don in R. Sweet “Blue Lettuce”

     

    Literature Cited:
    - Moulton, Gary E., 1999.
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, p. 502) published it as Sonchus pulchellus from an unknown dried specimen. It was not a Lewis & Clark collection, per Moulton (1999).

    Original Text
    pulchellus. 9. S. pedunculis squamosis, floribus corymboso-racemosis, foliis caulinis cordato-amplexicaulibus ovato-oblongis acutis integerrimus glabris.
      On the banks of the Missouri. ♃ Sept. v. s. Flowers large, beautiful blue.

    Literature Cited:
    - Bremer, Kare, 1994.  

    Frequently referred to, but outside my price range, Bremer (1994) apparently made a strong case for treating our plant as Mulgedium pulchellum.

    Literature Cited:
    - Flora of North America Editorial Committee, eds., 2006.  

    Flora of North America (2006), John Strother genus author, treats this taxon as Mulgedium pulchelllum (Pursh) G. Don

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittmann (2012) treat this taxon as L. tatarica (L.) Meyer subsp. pulchella (Pursh) Stebbins.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015) treats this taxon as Lactuca tatarica (L.) C. A. Mey var. pulchella (Pursh) Breitung.

    Literature Cited:
    - POWO, 2021 - 2023.  

    Plants of the World Online (Kew, 2021) treats it as Lactuca pulchella DC.

     

    Other articles:
    • Golden Checklist Flora:  Nothocalais cuspidata;  

    Nothocalais cuspidata (Pursh) Greene “Prairie False Dandelion”

     

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, suppl., p.742) published Troximon cuspidatum from a collection in upper Louisiana by John Bradbury.

    Original Text Comments and Interpretation
    Troximon cuspidatum. — T. scapo unifloro superne subtomentoso, foliis linearibus margine undato tomentosis, calycinis foliolis imbricatis cuspidatis glabris.

    In Upper Louisiana. Bradbury.v. s. in Herb. Bradbury. Flowers large, yellow. Resembles the preceding.

    The preceding was Troximon glaucum (=Agoseris glauca)

    Literature Cited:
    - Gray, Asa, 1884.

    Other articles:
    • Golden Checklist Flora:  Notes on Comandra umbellata ssp. pallida, DeCandolle, 1857;  

    Nothocalais was first published by Gray (1884, p. 420) as a section of Microseris.

    Original Text Comments and Interpretation
    § 4. Nothocálais. Pappus of 20 to 24 narrowly linear-lanceolate silvery-white paleae, occupying two or more series, with obscure mid-nerve, very gradually attenuate into a slender awn : akenes attenuate-fusiform : seed not reaching to the tapering summit : bracts of the oblong-campanulate involucre narrowly lanceolate, nearly equal, in about two series : perennial from a thick caudex. Intermediate between Microseris and Troximon ! — Gray, Proc. Am. Acad. xix. 65.  
    M. troximoides, Gray. Acaulescent or nearly so: leaves tufted on the caudex, rather fleslhy, narrowly linear-lanceolate, entire or undulate, 4 to 6 inches long : scapes a span to a foot high: involucre three-fourths inch high: ligules somewhat elongated: mature akenes half-inch long : pappus somewhat longer, its almost setiform paleae a quarter of a line wide below. — Proc. Am. Acad. ix. 211. — Wooded hills and open plains, Montana and Idaho (first coll. by Spalding), Washington Terr., and Oregon to N. W. California. Another first collection by Spalding was Comandra umbellata ssp. pallida, also in Washington Territory.

    Literature Cited:
    - Greene, Edward L., 1886.  

    Greene (1886, p. 55) elevated Gray's section Nothocalais to the rank of species.

    Original Text
    N. CUSPIDATA. — Akene little contracted, 3 lines long, filled by the seed : pappus of 40 — 50 unequal, very narrow, setose paleae and scabrous bristles: leaves all radical, longer than the flowering scapes: involucre glabrous. — Troximon Pursh, Fl. ii. 742; Torr. & Gray, Fl. ii. 489; Gray, Syn. Fl. ii. 437: T. marginatum, Nutt. Gen. ii. 127.

    On bleak, stony hills and fertile prairies, from Dakota and Colorado to Wisconsin and Illinois. Scarcely distinguishable from its far Western congeners except by the pappus. The undulate-crisped, white-hairy margins of the grassy leaves of this giving it an aspect so strikingly unlike the general appearance of the other species of his genus Troximon, were points not overlooked by that well traveled and most keenly observant botanist, Mr. Nuttall. That he noticed the peculiarity and was impressed by it is evinced by his effort to invest the species with a new specific name, marginatum, more appropriate than Pursh's cuspidatum, which was given to it in reference to the acuminate rather than cuspidate bracts, and has, therefore, no fitness, but which must needs be retained in deference to its priority. The name marginatum would, indeed, be equally and in the same way, applicable to each of the three known species of Nothocalais.

       

    Oligosporus Cassini “Tarragon, Sagewort”

     

    Literature Cited:
    - Cassini, Alexandre H. G., 1817.

    Other articles:
    • Golden Checklist Flora:  Notes on Artemisia campestris, Weber & Wittmann, 2012;  Notes on Artemisia dracunculus, Weber & Wittmann, 2012;
    • Glossary:  calathid;  synantheree;  

    Cassini (1817, p. 31) dscribed his new genus of Oligosporus

    Original Text Translation
    51. Oligosporus. Ce genre, ou sous-genre, de la tribu des anthemidées, comprend toutes les espèces d' Artemisia, L., dont la calathide est composée de fleurs femelles et de fleurs males. Telle est, par exemple, l' Artemisia campestris, L. 51. Oligosporus. This genus, or subgenus, of the anthemid tribe, includes all species of Artemisia, L., whose calathid is composed of female and male flowers. Such is, for example, Artemisia campestris, L.

       

    Packera A. Löve & D. Löve

    SEINet (2021+) data includes collections of Packera that have been determined the following species:

    • Packera cana, two collections, Genesee Park, and Pine Valley Ranch.
    • Packera fendleri
    • Packera multilobata, one collection, near Evergreen (GREE24339).
    • Packera plattensis
    • Packera pseudaurea, three collections, Rocky Flats, Pine Valley Ranch, and South Platte (town).
    • Packera tridenticulata, eleven collections, mostly high plains, one near Evergreen.
    • Packera werneriifolia, Two collections, Meyer Ranch, and Rampart Range.
    There are no collections of P. streptanthifolia made in Jefferson County.

    Literature Cited:
    - Harrington, H. D., 1964, 2nd ed..  

    Harrington (1964, 2nd ed., p. 610) distinguishes between Packera fendleri and P. plattensis by the hairiness of the achenes, those of P. fendleri being glabrous, and those of P. plattensis hispidulous along the angles.

    Literature Cited:
    - Löve, Áskell, and Doris Löve, 1976.  

    Love & Love (1976) published Packera

    Literature Cited:
    - Weber, W. A., and Áskell Löve, 1981.  

    Weber & Love (1981) published fifty-one new names in Packera

    Literature Cited:
    - Trock, Debra, 2006.  

    Trock (2006) key to Packera has two ways to get into P. fendleri and four ways to get into P. plattensis.

     

    Packera streptanthifolia (Greene) W. A. Weber & Á. Löve, Phytologia. 49: 48. 1981. Packera pseudaurea (Rydberg) W. A. Weber & Á. Löve, Phytologia. 49: 48. 1981. Packera plattensis (Nuttall) W. A. Weber & Á. Löve, Phytologia. 49: 48. 1981. Packera fendleri (A. Gray) W.A. Weber & Á. Löve Coll. No. 2064
    Rocky Mountain groundsel False-gold groundsel Prairie groundsel Fendler's Ragwort
    Perennials, 10–50+ cm; fibrous-rooted (caudices weak to stout, horizontal to suberect). Perennials, 20–70+ cm; fibrous-rooted (bases simple or branched, horizontal to erect). Biennials or perennials, 20–60+ cm; rhizomatous and/or fibrous-rooted (bases erect to suberect), sometimes stoloniferous (mostly eastern populations). Perennials, 10–40+ cm; rhizomatous (rhizomes horizontal to suberect, branched). Perennial herb, 25-35 cm., caudex , some, perhaps older ones, are stout and erect;
    Stems 1 or 2–5, clustered, usually glabrous, sometimes sparsely floccose-tomentose proximally and in leaf axils. Stems usually 1, sometimes 2–4, clustered, glabrous or sparsely tomentose proximally. Stems 1 or 2–3, clustered, floccose-tomentose proximally and in leaf axils, otherwise sparsely tomentose or glabrescent. Stems 1 or multiple (crowded to subcespitose), floccose-tomentose or glabrescent. Stem, tomentose, especially in axils, thinning distally;
    Basal leaves (and proximal cauline, relatively thick and turgid) petiolate; blades spatulate to oblanceolate, or ovate to orbiculate, 20–40+ × 10–30+ mm, bases tapering to abruptly contracted or subcordate, margins entire, crenate, dentate, or weakly lobulate (faces usually glabrous, sometimes hairy). Basal leaves petiolate; blades usually broadly lanceolate to ovate, sometimes subhastate, 20–50+ × 20–40+ mm, bases truncate to subcordate or obtuse, margins crenate, denticulate, bluntly dentate, or sharply dentate (proximal cauline leaves petiolate; margins usually pinnately lobed to laciniate, sometimes subentire). Basal leaves (and proximal cauline) petiolate; blades narrowly elliptic to elliptic-ovate or oblanceolate to suborbiculate or sublyrate, 20–70+ × 10–30+ mm, bases tapering to rounded or abruptly contracted, margins subentire to crenate, serrate-dentate, or pinnately lobed (abaxial faces floccose-tomentose, especially along midribs, ± glabrescent). Basal leaves petiolate; blades lanceolate to oblanceolate, 30–60+ × 10–30+ mm, bases tapering, margins shallowly, evenly pinnatifid to pinnatisect or wavy (adaxial faces floccose-tomentose or subglabrescent). Basal leaves, petiole, 35-40 mm, long, thin not winged, tomentose, blade 20-24 mm. × 7-9 mm. wide, ovate, some (7 of 23 = 30%) lyrate, 2-7-2.8 × longer than wide, mid-vein tomentose thinning distally, tips, mostly entire, some 3-toothed;
    Cauline leaves gradually to abruptly reduced (± petiolate or sessile; entire or subentire). Cauline leaves gradually reduced (becoming sessile, sometimes clasping). Cauline leaves gradually reduced (petiolate, sublyrate or pinnatisect, abaxial faces sparsely hairy; distals sessile, subentire to irregularly dissected). Cauline leaves gradually reduced (sessile; lanceolate to oblanceolate, pinnatisect to wavy). Cauline leaves, reduced distally, sessile, base not auriculate, inreasingly pinnatisect distally;
    Heads 2–20+ in loose, corymbiform or subumbelliform arrays. Peduncles bracteate, glabrous or sparsely tomentose. Heads 5–20+ in open or congested, subumbelliform arrays. Heads 6–20+ in open or congested, corymbiform arrays. Peduncles conspicuously bracteate, sparsely to densely tomentose. Heads 6–25+ in open or compact, corymbiform arrays. Inflorescence, heads #3-5 per stem, >leaves;
    Peduncles bracteate, glabrous. Peduncles bracteate, densely to irregularly floccose. Peduncles, 8-12 mm.;
    Calyculi conspicuous. Calyculi inconspicuous. Calyculi inconspicuous. Calyculi 0 or inconspicuous (bractlets red-tinged). (Calyculi inconspicuous)
          Involucre, 5 mm. × 9 mm. wide, ovoid, thinly tomentose;
    Phyllaries (8–)13 or 21, green (tips sometimes cyanic), 4–7+ mm, glabrous. Phyllaries (13–)21(–30+), light green, 3–8 mm, glabrous. Phyllaries 13 or 21, green (tips sometimes cyanic), 5–6+ mm, densely tomentose proximally, glabrescent distally. Phyllaries 13, green, 5–7 mm, floccose proximally to glabrescent distally. Phyllaries, in 2 equal series, 5.5 mm. × 1.0-1.3 mm. wide, green, thinly tomentose, margins, flat, scarious, tip, reddish brown;
      Receptacle, epaleate;
          Flowers, of 2 kinds;
    Ray florets 8 or 13; corolla laminae 5–10 mm. Ray florets 0, 8, or 13; corolla laminae 6–10+ mm Ray florets 8–10; corolla laminae 9–10 mm. Ray florets 6–8+; corolla laminae 5–7 mm. Ray flowers, #10-12, tube 3 mm. + blade 6.5-7.5 mm. × 2.7 mm. wide, color yellow, fertile;
    Disc florets 35–60+; corolla tubes 2–4 mm, limbs 2.5–4 mm. Disc florets 70–80+; corolla tubes 2.5–3.5 mm, limbs 2–3 mm. Disc florets 60–70+; corolla tubes 2.5–3.5 mm, limbs 3.5–4.5 mm. Disc florets 30–40+; corolla tubes 2.5–3 mm, limbs, 2.5–3.5 mm. Disk flowers many, tube 4 mm. + lobes 0.5 mm., throat expanding gradually, yellow, open, bisexual;
          Pappus, many, well-developed. bristles, 4.5 mm., ±equal;
    Cypselae 1–2.5 mm, glabrous; pappi 3–6 mm. Cypselae 1–1.5 mm, glabrous; pappi 4.5–5.5 mm. Cypselae 1.5–2.5 mm, usually hirtellous, sometimes glabrous; pappi 6.5–7.5 mm. Cypselae 2.5–3 mm, glabrous; pappi 4–5 mm. 2n = 46. Fruit, 1.2 mm. × 0.5 mm. wide, compressed front-to-back, color brown, margin, thinly pubescent;
    2n = 46, 92. 2n = 46, 92. 2n = 46.  
    Flowering late May–late Aug. Forests, open meadows, valleys, dry to damp and loamy soils; 1000–3400 m; Alta., B.C., N.W.T., Sask., Yukon; Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo. Flowering mid Apr–early Jun(–mid Jul, north). Prairies, meadows, open wooded areas, along highways, railroads, around mining and construction areas, usually on limestone; 50–1800 m; Ont., Sask.; Ark., Colo., Ga., Ill., Ind., Iowa, Kans., La., Mich., Minn., Miss., Mo., Mont., Nebr., N.Mex., N.Dak., Ohio, Okla., Pa., S.Dak., Tenn., Va., Wis., Wyo. Flowering late May–early Oct. Steep slopes, loose, dry rocky or gravelly soils, along streams, open forests, disturbed sites; 1600–3200 m; Colo., N.Mex., Wyo.  
    Packera streptanthifolia is widespread and variable throughout the Western Cordillera. It includes weakly defined phases that have been treated as distinct species or as varieties. Characteristics used to delimit those taxa often overlap and are difficult to score; some ""phases"" grade into each other. Northern populations are sometimes segregated as a distinct taxon (e.g., Senecio streptanthifolia var. borealis; J. F. Bain 1988). Packera plattensis is abundant, widespread, and almost weedy. Putative hybrids with other species are known. Plants in mesic, remnant prairies in the east are sometimes stoloniferous. Packera fendleri is abundant, almost weedy in the southern Rocky Mountains. It thrives in a wide range of elevations and in a wide variety of habitats; flowering times vary. It frequently grows in close association with other species of Packera and may hybridize with them.  
    Filename: 280Packera_plattensis_streptanthifolia.html

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittmann (2012) key focuses mainly on the shape of the basal leaves …

    Original Text Interpretation
    3a. Basal leaves and most stem leaves deeply pinnatifid or runcinate-pinnatifid. P. fendleri (A. Gray) Weber & Love. Abundant in gravelly soil, open forests of the foothills. Producing rosettes from long, slender rhizomes; leaf lobes uniform, shallow; leaves white tomentose.  
    3b. Basal leaves oval, rately pinnatifid except at the very base. … 4  
    4b. Basal leaves narrower, on winged petioles, irregularly toothed, lobed or entire. … 10 Leads eventually to P. plattensis.
     

    Ackerfield (2015) key also focuses on the shape of the basal leaves …

    Original Text Interpretation
    2a. Basal and stem leaves nearly all deeply and evenly pinnately dissected or runcinate-pinnatifid … 3 This couplet leads to P. multilobata and P. fendleri.
    2b. Basal leaves with entire, toothed, or crenate margins (in P. plattensis and P. tridenticulata a few basal leaves may have pinnatisect margins, but the majority of the leaves will have entire or merely toothed margins), stem leaves entire to pinnatisect or sublyrate … 4 This couplet eventually leads to P. plattensis after elininating seven other taxa, and arriving at a group including P. plattensis, P. streptanthifolia, and P. pseudaurea.

     

    Other articles:
    • Golden Checklist Flora:  Packera fendleri;

    Locations: Schweich Hill.  

    Packera fendleri (A. Gray) W.A. Weber & Á. Löve. (Syn: Senecio fendleri A. Gray ) Fendler"s Ragwort.

    Fendler's Ragwort — Packera fendleri (A. Gray) W.A. Weber & Á. Löve — is widespread and fairly common in dry meadows and slopes of Golden s.l.. The author has collected it on Schweich Hill, and Deadman Gulch. Others have collected the plant on North and South Table Mountains.

    P. fendleri is frequently confused with P. plattensis which is more a plant of the great plains.

    Literature Cited:
    - Gray, Asa, 1849.

    Other articles:
    • Golden Checklist Flora:  Gray, 1849, publication details;

    Locations: Santa Fe River.  

    Gray (1849, p. 109) writing in Plantae Fendlerianae Novi-Mexicanae described P. fendleri from a collection made “ ...along the Creek, twelve miles above Santa Fé …” The current accepted name is Santa Fe River.

    Original Text Comments
    444. S. Fendleri (sp. nov.) : perennis, flocoso-incanus, demum subglabratus ; caule folioso erecto pedali corymbosi-ramoso ; foliis oblongis omnibus pinnatifidis supra glabratis inferioribus in petiolum nudum attenuatis summis sessilibus haud amplexicaulibus, segmentis 11-21 confertis oblongis obtusissimis plerisque inciso-dentatis seu 2-4-lobatis ; corymbis compositis polycephalis ; involucro campanulato fere ecalyculato 12-phyllo multifloro ; ligulis 7-8 oblongis disco duplo longioribus ; acheniis glaberrimis. — Foot of mountains along the Creek, twelve miles above Santa Fe ; June, July. (478, † 480.) — A well-marked species, related to S. eremophilus. Stem stout. Leaves from 2 to 4 inches long including the petiole, some of the lower often only sinuate-pinnatifid, but commonly all deeply pinnatifid or pinnately parted, the crowded lobes from one fourth to half an inch long, early glabrate above ; but still floccose of white-wooly underneath. Involucre 3 lines long. Rays 4 or 5 lines long. *  

    There follows a half page discussion of similar Senecio found throughout the southwest, such as Senecio multilobatus (Torr. & A. Gray, ined.)

     

    Other articles:
    • Golden Checklist Flora:  Packera plattensis;  

    Packera plattensis (Nuttall) W. A. Weber & Á. Löve. “Prairie Groundsel”

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;  

    Original Text
    Senecio * Plattensis; ♃, somewhat pubescent; base of the stem arachnoidly tomentose; leaves all pinnatifid, the radical petiolate, cauline amplexicaule, lobes oblong, denticulate, the centre lobe sublanceolate; corymb nearly simple; involucrum subcampanulate, minutely bracteolate; sepals about twenty, scute; rays usually twelve, oblong, a little longer that the short involucrum; achenium puberulous; pappus about the length of the florets.
    Hab. In the Rocky Mountain range, and in Arkansa. About ten to fourteen inches high ; stem simple, striated. Corymb nearly simple, with ten to twelve heads of flowers, pedicels one to two inches long, slightly bracteolate. The Arkansa specimen is taller and more slender, with the primary small radical leaves entire and smooth, the leaves more elongated, and less denticulate. The whole habit of the plant, as well as the flowers, are very similar to S. tomentosus, at lease the smoother variety, but the achenium is less pubescent.

     

    Other articles:
    • Golden Checklist Flora:  Packera tridenticulata;  

    Packera tridenticulata (Rydb.) W.A. Weber & Á. Löve “Three-Tooth Ragwort”

     

    Literature Cited:
    - Gray, Asa, and Charles Wright, 1852.
    - Rydberg, Per Axel, 1900b.

    Other articles:
    • US Highway 24:   in Buena Vista;

    Locations: Cottonwood Creek.  

    Rydberg (1900, v. 27, p. 175) ...

    Original Text Comments
    7. Senecio tridenticulatus sp. nov.  
    Senecio aureus var. borealis A. Gray, Pl. Wright. I : 125. 1852 ; Not T. & G. 1843 ; S. aureus var. compactus A. Gray, Syn. Fl. I2: 391 ; in part.  
    Perennial with a branched caudex, in age perfectly glabrous, or slightly floccose at the base of the leaves : stems very slender, about 3 dm. high : basal leaves very narrowly oblanceolate, thick with a slender petiole, slightly 3-toothed at the apex or wholly entire, 4-8 cm. long and 4-5 mm. wide: stem leaves linear and subentire : cyme open and corymbiform : heads 7-8 mm. high bracts lanceolate, acute ⅔ or ¾ as long as the disk : rays light yellow, about 8 mm. long and 1—1.5 mm - wide > 3-nerved ; achenes hispid-puberulent on the angles.  
    This species differs, from the preceding in the slender stem and the open cyme, and also in the form of the leaves. The latter character also separates it from the next following. It grows in wet sandy soil. The type was growing at an altitude of 2400 m. Wright's specimens are past blooming, and good characters could not be taken from them, wherefore I have made Sheldon's speci- mens the type. The latter were mixed with some of the next. [Plate 5, f. 12.] The preceding was Senecio compactus (A. Gray)Rydb. and the following was Senecio oblanceolatus Rydb., both of which are now treated as synonyms of Packera thurberi (A.Gray) B.L.Turner.
    Colorado : Cottonwood Creek, Buena Vista, 1892, C. S. Sheldon (type in the herbarium of N. Y. Botanical Garden).  
    Texas: Mountains beyond the Limpia, 1849. Wright, 403. See Plantae Wrightianae (Gray & Wright, 1852).

    Literature Cited:
    - Weber, W. A., and Áskell Löve, 1981.  

    Weber & Love (1981, p. 48) published the name Packera tridenticulata.

     

    Other articles:
    • Golden Checklist Flora:  Picradeniopsis oppositifolia;  

    Picradeniopsis oppositifolia (Nutt.) Rydb. ex Britton. “Opposite Leaf Bahia”

     

    Literature Cited:
    - Moulton, Gary E., 1999.
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, v. 2, p. 167-168) ...

    Original Text Comments
    568. * TRICHOPHYLLUM. †  
    Calix oblong-cylindric, many-leaved, equal. Radial florets oblong. Receptacle naked. Pappus paleaceous, minute, 5 to 8-leaved, leaflets obtuse, awnless.  
    Herbaceous; leaves alternate? or opposite, palmately pinnatifid, tomentose or villous; peduncles 1-flowered, dichotomal and terminal.  
    Species.  
    1. T. lanatum. Actinella lanata, Ph. 2. p. 560. Every where whitely and lanuginously tomentose; leaves alternate, those of the stem subpalmately pinnatifid, of the branches linear and entire; peduncle elongated, the summit thicker. Hab. Near the sources of Columbia river. M. Lewis. Flowering in June and July. v. s. in Herb. Lambert. — Perennial; stem erect and branching, about a foot high. Leaves alternate? (perhaps not constantly so) those of the stem elongated, narrow at the base, dilated and divided pinnatifidly above, division ligulate and somewhat toothed, uppermost entire. Calix oblong-cylindric, composed of a simple series of leaves, about 12 to 14, linear-lanceolate, acute. Rays about the same number, oblong, bidentate. Pappus 5 to 8-leaved. Seed pentangular? glabrous, attenuated downwards. — The flowers are bright yellow, and in form and character strongly resemble those of the genus Tagetes. This would become Eriophyllum lanatum (Moulton, 1999).
    2. * oppositifolium. Decumbent and much branched, shortly and canescently pubescent; leaves opposite, all palmately trifid, segments ligulate, simple or divaricately subdivided; peduncle filiform, mostly dichotomal, scarcely longer than the leaves. Hab. On denudated sterile hills, near Fort Mandan; abundant. Flowering in July and August. — Perennial? stem diffuse, 6 to 12 inches high, grooved; oppositely branched. Leaves petiolate, trifid, canescent, pubescence very short, segments about an inch long, thickish and opaque, the lateral ones mostly bifid, the central one trifid, all somewhat obtuse and linear. Peduncle slender, 1 to 2 inches long, a little thicker under the calix. Calix oblong-cylindric, simple, leaflets 5 to 8, oblong-ovate, erect; rays about the same number, very short. Pappus paleaceous, 5 to 8-leaved, minute, leaflets partly obtuse and somewhat lacerate. Seen nearly smooth, rather long, and attenuated downwards, or inversely conic. Receptacle small and naked. — The whole of this plant is very sensibly bitter and destitute of aroma.  
    There is nothing in the habit of this genus which would lead us to suppose it allied to Actinella of Jussieu, and scarcely more in the generic character. In Actinella the calix is very short, flat, and horizontally spreading; the leaflets of the paleaceous pappus awned, and the seeds villous; the leaves are also alternate and entire. The proximate affinity of the present genus is to Tagetes.  

    † The copious pubescence, particularly distinguishing these plants from the genus Tagetes.
     

    Literature Cited:
    - Britton, Nathaniel Lord, 1901.  

    Britton (1901, p. 1008) Flora of the Northern States and Canada ...

    Original Text Comments

    76. PICRADENIOPSIS Rydb.
    [Bahia Nutt., not Lag.]
    Britton gives Rydberg credit for the genus name, but I don't see a place that Rydberg ever published it himself.
    … [English genus diagnosis not reproduced.] … [Name referring to resemblance of this genus to Picradenia.]  
    1. Picradeniopsis oppositifolia (Nutt.) Rydb. False Bahia. (I. F. f. 3967.) …. [English description not reproduced.] … On plains, S. Dak., Neb. and Mont. to Kans. and N. Mex. June-Sept. [Bahia oppositifolia Nutt.] Reference is to the “Illustrated Flora” figure 3967.

     

    Other articles:
    • Golden Checklist Flora:  Prenanthes racemosa;  

    Prenanthes racemosa Michx. “Purple Rattlesnake Root”

     

     

    Other articles:
    • Golden Checklist Flora:  Pseudognaphalium canescens;  

    Pseudognaphalium canescens (DC.) Anderberg “Wright's Rabbit-Tobacco”

     

    Literature Cited:
    - DeCandolle, Augustus Pyramus, 1836.  

    DeCandolle (1837, v. 6, p. 228) ...

    Original Text Translation or Comments
    40. G. canescens, totum villis sericeo-lanatis canum, canle herbaceo erecto ramoso, foliis lineari-lanoeolatis acuminatis , infer. basi attenuatis, super. breviter adnatis , capitulis ad apices ramulorum congestis sessilibus , involucre ovato-campanulati squamis scariosis albis lanceolatis acuminatis disco aequalibus. ♃ in Mexico ad Leonem ultra Guanaxuato legit cl. Mendez. Folia 1 ½ poll. longa, 2-3 lin. lata, patula. Inv. circ. 25-phyllum. Flor. herm. 5-7, foem. circ. 25. (v. s.) León, Guanajuato, Mexico collected by Mendez. Seen in the dried state by DeCandolle.

    Literature Cited:
    - Anderberg, Arne A., 1991.  

    Anderberg (1991) proposed moving G. canescens to Pseudognaphalium Kirp.

     

    Other articles:
    • Golden Checklist Flora:  Pseudognaphalium macounii;  

    Pseudognaphalium macounii (Greene) Kartesz “Macoun's Rabbit-Tobacco”

     

    Literature Cited:
    - Greene, Edward L., 1902b.  

    Greene (1902, v. 15, p. 278) published Gnaphalium maccounii

    Gnaphalium macounii. Apparently biennial, the stems rigidly erect, about two feet high, rather loosely leafy and clothed with a somewhat hirsute and viscid glandular-pubescence : leaves narrowly oblanceolate, acute, 3 inches long, the upper decurrent, all white-woolly beneath, light green and merely glandular-pubescent above : branches of the subpyramidal close panicle and the main stem for some distance below it densely white-woolly : involucres of middle size, their pearly scarious bracts all ovate, very acute : flower and fruit not seen.
    Collected in the Chilliwack Valley, B.C., 29 July, by Mr. Jas. M. Macoun, No. 26,847 ; also earlier at Revelstoke, No. 11,334, and again from the Warm Springs, Kootenay Lake, both in British Columbia, in the year 1890. No. 34,053 from Salmon Arm, J. R. Anderson, 1899, is also the same. The species is related to G. decurrens, yet very distinct in habit and inflorescence, the dense white-woolly pubescence of the upper part of stems and branches of the panicle being very peculiar.

     

    Other articles:
    • Golden Checklist Flora:  Ratibida columnifera;
    • Water Tank Road:   above the curve;
    • Field Notes:  Coll. No. 1198, 12 Jul 2015;
    Full Size ImageColl. No. 1198, Ratibida columnifera  

    Ratibida columnifera (Nutt.) Woot. & Standl. “Upright Prairie Coneflower”

    “Upright Prairie Coneflower” Ratibida columnifera is predominantly a Great Plains species which extends from southeastern British Columbia to Manitoba and Michigan, south through Illinois to Louisiana, and west through Texas and northern Mexico to Arizona.

    [So, is the cypsela ciliate on the abaxial side, or the adaxial side?]

       

    Literature Cited:
    - Fraser, John, 1813.
    - Greene, Edward L., 1889.

    Other articles:
    • Golden Checklist Flora:  Fraser's Catalogue, publication details;
    • Glossary:  flosculi;  fulvous;  

    Fraser's (1813) catalogue of new and interesting plants published for the first time a number of new taxa for North America. Many of these were Thomas Nuttall collections that were subsequently published in Pursh (1814) Flora Americae Septentrionalis and Nuttall (1818) Gen. Am.

    Original Text Comments

    75 * Rudbeckia columnifera. ‡ Spontaneous varieties of this plant sometimes occur with bright fulvous flowers, colored like Tagetes patula : the stem is simple, seldom producing more than three flowers, which are of an uncommon length, appearing like a column of flosculi, subtended by 5-8 neutral florets, and a simple calyx.

    Tagetes patula is commonly called the French Marigold, also in the Asteraceae, and native to Mexico and Guatemala.

    76 Rudbeckia purpurea. * serotina. ‡ Stem somewhat branching and hirsute, flowers brighter and more numerous.

    I think this might be what we now call Echinacea purpurea, not native to Colorado, but widely cultivated, and found occasionally as a garden escapee.
    * New Species. — † A Shrub. — ‡ Perennial. — M. from Missourie.  

    Copies of Fraser's (1813) catalogue were quite rare, so Greene (1889) republished it. It is unclear who actually wrote the catalogue. Greene was convinced that Nuttall wrote it having seen an original copy sent to Zaccheus Collins from Messrs. Fraser, saying

    It had passed through the hands of Nuttall, and had received one or two slight corrections from his pen. Moreover, he had written his name in ink, as the author of the Catalogue …

    Literature Cited:
    - Pursh, Frederick, 1816.

    Other articles:
    • Golden Checklist Flora:  Pursh, 1814, publication details;  

    Pursh (v. 2, 1816) published the name, citing Fraser's (1813) catalogue.

    Original Text
    657. RUDBECKIA. Gen. pl. 1324.
    11. R. caule stricto simplici summutate paucifloro, pedunculis elongatis, foliis pinnatifidis incisis, laciniis linearibus, calyce simplici 5-phyllo, radiis 5-8., disco cylindraceo elongato.
    R. columnifera, Fraser catal. 1813
    On the Missouri. v. s. The singular appearance of the receptacle of this plant distinguishes it from all the other known species.

    Literature Cited:
    - Fraser, John, 1813.
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Nuttall (1818) published Rudbeckia columnaris

    Original Text
    590. RUDBECKIA. L.
    Calix subequal, mostly consisting of a double series of leaflets. Receptacle paleaceous, conic. Pappus a 4-toothed margin.
    Herbaceous; leaves alternate, entire, lobed of pinnaifid ; flowers terminal ; disk often dark, rays yellow, rarely brown, in R. purpurea purple. — Stigma often obtuse.
    Species. ....
    12. columnaris. Hispid; stem nearly simple, 1 or few-flowered, peduncles very long; radical leaves nearly entire, cauline pinnatifid, segments linear-lanceolate; calix simple, 5 to 8-leaved, rays 5 to 8; disk cylindric, elongated. Hab. On the plains of Upper Louisiana. Flowering in July. Perennial: 1 to 2 feet high. Rays sometimes brown-red, as in Tagetes patula.
    13. ...
    A North American genus, with the exception of R. nudicaule of Monte Video, which appears to be scarcely distinct from R. spathulata. The seeds of R. purpurea are pungently aromatic.

    In this publication, Nuttal has altered R. columnifera to R. columnaris not referred to R. columnifera as published in Fraser's (1813) Catalogue. I suspect this might be enough to render R. columnaris and illegitimate name. If so, this will have an impact on Rafinesque's proposal of Ratibida columnaris Raf., nextunder.

    Literature Cited:
    - Rafinesque, C. S., 1818a.
    - Rafinesque, C. S., 1818b.

    Other articles:
    • Golden Checklist Flora:  Amer Monthly Mag Crit Rev, publication details;  

    Rafinesque (1818) wrote a two-part review of Pursh (1814) Flora Americae Septentrionalis.

    Original Text
    125. Rudbeckia columnaris must form the genus Ratibida of Raf. Fl. Miss.
    Ratibida columnaris Raf. is treated as a synonym of R. columnifera (Nutt.) Wooton & Standl.

    The reference to “Fl. Miss.“ is a puzzle. I have seen another reference to this publication that it was “ined.” However, it is unclear whether “Miss.” refers to either Mississippi or Missouri.

    Literature Cited:
    - Wooton, E. O., and Paul C. Standley, 1915.  

    Original Text
    2.Ratibida columnifera (Nutt.) Woot. & Standl.

    Rudbeckia columnifera Nutt. Fraser's Cat. No. 75. 1813.
    Rudbeckia columnaris Pursh, Fl. Amer. Sept. 575. 1814.
    Ratibida columnaris D. Don in Sweet, Brit. Flower Gard. II. 4: pl. 361. 1838
    Lepachys columnaris Torr. & Gray, Fl. N. Amer. 2:313. 1842.

    Type locality: Upper Louisiana.

    Range: British Columbia and Saskatchewan to Arizona, Texas, and Tennessee.

    New Mexico: Sierra Grande; mountains west of Grants Station; Santa Fe and Las Vegas mountains; Clayton; Lower Plaza; White and Sacramento mountains. Plains and low hills, in the Upper Sonoran and Transition zones.

    2a. Ratibida columnifera pulcherrima (DC.) Woot. & Standl.

    Obeliscaria pulcherrima DC. Prodr.5:559. 1836.
    Ratibida columnaris pulcherrima D. Don in Sweet, Brit. Flower Gard. II. 4: pl. 361. 1830
    Lepachys columnaris pulcherrima Torr. & Gray, Fl. N. Amer. 2: 313. 1842.

    Type locality: “In Mexici provinc. Texas as San-Fernando de Bejar, et in sinu Spiritus-Sancti ad lacum Sancti-Nicolai.”

    Range: With the species, but more common in New Mexico.

    New Mexico: Dulce; Chama; Pecos; Santa Antonita; Ramah; near Las Vegas; mountains west of Grants Station; El Cedro; Tucumcari; Mongollon Mountains; White Mountains; Buchanan; Redlands; Queen; Knowles; Artesia.

    This is a mere form of the type and hardly deserves a name. Both forms almost invariably occur together, although occasionally they grow alone, It is possible to find in a single patch every possible gradation in the color of the rays from pure bright yellow to solid brown-purple. The same variation in color occurs in R. tagetes, but since that has very small and inconspicuous rays no one has yet thought to distinguish the various forms by name.

    Literature Cited:
    - Reveal, James L., 1968.

    Other articles:
    • Golden Checklist Flora:  Reveal (1968) comments on Fraser (1813) Catalogue - 1;  

    Reveal (1968) summarized the evidence and opinions on the author of the names in Fraser's (1813) Catalogue and the legitimacy of them. Excerpts from Reveal's introduction were presented above, whereas his specific comments on Ratibida columnifera are below.

    75. Rudbeckia columnifera Nutt. In Fras. Catal. 1813. In my opinion, this species is adequately described in Fraser's Catalogue, as Nuttall states:
    “Spontaneous varieties of this plant sometimes occur with bright fulvous flowers, coloured like Tagetes patula: the stem is simple, seldom producing more than three flowers, which are of an uncommon length, appearing like a column of flosculi, subtended by 5-8 neutral florets, and a simple calyx.”
    This species was also described as R. columnaris Sims, Bot. Mag. 39: 1601. 1813, from cultivated plants given to Sims by the Fraser Brothers who had grown the species from seeds collected by Nuttall. The Pursh name, R. columnaris Pursh, Fl. Am. Sept. 2: 575. 1814, is an illegitimate substitute for Nuttall's R. columnifera and is thus an exact synonym of it. The species is now commonly known as Ratibida columnifera (Nutt. In Fras.) Woot. & Standl., Contr. U.S. Nat. Herb. 19: 706. 1915.

    Literature Cited:
    - Richards, Edward Leon, 1968.  

    Monograph on Ratibida

     

    Other articles:
    • Golden Checklist Flora:  Senecio eremophilus var. kingii;  

    Senecio eremophilus Richardson var. kingii (Rydb.) Greenm. “Cut-Leaved Groundsel”

     

    Literature Cited:
    - Rydberg, Per Axel, 1910b.  

    Rydberg (1910, v. 37, p. 467-8) published S. ambrosioides as a primarily Colorado species and S. kingii as a primarily Utah species.

    Original Text
    Senecio Kingii sp. nov.
    Senecio eremophilus D. C, Eat., Bot. King's Exp. 191. 1871. Not S. eremophilus Richards. 1823.
    Perennial, with a thick rootstock; stems glabrous, 3-6 dm. high, rather stout, leafy; leaves obovate or oblanceolate In outline, 4-7 cm. long, the lower petioled, all pinnately lobed one third to one half the distance to the midrib, with ovate or lanceolate, more or less toothed lobes; heads numerous, corymbose-paniculate, 9-11 mm. high; involucres glabrous, campanulate, 7-8 mm, high, 6-7 mm. broad; bracts linear, acute, carinate, sometimes with small black tips; the calyculate ones few, subulate; rays 5-7 mm. long; achenes scabrous-papillose on the rounded angles.
    This species is related to S. eremophilus, but differs in the somewhat smaller and erect heads, less deeply dissected leaves, and their broad and short divisions, and shorter rays. One of the specimens cited below was determined some years ago as S. glaucifolius, but that species differs from this as well as from the rest of the group in the narrower and scarcely carinate bracts.
    Utah: Cottonwood Canon, Aug. 1869, S. Watson 676 (type, in herb. Columbia University); Alta, Wahsatch Mountains, 1879, M. E. Jones 1144; American Fork Canon, July 1895, M. E. Jones; Big Cottonwood Canon, Aug. 1905, Garrett 1591; near Marysvale, 1905, Rydberg & Carlton 7068; Mount Barrette, 7206; Fish Lake, 7206.

     

    Other articles:
    • Golden Checklist Flora:  Senecio integerrimus;  

    Senecio integerrimus Nutt. “Columbia Ragwort”

     

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;
    • Glossary:  carnose;  sphacelate;  

    Original Text Comments
    564. SENECIO. L. (Ragwort.)  
    Calix cylindric, subcaliculate: scales sphacelate at the point. Receptacle naked. Pappus simple, capillary, and copious. sphacelate = undergo necrosis.
    Suffruticose or more commonly herbaceous; leaves entire or pinnatifid; flowers mostly corymbose or terminal; yellow or rarely purple. A few species are destitute of rays.  
    Species. 1. …  
    15. * integerrimis Smooth; stem simple and attenuated; leaves perfectly entire; radical ones long petiolate, lanceolate, acute, cauline, sessile, acuminate, uppermost minute; corymb simple, 8 to 12-flowered? peduncles 1-flowered, rays shorter than the hemisperical caliculate calix. HAB. In depressed and moist situations on the plains of the Missouri, near the Great Bend. Flowering in June. Flower large and yellow. Stem 12 to 18 inches high. Lower leaves thickish and somewhat carnose, very smooth, uppermost minutes, slightly tomentose; corymb coarctate. Seeds smooth. Nearly allied to S. aquaticus. carnose = of a fleshy consistence —used of succulent parts of plants.

    coarctate = crowded together.

    A genus of more than 140 species principally indigenous to Europe and the Cape of Good Hope.  

     

    Other articles:
    • Golden Checklist Flora:  Senecio riddellii;  

    Senecio riddellii (Torr. & A. Gray) Greenm. ex L.O. Williams “Riddell's Ragwort”

     

     

    Other articles:
    • Golden Checklist Flora:  Senecio spartioides;
    • Field Notes:  6 Sep 2020;

    Locations: Schweich Hill.
    Full Size ImageColl. No. 2231, Senecio spartioides with a praying mantis and a beetle.  

    Senecio spartioides Torr. & A. Gray. “Broomlike Ragwort”

    Known from Apex Park and Tin Cup Ridge, North and South Table Mountains, and Schweich Hill.

    Full Size Image
    Senecio spartioides adventive in my garden.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1838-1843.

    Other articles:
    • US Highway 287:   at Jeffrey City;

    Locations: Jeffrey City.  

    Senecio spartioides was described by Torrey and A. Gray (1843) from a Lt. John Fremont collection.

    Original Text
    * * * * Perennial : heads corymbose, chiefly radiate
    † Leaves entire or denticulate.
    8. S. spartioides : glabrous throughout; stems suffruticose, very numerous from the same ligneous tap-root, rigid, corymbose at the summit, leafy; leaves fleshy, narrowly linear, perfectly entire, rather obtuse, sessile; heads (large and showy) fastigiate-corymbose, on short minutely bracteolate peduncles ; the calyculate scales subulate, minute ; scales of the cylindrical involucre about 12, lanceolate-linear, acutish; rays mostly 7, oblong-linear, elongated ; achenia silky-canescent.
    Upper Platte; on a steep sand-bank of the Sweet-water River, Lieut. Fremont! Aug.–Sept. — Stems a foot high, forming a dense tuft. Leaves 1-3 inches long, about a line wide, very numerous. Heads half an inch in length. Rays golden-yellow. Pappus as long as the disk-corolla. — A remarkable and handsome species.

     

    Other articles:
    • Golden Checklist Flora:  Senecio wootonii;  

    Senecio wootonii Greene “Wooton's Ragwort”

     

       

    Solidago L. “Goldenrod”

     

    Literature Cited:
    - Semple, J. C., 2018.  

    Semple (2018) is a frequently updated resource on Goldenrods (Solidago sp.).

     

    Other articles:
    • Golden Checklist Flora:  Solidago canadensis;  

    Solidago canadensis L. “Canada Goldenrod”

     

     

    Other articles:
    • Golden Checklist Flora:  Solidago gigantea;  

    Solidago gigantea Aiton “Giant Goldenrod”

     
     

    Aiton (1789, v. 3, p. 211) published Solidago gigantea from plants grown at the Physick Garden at Chelsea.

    Original Text Comments
    4. S. caule erecto glabro, foliis lanceolatis glabris ferratis margine feabris, racemis paniculatis fecundis, pedunculis hirtis, ligulis abbreviatis. gigantea. 4. Solidago smooth stiff stems, leaves are smooth, margins serrate; panicles with clusters of flowers; peduncles rough, ligules short.
    Gigantic Golden-rod.    
    Nat. of North America.    
    Cult. 1758, by Mr. Philip Miller.   Mr. Philip Miller was the curator at the Physick Garden at Chelsea.
    Auguft and September. H. ♃   H. – Hardy.
    ♃. – Perennial.

     

    Other articles:
    • Golden Checklist Flora:  Solidago missouriensis;
    Full Size ImageHabitat of Coll. No. 1256, Solidago missouriensis at Ranson/Edwards.
    Full Size ImageColl. No. 1732, Solidago missouriensis at North Table Mountain.  

    Solidago missouriensis Nutt. “Missouri Goldenrod”

     

    Literature Cited:
    - Nuttall, Thomas, 1834a.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1834a, publication details;  

    Nuttall (1834a, p. 32-33) published Solidago missouriensis

    Original Text
    55. Solidago * Missouriensis. Pumila, glabra, recemis erectis, foliis lineari-lanceolatis, acutis, inciso-subserrularis, superioribus integris, panicula brevi laxa, floribus majus-culis
    Stem slender, smooth, leafy, about a foot or so high. Leaves scabrous at the margin. Panicle about three inches long, the branchlets slender, the flowers pedicellate, and brought together in a somewhat rhomboidal raceme. Rays as long as the calyx.
    Hab. in the upper branches of the Missouri and in Arkansas.
    Even though this was described with collections by Wyeth, the locations noted seem much more like Nuttall collections from his 1811 trip on the Missouri River and 1819 trip to Arkansas. Wyeth collections would have been along the Platte, Snake, or Columbia.

     

    Other articles:
    • Golden Checklist Flora:  Solidago nana;  

    Solidago nana Nutt. “Baby Goldenrod”

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1841, publication details;  Crepis occidentalis, Weber & Wittman, 2012;
    Full Size ImageColl. No. 2439, Solidago nana.  

    Nuttall's (1841) description of S. nana.

    Original Text
    Solidago * nana; somewhat cinereous and pulveulently pubescent, dwarf, many stems from the same root; lower and radical leaves spathulate, obtuse, entire, or subserrulate at the apex, stem leaves linear, narrowed below; ramuli fastigiate, subcorymbose; bractes linear; involucrum nearly smooth, scales ovate; rays about seven, oblong, as long as the disk; achenium pubescent.
    HAB. In the Rocky Mountain range, near Lewis' River of the Shoshonee. About a span high, with a large, black, almost woody root. Stem leaves small, radical ones about one and a half inches by half an inch wide; scales of the involucrum unusually broad, pubescent on the margin, rays conspicuous. Apparently allied to S. nemoralis, though very distinct and alpine.

     

    Other articles:
    • Golden Checklist Flora:  Solidago nemoralis ssp. decemflora;  

    Solidago nemoralis Aiton ssp. decemflora (de Candolle) Brammall ex Semple “Gray Goldenrod”

     
     

    DeCandolle (1836, v. 5, p. 332) ...

    Original Text Translation
    15. S. decemflora, caule erecto tereti foliisque pube brevissima subscabris, foliis oblongis mucronulatis integerrimis subtriplirierviis , radicalibus basi attenuatis, racemis subsecundis in paniculam corymbosam dispositis, invol. glabri squamis linearibus , capitulis 10-floris, ligulis 5 minutissimis. ♃ in Mexicanae prov. Texas districtibus orientalibus legit cl. Berlandier (pl. exs. n. 1924). Achaenia sericeo-villosa. Herba pedalis. (v. s.) 15. Solidago decemflora stems erect, leaves rounded and short scabris; oblong leaves mucronulate entire subtri-nerved; Radical base attenuate, corymbose panicle, subsecundis in the clusters have been arranged, involved, smooth, linear phyllaries, heads 10-flowered, ligules 5 minute. ♃ (perennial) In Mexican Prov. East Texas, eastern districts per the collector Berlandier (Pl. Exs. N. 1924). Achaenia silky-villous. The plant a foot tall. (Seen in the dried condition.)

     

    Other articles:
    • Golden Checklist Flora:  Solidago simplex;  

    Solidago simplex Kunth “Sticky Goldenrod”

     

    Literature Cited:
    - Kunth, Carol Sigismund, 1815-1825.  

    Kunth (1818, t. 4(15), p. 81) ....

    Original Text Translation and Comment
    1. SOLIDAGO SIMPLEX. †  
    S. caule erecto, simplici, subangulato, striato, superne hirtello; foliis lineari-lanceolatis, superne obsolete crenatis, inferne valde angustatis, glabris , margine denticulato-scabris ; panicula terminali, ramis subtrifloris.  
    Crescit prope pagum Sanctce Rosae Mexicanorum , alt. 1300 hex. ♃ Floret Septembri. It grows near the Mexican village called Santa Rosa, altitude 1300 hex. perennial. Flowers in September.
    Radix crassa, perpendicularis, fibris crebris obsita. Caulis erectus, i 4 — i 5 -pollicaris , simplex, striatus, subangulatus, puiqrarascens, superne hirtellus. Folia radicalia sessilia , lineari-laneeolata , acutiuscula , inferne valde angustata , adeo ut petiolata appareant, subintegerrima aut apicem versus remote et obsolete crenata, reticulato-venosa, crassiuscula, glabra, margine denticulis minutissimis scabra, 26—27 lineas longa , tres lineas lata. Folia caulina alterna , radicalibus simillima, magis tamen acuta , summa subacuminata. Panicula terminalis, subramosa , bracteata5 ramis subternis, patulis, subtrifloris, angulatis , hirtellis; floribus pedicellatis , magnitudine floris Solidaginis Virgaureae; bracteis linearibus. Involucrum campanulato-hemisphaerieum; foliolis circiter 25 , laxe imbricatis , linearibus, acutis , planis , uninerviis , margine scariosis et diaphanis , glabris, nitentibus; exterioribus brevioribus. Receptaculum convexum, scrobiculatum, glabrum. Flosculi disci circiter 20, tubulosi hermaphroditi; radii 8— g, ligulati, feminei. Flosculi hermaphroditi : Corolla flava, glabra, superne ampliata , quinquefida ; laciniis lineari-lanceolatis , acutis, enerviis , patulis. Stamina tubo corollae inserta. Antherae subinclusae. Ovarium lineare , hispidum. Stylus glaber. Stigma bipartitum, exsertum; laciniis patentibus , glabriusculis. Akenium immaturum. Pappus pilosus, sessilis; pilis scabris, apice vix incrassatis, albis, corollam subsequantibus, patulis. Flosculi feminei: Corolla flava ; tubo tenui, glabro; ligula lanceolato-oblonga, tridentata, plana, patente , sesquilineam longa. Ovarium, Stylus, Stigma et Pappus prorsus ut in flosculis hermaphroditis. Pappus tubo paulo longior.  
    Solidagini Virgaureae habitu affinis. Solidago virgaurea has similar habitat.

     

    Other articles:
    • Golden Checklist Flora:  Solidago speciosa;  

    Solidago speciosa Nutt. “Showy Goldenrod”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Original Text
    560. SOLIDAGO. L. (Golden-Rod.)
    ...
    †† Racemes erect
    27. *speciosa. Stem tall and smooth, simple or virgately branched; leaves lanceolate, entire, somewhat carnose. scabrous on the margin, the lower very broad, radical ones subserrate; racemes mostly terminal, erect and compound, pubescent; pecuncles mostly shorter than the calix; rays elongated about 5; seed smooth. Hab. In shady woods, on the banks of the Schuykill, also in Jersey; near Philadelphia, but rare. S. sempervirens. Mich. S. integrifolia? Persoon, 2. p. 449. Allied to S. petiolaris. Stem often 6 feet high, smooth and sulcate. Lowest leaves a span long, and 3 inches broad, irregularly and remotely subserrate, upper leaves very entire. gradually diminishing upwards, in dry and shady situations, membranacei=ous and veined, in gardens subcarnose and smaller, with the veins partly obliterated, racemes also numerous, but always rigid, terminal and erect. Flowers larger than the preceding, with the calix also coloured (which in the preceding is green); rays bright yellow, unusually broad. The seeds in this species are perfectly smooth, in our sempervirens pubescent. This is one of the most ornamental plants of the genus.

     

    Other articles:
    • Golden Checklist Flora:  Solidago velutina ssp. sparsiflora;  

    Solidago velutina DC. ssp. sparsiflora (A. Gray) Semple. “Three-Nerve Goldenrod”

     

    Literature Cited:
    - Gray, Asa, 1876a.
    - Wheeler, George M., 1878.  

    Gray (1876, p. 58-59) publisher S. sparsiflora from a collection on the Wheeler Expedition.

    Solidago sparsiflora. Virgaurea, Virgatae : scabrido-puberula ; foliis inferioribus ignotis, su; erioribus floralibusque parvulis lanceolatis (lin. 6-12 longis); racemulis oligocephalis laxis laxeque thyrsoideis ; involucri squamis linearibus puberulis apice viridulis acutiusculis ; floribus radii circiter 10 ligulis parvulis, disci 4-5 ; acheniis sericeo-pubescentibus. — Arizona, near Camp Lowell, Sept., 1874. Rothrock, in Wheeler Expedition, 1874.

     

    Other articles:
    • Golden Checklist Flora:  Stephanomeria pauciflora;  Notes on Lygodesmia juncea;  

    Stephanomeria pauciflora (Torr.) A. Nelson “Brownplume Wirelettuce”

     

    Literature Cited:
    - Torrey, John G., 1828.  

    Torrey (1827, p. 210) published several of Dr. James collections, placing them all tentatively in Prenanthes.

    Original Text Comments
    202. Prenanthes juncea, Pursh fl. ii. p. 498. Nutt. gen. ii. p. 123. On the Platte and Missouri. This is a synonym of Lygodesmia juncea (Pursh) D. Don ex Hooker
    Obs. Near the Rocky Mountains, Dr. James collected two species of a genus allied to Prenanthes, but differing in their plumose and sessile pappus. They will probably hereafter be referred to a new genus. Their principal characters are as follows:  
    1. P.? pauciflora, caule ramoso, flexuoso, superne paniculato, ramulis unifloris, floribus erectis ; foliis lineari-lanceolatis, runcinatis, glabris ; calycibus quinquefloris. This will become Stephanomeria pauciflora (Torr.) A.Nelson in J.M.Coult. & A.Nelson, New Man. Bot. Centr. Rocky Mt. 588 (1909)
    2. P.? tenuifolia, caule ramoso, glabro ; foliis linearibus, integerrimis; panicula laxa terminali; floribus erectis; calycibus quinquefloris. This is a synonym of Stephanomeria tenuifolia (Torr.) H.M.Hall, of which most Colorado collections are on the western slope.

    Literature Cited:
    - Coulter, John M., and Aven Nelson, 1909.  

    Coulter & Nelson (1909, p. 588) placed P. pauciflora Torr. in Stephanomeria

    Original Text Comments
    2. Stephanomeria pauciflora (Torr.) Nels. Similar to the preceding, usually lower, smaller in all of its parts, and somewhat woody at the base: the lower leaves at least more or less runcinate-pinnatifid: pappus plumose nearly to the base. S. runcinata in part. (Ptiloria pauciflora Raf. Atl. Journ. 145. 1832.) Sparingly if at all in our range; on the plains; from Nebraska to Texas. The preceding was S. tenuifolia (Torr.) Hall.

     

    Other articles:
    • Golden Checklist Flora:  Symphyotrichum ericoides;  Dill. elt., 1732;  Gron. virg., 1738;  Roy. lugdb., 1740;  

    Symphyotrichum ericoides (Linnaeus) G. L. Nesom “White Heath Aster”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 875) ...

    Original Text Translation and Comments
    16. ASTER caule paniculato, pedunculis racemofis : pedicellis foliofis : foliolis linearibus integerrimus. Gron. virg. 100. ericoides 16. ASTER stem paniculate, peduncles racemose : pedicels leafy : leaves linear entire.

    Flora Virginica, published as two volumes by Jan Frederik Gronovius in 1739 and 1743, based on the work of colonial Virginia botanist John Clayton.

    After caule paniculato, floribus racemofis, calycibus patulis oblongis, foliis fubulato-linearibus. Roy. lugdb. 168.   Aster with paniculate stems, flowers recemose, calyx wide oblong, leaves subulate-linear.

    Flora Leydensus Prodromus, published by Adriaan Royen in 1740. Royen was a professor of botany at Leyden and well aquainted with both Cliffort and Linnaeus.

    After ericoides dumofus. Dill. elt. 40. t. 36. f. 40.   Aster ericoides dumosus.

    Johan Jacob Dillenii, 1732. Hortus Elthamensis, a catalogue of the rare plants growing at Eltham, London.

    Habitat in America feptentrionalis. ♃   Lives in North America.

    Literature Cited:
    - Nesom, Guy L., 1994.  

     

     

    Other articles:
    • Golden Checklist Flora:  Symphyotrichum falcatum;  

    Symphyotrichum falcatum (Lindl.) G.L. Nesom “White Prairie Aster”

     

     

    Other articles:
    • Golden Checklist Flora:  Symphyotrichum laeve;  

    Symphyotrichum laeve (L.) Á. Löve & D. Löve var. geyeri (A. Gray) G.L. Nesom “Smooth Blue Aster”

     

     

    Other articles:
    • Golden Checklist Flora:  Symphyotrichum lanceolatum ssp. hesperium;  

    Symphyotrichum lanceolatum (Willd.) G. L. Nesom ssp. hesperium (A. Gray) G. L. Nesom Western Lance-Leaved Aster

     

    Literature Cited:
    - Gray, Asa, 1884.  

    Gray (1884, v. 1, pt. 2, p. 192) ...

    Original Text
    A. hesperius. Resembles A. paniculatus and A. salicifolius of the East, equally variable, from nearly glabrous and smooth to scabrous-pubescent . leaves lanceolate, entire or the larger with a few denticulations (2 to 5 inches long, 3 to 8 lines wide) : heads rather crowded, 4 or 5 lines high : involucre of narrowly linear or more attenuate acute or gradually acuminate erect bracts, either unequal and imbricated, or with some loose and slender herbaceous exterior ones which equal the inner : rays either white or violet, 3 or 4 lines long. — Damp soil and along streams, S. Colorado and New Mexico to Arizona and S. California. Has been variously taken for A. longifolius, Novi-Belgii, aestivus, &c., and coll. by Wright, Greene, Rothrock, Cleveland, Parish, Lemmon, &c.

     

    Other articles:
    • Golden Checklist Flora:  Symphyotrichum porteri;  

    Symphyotrichum porteri (A. Gray) G.L. Nesom “Smooth White Aster”

     

    Literature Cited:
    - Porter, Thomas C., and John M. Coulter, 1874.  

    Publication of Aster ericoides L. var strictus Porter in Porter and Coulter (1874, p. 56) ...

    Original Text
    Aster ericoides, L., var. strictus, Porter. Low, ¾°-1° high, glabrous, except the scabrous margins and ciliate bases of the leaves, erect, slender, paniculately branched above, branches short; scales of the involucre narrowly linear, lax, outer ones very acute, often entirely green, inner ones scarious with a central green line; radical leaves narrowly oblanceolate. — &ldquo';In the mountains at middle elevations,” Hall & Harbour, 254. Near Denver, Coulter. Foot-hills west of Denver, Porter; Meehan; Hoopes.

    Literature Cited:
    - Gray, Asa, 1880.  

    The basionym: Asteraceae Aster porteri A.Gray Proc. Amer. Acad. Arts xvi. (1880, p. 99) ...

    Among the true Asters are several forms which have to be named, such as A. Porteri for A. ericoides, var. strictus, Porter & Coult. Fl. Colorad. 56, …

     

    Other articles:
    • Golden Checklist Flora:  Thelesperma megapotamicum;  

    Thelesperma megapotamicum (Spreng.) Kuntze “Hopi Tea Greenthread”

     

    Literature Cited:
    - Sprengel, Curt Polycarp Joachim, 1826.  

    Sprengel (1826, v. 3, p. 454) ...

    Original Text Translation and Comments
    454 Syngenesia. II. Eupatorinae. Bidens 454 Syngenesia. II. Eupatorinae. Bidens
    megapotamica* 33. B. foliis omnibus 2pinnatifidis lineari-filiformis glabris, floribus fubgeminis pedunculatis erectis difcoideis, involucro colorado anthodium aequante. Ad fl. magnum Amer. auftr. (Rio grande. Sello.) megapotamica* 33. Bidens leaves all 2-pinnatifid, linear-filiform, glabrous, peduncle erect, heads equal, sub-budded (refers to calyculi??), involucre reddish-brown, flowers discoid. On large rivers of the American west. (Rio grande. Sello.)

    Literature Cited:
    - Kuntze, Otto, 1891-1898.  

    Kuntze (1898, Pt. 3[3], p. 182) ...

    Original Text
    Thelespermum megapotamicum OK. (Bidens m. Spr. 1826 — Th scabiosodes Less. 1831.) Argentina: Villa Merceded, Ceres.

    Plants of the World (Kew) does not accept Kuntze's authorship of this name, instead accepting Thelesperma megapotamicum (Spreng.) Herter, Revista Sudamer. Bot. 7: 235 (1943). I am unable to find that publication online.

     

    Other articles:
    • Golden Checklist Flora:  Townsendia exscapa;  

    Townsendia exscapa (Richards.) Porter “Stemless Townsend Daisy”

     

    Literature Cited:
    - Hooker, Sir William Jackson, 1829-1840.
    Full Size ImageHooker (1834), Tab. CXIX, Townsendia exscapa  

    Hooker (1834,v. 2. p. 16) described Townsendia and T. sericea that will become a synonym of T. exscapa.

    Original Text
    58. TOWNSENDIA. Nov. Gen.
    Capitulum heteroganium. Flosculi radii ligulati foeminei; disci hermaphroditii regulares. Involucri foliola plurisermlia imbricata. Pappus uniserialis; radii brevis, paleaceo-subulatus, inaequalis; disci pilosus, pilis elongatis scabriusculis. — Herba Americae borealis humilis acaulis. Radix fusiformis, lignosa, longe descendens ; superne saepe divisa, reliquiis foliorum vetustorum obtecta. Folia omnino radicalia, numerosa, erecto-patentia, linearia, subspathulata, integerrima, utrinque adpresso-sericea, basi dilatata, et utrinque ciliata. Flos solitarius, foliis immersus et obtectus, ratione plantae magnus. Involucrum ovatum, foliolis sericeis, lineari-subulatis, erectis, imbricatis. Flosculi radii ligulati, erecti, pallide rosei, marginibus involuti, apice dentibus tribus conniventibus. Stigma bifidum, purpureum, glabrum. Ovarium oblongum, compresso-triquetrum, pubescens. Pappus brevis sed valde inaequalis, uniierialis, e pilis seu paleis subvlatis, basi membranaceis. Flosculi disci tubulosi, lutei. Pappus pilosus, pilis scabriusculis longitudine corollae. Ovarium sericeum. Receptaculum alveolato-punctatum, nudum.
    1. T. sericea. (Tab. CXIX), Aster? exscapus. Rich, in Frankl. 1st. Journ. ed. 2. App. p. 32.
    Hab. Carlton House upon the Saskatchawan. Dr. Richardson. Rare, upon the dry banks of the Saskatchawan and among the Rocky Mountains. Drummond. — This highly interesting little plant, no less on account of its habit than its pappus, deserves to be separated from Aster, of which, indeed, it was by Dr. Richardson, considered a doubtful species. “the bud is formed in the autumn,” and what Dr. Richardson further observed in the living plant, I find to be characteristic of all the specimens in this Collection, that “the florets of the ray are mostly involute, rarely expanded, and always narrow, nearly of the same colour with the pappus and inconspicuous; the flowers, indeed, n ever fully expanding.” I have named the Genus in compliment to David Townsend, Esq. of West Chester, Pennsylvania, who having imbibed the most ardent love of Botany from his friend and instructor Dr. Darlington of the same city, has devoted his leisure hours to the science with eminent success. The plant now under consideration is peculiarly worthy of bearing his name, because he has studied and ably discriminated the numerous Pennsylvanian species of the allied Genus Aster.
    Tab. CXIX. — Fig. 1, Flowers ; fig. 2, Floret of the ray; fig. 3, Portion of the pappus from the ray; fig. 4, Floret of the disk; fig. 5, Portion of a hair from the pappus of the disk; fig. 6, Stigma from the disk; fig. 7, 7, Scales of the involucre; fig. 8: — all more or less magnified.

       

    Townsendia grandiflora Nutt. “Largeflower Townsend Daisy”

     

    Other articles:
    • Golden Checklist Flora:  Townsendia grandiflora;  

    (Nuttall, 1840, p. 306) ...

    Original Text Comments
    Townsendia * grandiflora; stem canescent, divaricately branching from the base, branches one or few-flowered, leaves linear-sublanceolate, very acute, nearly smooth, or minutely pubescent, green ; capitulum hemispherical ; involucrum of three series, the sepals lanceolate, filiformly acuminate, minutely fringed ; rays twenty-eight to thirty, or more, bidentate.  
    Hab. With the preceding, which it resembles wholly in habit, but with the flower as large nearly as that of the China Aster, (Callistephus Chinensis) Branching from the base, and spreading out sometimes from six to ten inches along the ground, Leaves linear, much attenuated below, and very acute, when green rather succulent, and appearing smooth, though somewhat pubescent beneath, (seen through a glass.) Sepals elegantly imbricated, perfectly lanceolate, much acuminated, scariose, except the centre, which is green, the margin minutely lacerate-ciliate. Rays pale lilac, longer than the disk. — A plant which well deserves cultivation, from its large, showy flowers. The prededing was Townsendia strigosa which was found “On the Black Hills, (or eastern chain of the Rocky Mountains,) near the banks of the Platte.” In Colorado, it is found only in westernmost counties.

     

    Other articles:
    • Golden Checklist Flora:  Townsendia hookeri;  

    Townsendia hookeri Beaman “Hooker's Townsend Daisy”

     

    Literature Cited:
    - Nuttall, Thomas, 1840-1841.  

    Nuttall (1840-1841, p. 304-305) describes a collection as T. sericea which is later to be determined T. hookeri

    Original Text
    TOWNSENDIA. (Hooker.)
    Townsendia sericea; caespitose; leaves narrow linear, acute, scarcely half a line wide, canescently sericeous; capituli sessile on the caudex: scales of the involucrum numerous, very narrow and acuminate. — Achenium as in the rest of the genus, obovate, margined, and flatly compressed, sericeous, with a numerous connate series of white, silky pappus, almost plumosely barbellate, and remarkably attenuated above.
    Hab. On the Black Hills, (an alpine chain toward the sources of the Platte.) Flowering probably in April. By the achenium, this genus makes some approach to Calimeris, though totally unlike in habit.
    Nuttall goes on the describe Townsendia incana that he describes as with the above.

    Literature Cited:
    - Beaman, John H., 1957.  

    Beaman (1957, p. 95-100), in a six-page section described Townsendia hookeri from an Ira W. Clokey collection in Mount Vernon Canyon.

    10. Townsendia Hookeri Beaman, sp. nov.
    Type: Clokey 4338, dry hills, Mt. Vernon Canon, 1730 m. alt., Jefferson Co., Colorado, 13 April 1920 (*COLO 12061, holotype; CAN, CAS, DS, F, GH, MICH, MO, MONTU, NA, PH, POM, *RM, UC, US, UTC, WS, WTU, isotypes).

    This seems to be the only collection made by Ira W. Clokey on 14 April 1920, indeed the only collection Mr. Clokey made in the week from 10 April to 16 April 1920. Clokey also collected a large number of duplicates, i.e. 18 duplicates according to Beaman (1957). SEINet reveals 3 more duplicates, at IND, NO, and MIN.

     

    Other articles:
    • Golden Checklist Flora:  Xanthisma spinulosum;
    Full Size ImageColl. No. 2438, Xanthisma spinulosum  

    Xanthisma spinulosum (Pursh) D. R. Morgan & R. L. Hartm. “Spiny Goldenweed”

     

    Literature Cited:
    - Nuttall, Thomas, 1813.
    Full Size ImageColl. No. 2434, Xanthisma spinulosum  

    Nuttall (1813) may have been first into print with:

    81 * [Sideranthus] pinnatifidus. Fl. of both these species yellow, and considerably like those of the genus Aster, but from which they are sufficiently distinct. M

    Unfortunately, Sideranthus was a nomen nudum when this was published, as the name was not validly published until 1840.

    Literature Cited:
    - Pursh, Frederick, 1814.
    Full Size ImageDetail of Coll. No. 2434, Xanthisma spinulosum  

    Pursh (1814, v. 2, p. 564) ...

    Original Text
    650. AMELLUS. Gen. pl. 131 5.
    ... ...
    spinulosus. 2. A. canescens ; foliis bipinnatifidis inciso-dentatis, laciniis linearibus rigido-mucronatis, floribus lateralibus et terminalibus congestis, radiis bidentatis, paleis setaceis.

    In open prairies on the Missouri. M. Lewis. ♃. Aug. Sept. v. s. in Herb. Lewis. Rays yellow.

    Tota planta rigida, scabra, cano-viridescens. Caulis ramosissimus. Rami angulosi. Ramuli laterales breves, uniflori. Folia alterna, bipinnatifida, rigida, laciniis linearibus, spinula terminatis, inferioribus inciso-dentatis. Flores laterales solitarii et terminales congesti, subcorymbosi, lutei. Calyx imbricatus : squamis exterioribus brevioribus, acutis, hirsutis ; interioribus linearibus, scariosis. Corollulae radii lineari-lanceolatae, bidentatae. Receptaculum : Paleis brevibus, subuiatis.

    Both species are an intermediate link between Aster and Inula. They approach the nearest to Amellus ; but the receptacle is not, properly speaking, paleaceous, but only setaceous.

    Literature Cited:
    - Moulton, Gary E., 1999.  

    Moulton (1999) lists the Lewis & Clark collection as Macheranthera pinnatifida

    Literature Cited:
    - Morgan, David R., and Ronald L. Hartman, 2003.  

    Morgan & Hartman (2003, v. 20, p. 1406) ...

    14. Xanthisma spinulosum (Pursh) D. R. Morgan & R. L. Hartman, comb. nov. Basionym: Amellus spinulosus Pursh, Fl. Amer. Sept. 2:564. 1813. … Type: U.S.A. [South Dakota. Lyman or Brule Co.: Reveal et al. 1999], in open prairies on the Missouri, 15 Sep 1804, M. Lewis s.n. (Lectotype: PH! [designated by Hall 1928]; isolectotype: UC!).
    The following two binomials are not valid as the genus Sideranthus was a nomen nudum when the combinations were published: Sideranthus pinnatifidus Nutt., Fraser's Catalogue, no. 81, 1813, non Aplopappus pinnatifidus Nutt., 1840 and Sideranthus spinulosus (Pursh) Sweet, Hort. Brit., 227. 1826.

     

    Other articles:
    • Golden Checklist Flora:  Euphorbia dentata;  

    Euphorbia dentata Michx. “Toothed Spurge”

     

    Literature Cited:
    - Ackerfield, Jennifer, 2015.
    - Michaux, Andre, 1803.
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Described by Michaux (1803).

    Original Text
    dentata. E. annua; pumila, hirsuta : foliis oppositis , ovalibus , dentatis : floribus ad summitates congestis.
    Obs. Folia superiora maculosa.
    Hab. in Tennasee , juxta Nashville.
    Full Size Image
    Coll. No. 2680, Euphorbia dentata.

       

    Native Monocots in the Golden Flora

     

     

    Other articles:
    • Golden Checklist Flora:  Typha angustifolia;  

    Typha angustifolia L. “Narrow Leaf Cattail”

     

     

    Other articles:
    • Golden Checklist Flora:  Typha latifolia;  

    Typha latifolia L. “Cattail”

     

     

    Other articles:
    • Golden Checklist Flora:  Zannichellia palustris;  

    Zannichellia palustris L. “Horned Pondweed”

     

     

    Other articles:
    • Golden Checklist Flora:  Potamogeton nodosus;  

    Potamogeton nodosus Poir. “Longleaf Pondweed”

     
       

     

    Other articles:
    • Golden Checklist Flora:  Agropyron cristatum;  

    Agropyron cristatum (L.) Gaertn. “Crested Wheat Grass”

     

     

    Other articles:
    • Golden Checklist Flora:  Alopecurus aequalis;  

    Alopecurus aequalis Sobol. “Shortawn Foxtail.”

     

    Literature Cited:
    - Sobolewski, Gregor, 1799.  

    Sobolewski (1799, p. 16) ...

    Original Text Translation
    45. ALOPECURUS AEQUALIS. Ariftis gluma aequalibus (Sob.)

    Habitat in locis uliginosis cum praecedente. Floret Junio, Julio. In lacubus natans eft. ♃. Sobol.

    45. Alopecurus aequalis. Glume awns equal. (i.e., glumes equal).

    Lives in a damp place with the first, i.e., A. geniculatis. Flowers June, July. Swimming in lakes. Perennial. [Belarusian] Fox horsetail is smooth. Sobolewski.

     

    Other articles:
    • Golden Checklist Flora:  Andropogon gerardii;  

    Andropogon gerardii Vitman. “Big Bluestem”

    The native range is North America from Utah to the east, and from Saskachewan south to Costa Rica (POWO, 2021). How it got the the Geneva Botanic Garden is unexplained.

    Literature Cited:
    - Vitman, Fulgenzio Antonio Maris, 1789-1792.  

    Original Text Interpreted Latin Interpreted English Comments
    1365. ANDROPOGON. Hermaphr. Cal. gluma 1-fl. Cor. gluma bafi ariftata . Stam 3. Styl. 2. Sem. 1. follic. involutum , ariftatum .

    Maf. Cal. Cor. Stam. prioris .

    1365. ANDROPOGON. Hermaphroditic. Calyce gluma 1-floris. corolla gluma basi aristata. Stamina 3. Stylis 1. Folliculus involutum, aristatum.

    Maf. calyce corolla stamina prioris .

    1365. ANDROPOGON. Hermaphroditic. Calyx glume single-flowered, corolla glume (i.e., lemma) bristled from base (???) Stamens 3. Style 1. Follicles rolled, bristled.

    Maf. calyx of the corolla less than the stamens.

    Where on the lemma does the awn arise???

    What does “Maf.” mean?

    ...        
    Gerardi. A. fpicis digitatis : fl. alterne geminis ; hermaphrodito fefilli , ariftato ; mafculo pedunculato , mutico . Gerard. gallo-prov. cum. fig.

    In Gallo provincia.

    Gerardi.

    Andropogon spicis digitatis: floris alterne geminis; hermaphrodito sesilli, aristato, masculo pedunculato, mutico.
    Gerard. Flora Gallo-Provincialis cum figura.

    In Gallo provincia.

    Gerardi. Andropogon with spikes digitate, flowers alternately twinned, the sessile flower is hermaphroditic and bristled, the pedunculate flower is masculine and pointless.
    [Published in] Gerard, 1760. Flora Gallo-Provincialis with figure. In Provence.

    The location literally translates to “In Provence,” but I think it might really mean “In the French provinces [of Canada.]”

    The honoree's name is Gérard, Louis, 1760, author of Flora Gallo-Provincialis. Parisiis : Ad Ripam PP. Augustinorum, Apud C. J. B. Bauche, Bibliopolam, Ad insigne St.ae Genovefae, & S.ti Joannis in deserto 1761

     

    Other articles:
    • Golden Checklist Flora:  Apera interrupta;  

    Apera interrupta (L.) Beauv. “Italian Windgrass”

     

     

    Other articles:
    • Golden Checklist Flora:  Aristida purpurea;  

    Aristida purpurea Nutt. “Purple Threeawn. ”

     

    Literature Cited:
    - Nuttall, Thomas, 1834b.
    Full Size ImageColl. No. 1140, Aristida purpurea
    Full Size ImageColl. No. 1140, Aristida purpurea var. longiseta  

    First collected and described by Nuttall (1834) in his report in his near-fatal trip to Arkansa Territory.

    Original Text
    1. Aristida stricta. 2. A. oligantha. 3. A. dichotoma. 4. A. pallens
    5. A. * purpurea. Panicula erectiuscula gracili ; cal. Valvulis remotis aristulatis apice bifidis ; aristis capillaribus longissimis ; foliis brevibus scabris. — Hab. On the grassy plains of Red river, in arid situations. Flowering in May. — Obs. Perennial ' leaves narrow, short and scabrous ; ligula pilose ; culm about one foot high ; panicle many flowered, a little spreading, branches capillary ; flowers commonly in pairs (after the manner of the genus), bluish purple ; one valve of the calyx nearly double the length of the other, both bifid at the summit and shortly awned, the longer valve exceeding the corolla ; awns equal, capillary, nearly three times the length of the corolla and scabrous ; corolla minutely stipitate.

     

    Other articles:
    • Golden Checklist Flora:  Arrhenatherum elatius;  

    Arrhenatherum elatius (L.) P. Beauv. ex J. Presl & C. Presl. “Tall Oatgrass”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 1, p. 79) ...

    Original Text Translation and Comments
    2. AVENA calycibus bifloris, flofculo hermaphrodito mutico, mafculo aristato. Fl. fuec. 98.

    Avena calycibus bifloris, panicula laxe fpicata: pedunculis prioribus fafciculatis. Guett. ftamp. 1. p. 182.

    Gramen avenaceum, panicula acerofa, femine papofo. Dill. app. 48.

    Gramen avenaceum elatius, juba longa fplendente. Raj. meth. 179. angl. 3. p. 406. Morif. hift. 3. p. 214. f. 8. t. 7. f. 37. Scheuch. gram. 239. Hall. helv 225.

    β. Avena panicula nutante, calycibus bifloris: altero flofculo ariftato. Roy. lugdb. 66.

    Gramen nudofum, avenacea panicula, radice tuberibus praedita. Bauh. pin. 2. prodr. 3. theatr. 18. Scheuch. Gram. 237.

    Habitat in Europae maritimis & apricis.

    elatior. Dwelleth in the European maritime and sunny.

    Literature Cited:
    - Presl, Jan Svatopluk, anf K. B. Presl, 1819.  

    J. Presl & C. Presl (1819, p. 17) published A. elatius adding our grass to Beauvois' (1812) Arrhenatherum.

    Original Text Translation and Comments
    113. A. elatius B., O. wywysseny; flosc. superiore submutico, inferiore arista geniculata, culmi geniculis glabris, radice nodosa. H. G. 2. 49. Prata, pascua. ♃. 6, 3.

    B. = Palisot de Beauvois. Essai d'une nouvella Agrostographie. Our current citation is “P.Beauv. ex J.Presl & C.Presl” meaning the grass was described by Beauvois in Presl & Presl (1819).

    H.G. = Host. Gramina austriaca. Numerus prior tomum alter tabulam indigitat.

       

    Bouteloua Grama Grass

    Genus name honors two Spanish brothers Claudius (d. 1842) and Exteban Boutelou (d. 1813) professors of botany and agriculture respectively.

    Literature Cited:
    - Gould, Frank W., 1979.

    Other articles:
    • Glossary:  Kranz Syndrome;  

    Abstract. Bouteloua was established in 1805 by Mariano Lagasca. The type species is B. curtipendula (Michaux) Torrey, originally named B. racemosa by Lagasca. In the present treatment, 39 species are recognized, 29 of these restricted to North America and Central America, 2 species are endemic to the Antilles, 2 species occur in the Antilles, as well as at other locations, 5 species are distributed in both North and South America, and 1 species, B. megapotamica, is represented only in southern South America. Bouteloua is a characteristic member of the tribe Chlorideae of the subfamily Eragrostoideae (Chloridoideae). The species all are C4 plants with typical Kranz leaf anatomy and starch storage features. Chromosome numbers have been reported for 29 species with most species being diploid (2n = 20) or tetraploid (2n = 40). Aneuploid records or series of counts have been reported for 9 species.

    All species of Bouteloua are characterized by features of the Kranz Syndrome. They are C4 in their photosynthesis and have a characteristic chloridoid leaf anatomy. Starch storage is in specialized plastids of the leaf sheath bundles, and the arrangement of cells in the leaf blade is typically Kranz. The 39 recognized species are all variously adapted to shortgrass prairies, desert grasslands, and xeric sites along desert shrub areas, and sandy shores.

    The genus Bouteloua was described in 1805 by Mariano Lagasca. Lagasca proposed five species, B. racemosa, B. hirsuta, B. barbata, B. simplex, and B. prostrata, but did not designate a type species. Griffiths (1912), Hitchcock (1920), and Hitchcock et al. (1939) all have accepted the first species, B. racemosa, as the type. As plants of the type species had been named Chloris curtipendula by Michaux in 1803, the legitimate name for this taxon is Bouteloua curtipendula.

    Bouteloua was named in honor of two Spanish gardeners, the Boutelou brothers. Lagasca's original spelling of the genus name was Botelus. In a later publication (1816), he corrected this to Bouteloua.

    Two subgenera, Bouteloua and Chrondrosium (Desvaux) Gould. Weber & Wittmann (2012) retain Chrondrosium at the rank of genus, though spelling it Chrondrosum Desvaux, suggesting that we see Clayton, W. D., 1986, Genera Graminum: Grasses of the World. Kew Bull. Addit. Ser. XIII. 389 p. Peterson, et al. (2015) treat Chrondrosum as a section of Bouteloua containing B. gracilis and one other known in Colorado, B. simplex.

     

    Other articles:
    • Golden Checklist Flora:  Bouteloua curtipendula;  

    Bouteloua curtipendula (Michx.) Torr. in Marcy. “Side-Oats Grama”

     

    Literature Cited:
    - Michaux, Andre, 1803.  

    The basionym of B. curtipendula is Chloris curtipendula Michx.

    Original Text My Interpretation
    CHLORIS. Sw. CHLORIS. O. P. Swartz (1788) Nova Genera & Species Plantarum seu Prodromus
    Spicae unilateriflorae, spiculis subsessiliter biseriatis : quarum gluma communis 2-valvis, 2-6-flora : floribus dissimilibus ; valva altera ( saltem nonnullorum ) aristata : uno hermaphrodito , fertili ; caeteris inperfectis , masculis neutrisve ; ultimo pedicellato. Spikes unilateral flowered, darts subsessiliter biseriatis, of which the husk common 2-doors, 2-6-flowered, flowers are different; second valve (at least some) anstata one hermaphrodito, fertile; the rest of the imperfect, or the males neutrisve; last pedicellatae.
    curtipendula. C. racemo erecto longo ; e spicis plurimis , distiche alternis , e basí emittens rudimentum secundi floris inane , promisse aristatum ; intra quod bina alia rudimenta quasi in sola arista consistentia. curtipendula. Chloris, even on a long raceme; very many out of the ears of corn, distiche every other day, And I will send the initial stage of the second base of the flower out of the void, Retz promised; the rudiments of other things, and within it two of every sort as the only grain condition.
    Plantae cultae statura major; spicae 6-12-glumes. If cultivated plants are taller; 6-12-spike glumes.
    Hab. in aridis regionis Illinoensis ad Wabast et in rupibus ad prairie du rocher. ♃ Habitat. In arid regions of Illinois to Wabash River and watercourses of Prairie de Rocher (a town in southwest Illinois). Perennial.

    Prairie du Rocher is one of the oldest communities in the 21st century United States having been founded in 1722 by French colonists, mostly migrants from Canada. About four miles to the west, closer to the Mississippi River, is Fort de Chartres, site of a French military fortification and colonial headquarters established in 1720.

    Literature Cited:
    - Emory, William H., 1848.  

    Emory (1848) hinted as the existence of Bouteloua curtipendula as a name but this would be invalid because he really did not place B. racemosa in synonomy. I assume that Emory had heard from Torrey that B. curtipendula was the correct name but perhaps did not know that Torrey had not published the name.


    ____teloua racemosa, Lagasca. ? Culm erect, simple; spikes nu-
    ____s (20-40,) reflexed, 3-flowers; lower glume linear subulate;
    ____ one linear-lanceolate, scabrous, entire, nearly as long as the
    ____ts; lower palea of the perfect flower unequally tricuspidate,
    ____ent; abortive flower reduced to a slender awn which is nearly
    ____g as the perfect flower, furnished at the base with 2 short
    ____onspicuous bristles. Valley of the Gila, rare. This plant
    ____pretty well with Kunth's description of B. (Eutriana,) race-
    ____xcept in the pubescent lower palea, and the minute bristles
    ____base of the neiter flower. Whether it be the plant of La-
    ____r not is very difficult to determine from his brief character.
    It certainly is very different from B. racemosa of the United States,
    which has a large 3-awed neuter flower, and if distinct from La-
    gasca's, must receive another name. That of B. curtipendula
    would be appropriate.

    Literature Cited:
    - Marcy, Randolph B., 1854.  

    IPNI (2020) indicates that Bouteloua curtipendula was credit to Torrey in Marcy (1854), whereas Marcy seems to credit Torrey in Emory's report. IPNI also suggest that new combination was invalid, “Torrey (in Emory, Notes milit. Reconn. 154. 1848) mentioned this comb. Nov. (invalid)”

    Bouteloua racemosa, Lag. Var. Cienc. (1805) p. 141 ; Torr. In Emory's Rep., p. 154 ; not of Torr. Fl. N. York. Dinebra curtipendula, DC.? Kunth, Syn. Pl. Eq. 1, p. 281 ; excl. syn. Michx. Eutriana curtipendula, Trin. Fund. P. 161 (in part); Kunth, Enum. 1, p. 280, and Suppl. P. 233 ; excl. syn. Michx. And Willd. Main Fork of Red River; July. The detailed description of this species by Kunth, l. c., (drawn from a Mexican specimen collected by Humboldt) shows that the Chloris curtipendula of Michaux (Bouteloua curtipendula, Torr.) is a distinct species, as indicated in Emory's report, l. c.

     

    Other articles:
    • Golden Checklist Flora:  Bouteloua gracilis;  

    Bouteloua gracilis (Kunth) Lag. ex Griffiths. Blue Grama Grass.

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    110. ATHEROPOGON. Muhlenberg.
    ... [Description of A. apludoides, syn: Chloris curtipendula, Mich. = Bouteloua curtipendula (Michx.) Torrey – Ed.]
    2. A. * oligostachyum. Spikes 2 or 3, nearly terminal, many flowered; calix and corolla pilose; outer valve of the corolla distinctly 3-awned, the 2 lateral awns shorter, arising near the middle of the valve; neutral valve 3-awned.
    On the plains of the Missouri with the above. [Bouteloua curtipendula – Ed.] Common.
    Culm round, filiform, nearly naked, or with a single leaf, 8 to 12 inches high, smooth and erect. Leaves very short, smooth, and subulate, stipule and base of the spikes shortly bearded. Spikes 1, 2, or 3, about an inch long, usually curved backwards, unilateral, compressed, and pectinate, the second spikes bibracteate, rachis semiterete. Glumes in a double row, opposite; each 2-flowered; calix bluish-purple, exterior valve lanceolate, mucronate, with a single nerve; the nerve beset with a few scattered hairs arising from so many tubercles; inner valves shorter, very narrow. Corolla, outer valve lanceolate, carinate, 3-awned, pilose along the margins of the nerves, and at the base; inner valve smooth, shortly bi-cuspidate. Neutral flower 1-valved, obtuse, with 3 awns, and pubescent at the base.
    This species, though certainly a congener of the preceding, is considerably allied to Chloris, appearing to unite that genus and Sesleria, agreeing partly with the latter in the structure of the flowers, and with the former in its habitus. (A North American genus)

     

    Other articles:
    • Golden Checklist Flora:  Bromus briziformis;  

    Bromus briziformis Fisch. & C. A. Mey. “Rattlesnake Brome”

     

     

    Other articles:
    • Golden Checklist Flora:  Bromus lanatipes;  

    Bromus lanatipes (Shear) Rydb. “Wooly Brome”

     

     

    Other articles:
    • Golden Checklist Flora:  Bromus polyanthus;  

    Bromus polyanthus Scribn. ex Shear. “Great Basin Brome”

     

     

    Other articles:
    • Golden Checklist Flora:  Buchloe dactyloides;
    • Field Notes:  Coll. No. 1673, 16 Jun 2017;   Coll. No. 1674, 19 Jun 2017;  Coll. No. 1690, 29 Jun 2017;  

    Buchloë dactyloides (Nutt.) Engelm. Buffalo Grass.

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Glossary:  v. v.;  

    Buffalo Grass was first recognized in the field and published by Nuttall (1818). However, Nuttall only saw the staminate plants, and published the grass that way. He also was not sure of the genus, but settled on Sesleria as his best guess.

    95. SESLERIA. L. (Moor-grass.)
    Calix 2 to 5-flowered. Corolla 2-valved, valvea toothed at the point. Stigmata somewhat glandulous. — Flowers spiked, often purplish, base of the spike breacteate, or involucrate. Early flowering subalpine grasses, growing in calcareous mountains.
    Species. 1. S. Dactlyoides. Culm setaceous, leafy; leaves short, flat, subulate, and somewhat hairy; stipules bearded; spikes 2 or 3, few-flowered; flowers in 2 rows, disposed upon an unilateral rachis, calix mostly 2-flowered, and with the corolla acuminate and entire.
    Hab. On the open grassy plains of the Missouri; abundant. Flowers in May and Junes. v. v. Root after flowering resembling a bulb.
    Culm smooth and round, furnished with 2 or 3 leaves, about 4 or 5 inches high. Leaves flat, subulate, and somewhat hairy, 1 to 2 inhes in length, and about 2 lines wide; sheathes shorter than the internodes, very hairy around the stipules. Spikes 2 or 3, somewhat ovalm subtended by a single leaf, with which they are at first sheathed; rachis compressed, margined,spikelets 6 to 8, by pairs, inclined to one side. Calix 2-valved, 2 or 3-flowered, vales very unequal, each with a single nerve and carinate, the larger oblong-ovate, mucronulate. Outer valve of the corolla oblong-lanceolate, entire, 3-nerved, smooth, and menbranaceous, longer than the calix; inner 2-nerved, nearly the length of the outer. Anthers linear, entire, fulvous, exserted. Styles filiform, pubescept.
    This species appears on the one hand, alloed to Atheropogon, and on the other to Dactylis. Though rather a Sesleria than any other genus, it recedes from it in having the valves of the corolla entire at the apex, and thus it approaches Dactylis at least, the D. glomerata.
    With the exception of the present species, the genus Sesleria is confined to the alpine regions of Northern Europe.

    Literature Cited:
    - Rafinesque, C. S., 1819.

    Other articles:
    • Glossary:  nomen nudum;  

    Rafinesque (1819), recognizing that the grass must be distinct from the Old World Sesleria, published a genus name Bulbilis for Nuttall's grass.


    18. Sesleria dactyloides must form a peculiar genus by Mr. N's. own account, it may be called Bulbilis.

    The question will become whether this is a validly published name, or whether it is a nomen nudum.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1838-1843.  

    I don't see any grasses in Torrey & Gray's (1838-1843) Flora of North America.

    Literature Cited:
    - Marcy, Randolph B., 1854.  

    Sesleria dactyloides, Nutt. Gen, 1, p. 65; Kunth, Enum, 1, p. 323; Torr, in Emory's report, p. 323, t. 10. Upper tributaries of the Red River; July. This is the well known Buffalo-grass of the western prairies. It is remarkable that neither the grain nor the fertile flowers of this grass are known.

    Literature Cited:
    - Engelmann, George, M.D., 1859.  

    Original Text Comments and Interpretation
    TWO NEW DIOECIOUS GRASSES OF THE UNITED STATES.  
    By George Engelmann, M.D.  
    The grasses, though usually hermaphrodite, show a tendency to a separation of the sexes, and polygamous flowers are not rare among them. About 25 to 28 genera, one-twelfth of the whole number known, comprising only 75 to 80 species, about one seventy-fifth of all species, * are described as having monoecious and mostly heteromorphous flowers.  
    Only two genera of dioecious grasses are known to the books; of these, Spinifex, Lin., with 6 species from the East Indies and Australia, bearing on some plants staminate and on others complete flowers, is only incompletely dioecious; the other genus is Gynerium, H. B. K., five South American species. Some other dioecious species of genera, generally hermaphrodite, are noticed; such as Calamagrostis dioica, Lour., and Guadua dioca, Steud.  
    The unisexual grasses mostly belong to Oryzeae, Phalarideae, Paniceae, and Rottboellieae; none have been known among the tribes of Stipeae, Agrostideae, Chlorideae, Avenaceae, Festiceae, and Hordeeae.  
    They were unknown in the northern temperate zone, with the exception of Zizania and Tripsacum of North America and the cultivated Zea, all with heteromorphous staminate and pistillate flowers on the same plant. The dioecious grasses of our Flora are both species of Brizopyrum;† Eragrostis reptans is also frequently or mostly dioecious, and other species of this genus seem to be imperfectly so.  
    In the following pages, two new dioecious North American grasses are described, both types of new and very distinct genera, and both, it is believed, belonging to Chloridae.  

     
    * In the latest work on Grasses, Steudel's Glumaceae, published in 1855, about 6,000 species of Grasses are described, very unequally distributed in about 300 genera, many general containing only a single species, while Panicum alone comprises 864, Andropogon 461, Eragrostis 247, and Festuca 239 numbers.  
    Brizopyrum spicatum, Hook. Is from the eastern seacoast, and B. strictum from the saline soils of the Missouri region and of Utah. The flowers of both sexes are conform, but the staminate plants are readily distinguished from the pistillate ones by their more slender growth, the spikes overtopping the leaves; while in the pistillate plants the latter are longer than the spikes. = Distichlis spicata (L.) Greene
     

    Engelmann's description of the new genus:

    Original Text Comments and Interpretation
    BUCHLOË, Nov. Gen. BUCHLOË, New Genus
    Gramen plantitierum Americae Septentrionalis aridarum Missouriensium, Texensium, Mexicanarumque gregarium, perenne, stoloniferum, humile, sparse pilosum vel glabriusculum; ligulis barbatis. — Buchloe pro nimis longo Bubalochloe nomen vernaculum “Buffalograss,” graece reddit. Grass of arid North America, Missouri, Texas, Mexico, perennial stoloniferous, small, sparsely hairy or glabriusculus; ligules bearded. — Buchloe from a long form, Bubalochloe (Χορτ ο βου β αλ ου) The name of the breed “ Buffalograss” in Greek.

    Literature Cited:
    - Steudel, Ernst Gottlieb, 1855.  

    Description of the new species:

    Buchloe dactyloides.
    Syn. Plantae masculae: Sesleria dactyloides, Nuttall, Gen. I. p. 64. Sesleria (?) dactyloides, Torrey, in Emory's Rep. 1848, p. 153, Pl. X; id. In Whipple's Rep. Pacif, R.R. Expl., IV., p. 157. Calanthera dactyloides, Kenth (?) in Hooker's account of Geyer's Rocky Mountain plants, in Kew Journ. Bot., VIII., p. 18. Triodiae spec., Bentham, in Pl. Hartweg, nro. 250, p. 28. Lasiostega humilis, Rupprecht (ined) in Benth, Pl. Hartw. Corrig., P. 347. — Drummond Tex., Ill., nro. 378. Lindheimer, Pl. Tex. Exsicc. 569. Fendler N. Mex., 940. Berlandier, nro. 1612 and 1614. Hartw, 250 (fide Gray).
    Syn. Plantae faemineae: Antephora axilliflora, Steudel, Glum. I. . 111 — Drummond Tex., II., 359. Wright, 1849, 785; 1851-1852, 2079 (fide Torrey).
    This remarkable plant is found in our western pariries from the British possessions throughout the Mossouri Territory, Nebraska, Kansas, and New Mexico, down to Texas and Northern Mexico, and is, under the name of “Buffalo-grass,” well known to hunters and trappers as one of the most nutricious grasses, on which, for a part of the year, subsist and fatten immense herds of buffalo and the cattle of the hunter and emigrant. Since the time of Nuttall, who published an account of it, in his “Genera,” as early as 1818, the male plant has been collected by almost every botanist traversing those regions. The female plant had escaped the observers until it was described by Steudel, in the year 1855, from Drummond's Texan specimens, as a totally different plant and belonging even to a different tribe. Though Prof. Torrey had already, in Emory's Report, 1848, suggested the probability of the Buffalo-grass being a dioecious plant, the possibility that Nuttall's Sesleria dactyloides and Steudel's Antephora axilliflora could be the male and female of the same species was not even suspected, till finding both together in a collection sent by my brother, Henry Engelmann, who, as a geologist, accompanied the topographical corps attached to the army of Utah, I was struck with their similarity. My surmise, much doubted at first, became a certainty, when I discovered among some male plants, collected by A. Fendler, about Fort Kearny on the Platte River, a monoecious specimen, showing both male and female flowers on different stalks from the same rhizome. A figure of this important specimen is given on Pl. XII., fig. 3.
    That our plant is distinct from Sesleria has already been stated by Torrey (l. c. p. 154), and indeed by Nuttall himself (l. c. p. 65), and both have pointed to its affinity to Atheropogon or Chondrosium. The description now given fully confirms both positions. It also leaves no doubt that is is not an Antephora, nor at all paniceous. A new generic name, therefore, had to be given, and I have preferred to propose an abbreviated translation of the popular and widely known name of “Buffalo-grass,” retaining of course Nuttall's original specific appellation. The synonyms of the male plant, supplied through the kindness of Prof. Gray, are uncertain, Kunth never having published such a name as Calanthera, which, moreover, is quite unmeaning; nor can I learn that a genus Lasiostega has ever been described.
    The Buffalo-grass grows in dense tufts, sending out stolons. These, in most herbarium specimens, are only a few inches long, with internodes of 1/2-2 incles in length; Lindheimer, however, sends specimens from New Braunfels, Texas, with stolons 1-2 feet long, the internodes often measuring over 3 and even as much as 5 inches. The male plant seems to throw out more numerous runners than the female, and may often overspread and kill it out. Which would account for the much greater scarcity of the latter.
    Leaves 2-4 inches long, 1/2-1 1/2 lines wide, sparsely hairy or ciliate or glabrous; sheaths striate, glabrous, strongly bearded at the throat.
     

    Engelmann (1859) wrote a very long and detailed description of the staminate plant.

     

    Engelmann (1859) then wrote a very long and detailed description of the pistillate plant. The Engelmann article then continues with a description of Monanthochloë, and M. littoralis Engelm. This is also known as shore-grass and not known from Colorado, so the description is not included here.

    Literature Cited:
    - Plank, E. N., 1892.

    Other articles:
    • Golden Checklist Flora:  Buchloë dactyloides, Hitchcock, 1895;  

    E. N. Plank (1892) argued that the plant was actually monoecious. He based this on a single observation,

    During one of my botanical rambles in Kansas, while walking over soil newly moved by a freshet, I noticed the peculiar appearance of the individual plants of buffalo grass growing upon it. There were scores of them, if not hundreds. All of them appeared to be seedlings, having not yet sent out stolons. All of these plants were monoecious. That seems to be sexually the original character of the species. The fact of the unisexual flowering stems, proceeding from different parts of the plants, with its stoloniferous character generally increasing and spreading in that way, will fully account for its dioecious habit.

    Literature Cited:
    - Hitchcock, A. S., 1895.

    Other articles:
    • Golden Checklist Flora:  Buchloë dactyloides, Plank, 1892;  

    Hitchcock (1895) refuted Plank's (1892) observation that Buffalo Grass was monoecious.

    Note on buffalo grass. — I read with interest an article by Mr. Plank on “Buchloe dactyloides Englm., not a dioecious grass.” He asserts that the grass in question is not dioecious, as usually described, but monoecious, and in support records observations made in Kansas.
    Wishing to satisfy myself experimentally as to the correctness of this assertion, a few seeds were germinated in the greenhouse in the spring of 1893. A single seedling was transferred to an outdoor plat. This grew vigorously through the season, sending out stolons and forming a compact mat. During 1894 the mat became larger and denser, but no flowers appeared.
    However, the plant flowered this season (1895). Both staminate and pistillate flowers were present, the former preponderating. The flowers arose mostly from nodes that had taken root and thus become essentially independent plants. In no case did I find the two kinds of flowers from the same node, but from the interwoven state of the stolons I was unable to determine whether the two kinds of flowers were borne upon independent stolons.
    The plant was first described by Nuttall (Gen, 1: 65. 1818) from a staminate specimen, and named Sesleria dactyloides. He is evidently doubtful about the plant belonging to the genus Sesleria.
    Rafinesque having occasion to review Nuttall's Genera (Am. Monthly Mag. 2: 190. 1819) makes a note regarding this plant: “18. Sesleria dactyloides must form a peculiar genus by Mr. N.'s own account. It may be called Bulbilis.” It is upon this basis that Dr. Otto Kuntze establishes Bulbilis dactyloides (Nutt.) Raf. (Rev. Gen. Pl. 763).
    Nuttall remarks in his description: “Root, after flowering, resembling a bulb,” from which, doubtless, Rafinesque derives Bulbilis. Upon the margin of the copy of the American Monthly Magazine above quoted (in the library of the Missouri Botanical Garden), someone has suggested another derivation, “bull's bile!” — A. S. Hitchcock, Kansas Agricultural College, Manhattan.

    Literature Cited:
    - Lamson-Scribner, F., 1900.  

    Uses Rafinesque's name of Bulbilis dactyloides.

    Literature Cited:
    - Schaffner, John H., 1920.  

    Schaffner (1920) published an article on the dioecious nature of Buffalo Grass. He combined field observations and greenhouse experiments to support his contention that Buffalo Grass is strictly dioecious.

    Literature Cited:
    - Hitchcock, A. S., 1927.  

    Hitchcock (1927) proposed conservation of Buchloë against Bulbilis and others because Engelmann's Buchloë was the first name under which both staminate and pistillate plants were described.

    Buchloe Engelm. (1859) is conserved against Bulbilis Raf. (1819), Calanthera “Nutt.&rdquo: (1856), and Casiostega Rupr. (1857). Bulbilis was proposed by Rafinesque in a review of Nuttall's Genera as follows: “Sesleria dactyloides must for a peculiar genus by Mr. N's own account, it may be called Bulbilis.” Calanthera was mentioned by Hooker in a list of Geyer's plants from the Upper Missouri. “Calanthera dactyloides Kth.–Nutt. Sesleria.” Casiostega (of the Nomina Conservanda, a lisprint for Lasiostega Rpur.; Benth, Pl. Hartw. 347. 1857) is a nomen nudum. Nuttall's description of Sesleria dactyloides was based on the staminate plant only and until Engelmann gave his full description of both sexes of the buffalo grass there had been no mention of pistillate spikelets. Though in Bulbilis and Calanthera the type species is indicated and a previously published description is referred to, that description is so inadequate, compared to the detailed study published by Engelmann, that they may well be rejected in favor of Buchloe.

    Other articles:
    • Golden Checklist Flora:  Buchloë dactyloides, Hitchcock, 1971, 2nd ed.;  

    See Hitchcock, 1971, below.

    Literature Cited:
    - Anderson, Kling, and A. E. Aldous, 1937.  

    An article on monoecious Buffalo Grass by Anderson & Aldous (1937) is behind a paywall I have been unable to penetrate.

    Literature Cited:
    - Gernert, W. B., 1937.  

    An article by Gernert (1937) appears to address the height of pistillate spikes relative to harvesting them, but is behind a paywall I have not penetrated.

    Literature Cited:
    - Hensel, R. L., 1938.  

    Hensel (1938) wrote about perfect-flowered buffalo grass which might be interesting to read were it not, alas, behind a paywall.

    Literature Cited:
    - Spurlock, Clay, 1940.  

    Spurlock (1940) surveys monoecious and dioecious grasses in America, finding 17 genera and 54 species of monoecious, and 13 genera and 23 species of dioecious grasses of the Western Hemisphere, of which Buffalo Grass is one of the latter. Note is made of the controversy about the degree of dioecism seen in Buffalo Grass.

    Literature Cited:
    - Burr, Richard D., 1951.  

    Burr (1951) wrote about his observations in variations of sex along stolons of

    Literature Cited:
    - Rickett, H. W., and F. A. Stafleu, 1959.  

    In a review of conserved names, Rickett and Stafleu (1959) note that conservation of the name “Buchloë” was superfluous.

    † 308. Buchloë Engelmann, Trans. Acad. Sci. St. Louis 1 432. 1859 quadrim. 1. T.: B. dactyloides (Nuttall) Engelmann (Sesleria dactyloides) Nuttall).  
    Note: Conservation superfluous: Calanthera Nuttall ex W. J. Hooker, Journ, Bot. Kew Misc. 8: 18 (1856), Bulbilis Rafinesque, Am. Mon. Mag. Crit. Rev. 4: 190. (1819), and Lasiostegia Ruprecht ex Betham, Pl. Hartw. 347 (1857), are all nomina nuda.

    The meaning of the dagger (†) is unclear. All names with daggers were subjects of a superfluous conservation. However, not all with superfluous conservation were marked with a dagger.

    Literature Cited:
    - Hitchcock, A. S., 1971.

    Other articles:
    • Golden Checklist Flora:  Buchloë dactyloides, Hitchcock, 1935;  

    115. BÚCHLOË Engelm. (Bulbilis Raf.)
    Plants dioecious or monecious. … Type species, Buchloë dactyloides. Name contracted from Greek boubalos, buffalo, and chloë, grass, a Greek rendering of the common name, “buffalo grass.”
    1. Buchloë dactyloides (Nutt.) Engelm. Buffalo Grass. … The sod houses of the early settlers were made mostly from the sod of this grass. In 1941 it was planted at Boyce Thompson Institute, Yonkers, N. Y., and is proving to be an excellent cover for exposed dry banks.

    Literature Cited:
    - Bai, T. J., 1990.  

    Abstract : Seventy-five stands with different compositions of blue grama (B. gracilis) and buffalograss (B. dactyloides) were sampled in 1987. Multiple regression and principal component analysis techniques were used to investigate the relationship between blue grama and buffalograss composition and soil characteristics. Blue grama was often more abundant on sandy soil, while buffalograss was found on clay soil. Blue grama was also found on clay soils when lime content was high. The essential factor underlying sand, clay and lime content of soil was interpreted as water stress. Relative crowding coeff. calculated for blue grama and buffalograss showing that both species had greater height and produced more vegetation when found intermingled than when growing in monoculture.

    Literature Cited:
    - Quinn, James A., 1991.  

    Quinn (1991) examined multiple hypotheses for dioecy in Buffalo Grass and found support only for “out-crossing” as an advantage afforded by dioecy.

    Abstract. Buchloe dactyloides is a perennial dioecious grass in which male and female inflorescences are so strikingly dimorphic that they were originally assigned to different genera. The objective of this paper is to present the results of tests for sex-specific vegetative characters, ecological differences, and sexual niche-partitioning, combining them with prior information on the reproductive biology of Buchloe for an evaluation of the key factors leading to the evolution of dioecy and sexual dimorphism. Field and greenhouse data were collected from Oklahoma and Kansas populations on vegetative characters, allocation to reproduction, and relative growth and competitive success along resource gradients. Except for greater susceptibility to leaf rust by males, there were no significant differences between males and females in vegetative characters, total biomass, or reproductive effort. Field studies of spatial distributions of males and females failed to show any relation to soil, topography, or soil moisture. In a 45-month greenhouse experiment starting at the seedling stage, the relative growth and competitive success of randomly paired individuals showed no evidence for differential competitive success or for niche-partitioning of males and females. The "outcrossing advantage" and subsequent sexual specialization of the female inflorescence appear to be the major factors underlying this dimorphic system.

    Literature Cited:
    - Huff, David R., and Lin Wu, 1992.  

    Abstract. Variations of sex inconstancy were examined for vegetative and seed samples from eight natural populations of buffalograss located along two east-west transects crossing the shortgrass prairies of Oklahoma, New Mexico, and Texas. Each of the eight populations was found to contain inconstant (monoecious) sex forms. Sex form distributions ranged from the Guymon vegetative sample, having no inconstant sex forms, to the Chillicothe seed sample in which the frequency of inconstant sex forms was nearly 70%. Frequencies of inconstant sex forms were generally higher for seed samples than for vegetative samples. Male to female sex ratio of constant (dioecious) sex forms generally did not differ from 1:1 expectations. Inconstant sex forms were more common among peripheral populations where buffalograss vegetation coverage was sparse than for more central populations having a higher concentration of buffalograss vegetation. Quantitative measures of sex inconstancy from artificial crosses were significantly (P < 0.001) correlated with the additive linear model of general combining ability, suggesting that sex determination in buffalograss has high heritability. The possible selection forces affecting the frequency of monoecious sex forms among natural populations are discussed.

    Literature Cited:
    - Columbus, J. Travis, 1999.  

    An expanded circumscription of Bouteloua (Graminae: Chloridoideae): New Combinations and Names

    Literature Cited:
    - Columbus, J. Travis, Michael S. Kinney, Maria Elena Siqueiros Delgado, and J. Mark Porter, 2000.  

    Phylogenetics of Bouteloua and Relatives (Granineae: Chloridoideae): Cladistic Parsimony Analysis of Internal Transcribed Spacer (nrDNA) and trnL-F (cpDNA) Sequences.

    Literature Cited:
    - Peterson, Paul M., and Konstatin Romaschenko, 2015.  

    Phylogeny and subgeneric classification of Bouteloua with a new species, B. herrera-arrietae (Poaceae: Chloridoideae: Cynodonteae: Boutelouinae) … Buchloe is reduced to a section of Bouteloua.

     

    Other articles:
    • Golden Checklist Flora:  Calamovilfa longifolia;  

    Calamovilfa longifolia (Hook.) Scribn. “Prairie Sandreed”

     

    Literature Cited:
    - Hooker, Sir William Jackson, 1829-1840.  

    Hooker,____, v. 2, p. 241

    VI. Arundinaceae. Kth.
    19. CALAMAGROSTIS. Adans.
    5. C. longifolia ; laevissima,foliis anguste linearibus longissime acuminatissimis, panicula erecta stricta subspicata, glumae valvis inaequalibus perianthium acutum muticum superantibus, villis perianthio brevioribus.
    Hab. Saskatchawan. Drummond. — A species remarkable for the great length of its leaves, exceeding the culm, tapering into a long, slender apex, and for the pale, very smooth, glossy flowers. The base of the culm send out creeping shoots.

    Literature Cited:
    - Hackel, Eduard, 1890.  

    Hackel (1890) ... translated by F. Lamson-Scribner and Effie A. Southworth

    138. (141) Ammophila Host. (Psamma Beauv.) Panicles usually narrow and spike-like. Flowering glume and palea chartaceous, somewhat indurated, awnless; spikelets comparatively large. ...
    Obs. — Very nearly related to Ammophila are two N. American species, Calamagrostis brevipilis Gray and C. longifolia Hook., with the loose panicle of Calamagrostis, but the chartaceous flowering glumes of Ammophila, distinct from both genera by the one-nerved flowering glumes ; they may best be considered a separate genus, Calamovilfa (Gray as a sect. of Calamagrostis). (Hackel in MS.)
    [138a. Calamovilfa. Empty glumes unequal ; flowering glumes one-nerved ; rachilla not prolonged.

    Species two, in N. America (C. brevipilis of the Atlantic coast, and C. longifolia of the western interior). These species are referred to Ammophila in B. & H. Gen Pl., vol. III, p. 1153.]

     

    Other articles:
    • Golden Checklist Flora:  Cenchrus longispinus;  

    Cenchrus longispinus (Hack.) Fern. “Longspine Sandbur”

     

    Literature Cited:
    - Fernald, M. L., 1943.  

    The common northern Burgrass. — The common Burgrass of open sands, from New Hampshire to Oregon, south to North Carolina, Kentucky, Missouri, Kansas and New Mexico, is indigenous or spread from indigenous colonies through much of this range but, singularly enough, it has always passed under specific names which technically belong to other species. Long identified as Cenchrus echinatus L., it was forced to resign that name to the quite different tropical species. Similarly the name C. tribuloides L. soon proved to belong to the very coarse tropical and southern coastal species. Then C. carolinianus Walt., was despairingly grasped, but Walter's type is unknown, our plant is doubtfully in his territory, and Mrs. Chase, Contrib. U. S. Nat. Herb. xxii. 76 (1920), believes that C. carolinianus might have been C. incertus M. A. Curtis, which does grow in Walter's country. In her monograph of the genus she merges our plant with the Mexican C. pauciflorus Benth. and, until Dr. I. M. Johnston, working upon his Mexican and southern Texan material, segregated it off from the great bulk of specimens from the northern and Rocky Mountain region of the United States, it so rested. Dr. Johnston, not wishing to get involved with the more northern and northeastern plant, called the matter to my attention. There is no doubt that C. pauciflorus is the Mexican (and Texan) species, Bentham having described it from Lower California: "culmis suberectis, . . . Folia plerumque angustiora [quam in C. echinata], spinis dorsalibus marginalibusque validis basi dilatatis".
    Cenchrus pauciflorus, then, is the Mexican and Texan species with culms usually in erect or ascending tufts, the leaves 1.5-4 mm. broad, the summit of the sheath spreading as a chartaceous flange; spikes 1-5 cm. long, 1-1.5 cm. thick; the stramineous involucres 8-12 mm. broad (from tip to tip of mature spines), the dorsal and lateral spines compressed and broad-based. The characteristic fruit, from the type of C. pauciflorus, is shown by Mrs. Chase, 1. c. fig. 17, p. 68.
    Our plant, on the other hand, has more generally decumbent or rooting culms, up to 8 dm. long and strongly geniculate; leaves 3-8 mm. broad, the enlarged sheaths constricted at summit; spike 1.5-8 cm. long and 1.3-2 cm. thick, the mature stramineous to bronze or purplish burs 1-1.5 cm. in diameter, with the coarse spines subulate-subterete. The bur is beautifully shown, as that of C. pauciflorus, in Chase, 1. c. 69, fig. 18, and this figure, instead of that of true C. pauciflorus, was copied in Hitchcock, Man. fig. 1594. As compared with that of real C. pauciflorus it is too large, with more numerous spines, the stronger ones more slender and without deltoid and flattened bases.
    So far as I can find the only available name for the northern and transcontinental species is C. echinatus, forma longispinus Hackel, based upon Connecticut material collected by Harger and distributed by Kneucker (no. 426). As a diagnosis of a form of C. echinatus, in the loose sense, Hackel's brief description was sufficient. Treated as a species the plant demands a fuller account. It is fortunate that Hackel has supplied the basonym and type. Even so, I take no comfort in having even my name intimately associated with a Cenchrus and the keen taxonomist who called it to my attention would not care to have the species named for him!
    Cenchrus longispinus (Hackel), stat. nov. C. echinatus, forma longispinus Hackel in Kneucker, Allg. Bot. Zeitschr. ix. 169 (1903). Planta annua ; culmis decumbentibus vel adscendentibus ad 8 dm. longis geniculatis basi ramosis; foliis 3-8 mm. latis, vaginis distentis apice constrictis; spicis maturis 1.5-8 cm. longis 1.3-2 cm. crassis; involucris maturis stramineis vel aeneis vel purpurascentibus hirsutis 1-1.5 cm. diametro, spinulis majoribus tereti-subulatis. Type of C. echinatus, forma longispinus: Oxford, Connecticut, E. B. Harger in Kneucker, Gram. Exsicc. Lief. XV. no. 426.
    In southeastern Virginia, fortunately, Cenchrus longispinus is rare. Our only collections are the following. James City County: sandy field about 5 miles west of Toano, R. W. Menzel, no. 187. Southampton County: dry white sand of clearing in oak and pine woods bordering Assamoosick Swamp, south of Sebrell, no. 10,941; grassy roadside about 7 miles south of Franklin, no. 7297 (misidentified as C. incertus).

     

    Other articles:
    • Golden Checklist Flora:  Danthonia spicata;  

    Danthonia spicata (L.) P. Beauv. ex Roem. & Schult. “Poverty Oatgrass”

     

    Literature Cited:
    - Roemer, Johann Jacob, and Josef August Schultes, 1817-1830.  

    Roemer & Schultes (1817, v. 2, p. 690) — essentially the 9th edition of Linnaeus Species Plantarum — placed Avena spicata L. in Danthonia DC.

    Thanks to P. de Beavois on p. 18 of the Preface to volume 1:

    Original Text
    Illm. Palisot de Beauvois, experientissimum Botanicum, qui nec torridissimam Africam, nec Americam borealem ignotam sibi esse voluit.

    Publication of D. spicata from volume 2, page 690:

    Original Text
    2 * D. spicata P. de Beauv.; spicato paniculata, calyce flosculis sex longiore , corollae valvulâ exteriore apice aristatâ furcatâque. Avena spicata Sp. Pl. p. 119. Willd. Spec. I. p. 453. Avena glumosa, foliis subsetaceis collo vaginarum villoso ; paniculâ parvâ subspicatâ pauciflorâ ; calyce spiculam sexfloram superante, valvâ florum exteriore ex apice bicorni aristatâ. Mich. Fl. Bor. Amer. I. p. 72. Pers. Syn. I. p. 101. Pursh Fl. Amer. Sept. I. p. 86.
    „Gramen angustum; folia linearia; spica composita e spiculiis 3-4, remotis erectis, spiculae subpedunculatae, singulis calyx diphyllus sublatus aequalis ; flosculi sex sessiles erecti, laciniae glumae corollinae exterioris setaceae, arista articulata longotudine spiculae. Facies Festucae decumbentisLinn. Inflorescentiam re vera paniculam esse, nee spicam, aristamque basi spiralem, monuit b. Michaux. A nova Anglia ad Carolinam.
    Obs. An huc illa cum? allata Danth. spicaeformis, quae et Avena spicaeformis audit? sed Danthoniam glumosam sine dubio habet P. de Beauv.

     

    Other articles:
    • Golden Checklist Flora:  Dichanthelium oligosanthes ssp. scribnerianum;  

    Dichanthelium oligosanthes (J. A. Schultes) Gould ssp. scribnerianum (Nash) Freckmann & Lelong “Few-Flowered Panicgrass”

    Freckmann & Lelong (2002, v. 20, p. 170) ...

    Dichanthelium oligosanthes subsp. scribnerianum (Nash) Freckmann & Lelong, comb. & stat. nov. Basionym: Panicum scribnerianum Nash, Bull. Torrey Bot. Club 22:421. 1895. Panicum oligosanthes var. scribnerianum (Nash) Fernald, Rhodora 36:80. 1934. Dichanthelium oligosanthes var. scribnerinum (Nash)Gould, Brittonia 26:60. 1974. TYPE: U.S.A. Pennsylvania. Wysox, Jul 1836, J. Carey s.n. (Lectotype: GH [designated by Hitchc. & Chase, Contr. U.S. Natl. Herb. 15:283. 1910]).
    Panicum scoparium S. Watson in A. Gray, Man., ed. 6:632. 1890, nec Lam. (1798), nec Rudge (1805), nec Elliott. (1816).

     

    Other articles:
    • Golden Checklist Flora:  Digitaria sanguinalis;  

    Digitaria sanguinalis (L.) Scop. “Red-hair Crab-grass”

     

     

    Other articles:
    • Golden Checklist Flora:  Elymus albicans;  

    Elymus albicans (Scribn. & J.G.Sm.) Á.Löve. “Montana Wild Rye”

     

     

    Other articles:
    • Golden Checklist Flora:  Elymus canadensis;  

    Elymus canadensis L. “Canadian Wildrye”

     

     

    Other articles:
    • Golden Checklist Flora:  Elymus elymoides;  

    Elymus elymoides (Raf.) Swezey “Squirreltail”

     

    Literature Cited:
    - Graustein, Jeannette E., 1967.
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Thomas Nuttall collected a grass on arid plains of the Missouri that he described as Ægilops hystrix (Nuttall, 1818, v. 1, p. 85).

    Original Text Comments
    117. ÆGILOPS. L.  
    Calix, lateral, 2-valved, mostly 3-flowered, valves coriaceous, broad, with many awns; awns rigid and divergent. Corolla 2 valved, outer valve terminated by 2 or three awns. — Flowers spiked, intermediate masculine; lateral, hermaphrodite, sessile.  
    Small grasses, allied to Elymus, valves of the calix remarkably rigid and truncate, deeply divided into many flat and long scabrous awns; valves of the corolla also simularly divided and awned.  
    Species. 1. Æ. * Hystrix. Spike squarrose, with very long recurved and divergent awns: calix smooth, generally 4-parted to the base: segments mostly bifid, unequally 2-awned; spikelet about 4 flowered, the 2 masculine or neuter pedicellate, and intermediate; dorsal valve of the corolla terminated by about 2 or 3 unequal awns.  
    Considerably allied to Elymus. … [Long description omitted.]  
    On the arid plains of the Missouri. A search of SEINet for collections of Elymus elymoides in Missouri, Nebraska, South Dakota, and North Dakota, showed that there are almost no collections of the grass in proximity to the Missouri River. However, there are a few collections of the grass on or near the river from Fort Peck, Montana, and above, or Glendive, Montana, on the Yellowstone River, and above. Graustein (1967, p. 70) notes that Nuttall was very eager to reach the Rocky Mountains (the “Northern Andes” as Nuttall called them), and that a trail that struck almost directly west to the Yellowstone River from Fort Mandan doubtless enticed the naturalist. However, we do not really know how far above Fort Mandan that Nuttall ventured.
    Of this genus there are 2 species in the South of Europe, one of them also common to Barbary, and the other to Candia, there are likewise 2 other species peculiar to those places.  

    The next genus that Nuttall described was Elymus.

    Literature Cited:
    - Rafinesque, C. S., 1819.
    - Shaw, Robert B., 2008.
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Rafinesque's (1819) description of Sitanion elymoides Raf. is not available on Biodoversity Heritage Library, because Part 89 is missing. However, that part is available on the HathiTrust Digital Library.

    Original Text Translation
    PRODROME PRODROMO
    Des nouveaux Genres de Plantes observes in 1817 et 1818 dans l'interior des Etats-Unis d'Amerique; New Genres of Plants Observed in 1817 and 1818 in the Interior of the United States of America
    Par C. S. RAFINESQUE, By C. S. RAFINESQUE
    Professeur de Botanique et d'Histoire naturelle dans l' Universite de Lexington. Professor of Botany and Natural History at Lexington University.
     
    II. PARTIE. MONOCOTYLEES. PART II. MONOCOTYLEDONS.
     
    32. SITANION. (Graminee.) Fleurs polygames males en epi. Involucre lateral pentaphylle, multiflore. Glume univalve, convolutee, inegalement bifide et biaristee, contenant 4-5 fleurs, divisees en 2 spicules geminees. Glumelle bivalve, valves inegales, l'exterieure tres-grande, convexe, trifide, 3-aristee; ariste mediane tres-longue; valve interieure concavem bifide, mutique. 3 etamines. 2 styles. Fleur terminale communement male a 2 etamines et 2 setules ecailleuses. — Ce genre differe de l'Elymus par linvolucre 5-phylle, glume, glumelles, polygamie, etc. Une espece, S. elymoides. Chaume strie, scabre; feuilles scabres, glauques; epi droit, fleurs laches, involucres et aristes tres-longs, scabres, divariques; glumes lisses, dos uninerve, glumelles scabres sur les bords. — Missouri. 32. SITANION. (Graminee.) Polygamous flowers male in spike. Lateral involucre pentaphyll, multiflora. Glume univalve, convoluted, unevenly bifid and biaristous, containing 4-5 flowers, divided into 2 gemine spicules. Bivalve umbilicus, valves unequal, outer very large, convex, trifid, 3-aristate; very long median awn; inner valve concavem bifid, mutic. 3 stamens. 2 styles. Commonly male terminal flower with 2 stamens and 2 scaly setules. — This genus differs from Elymus by 5-phylinvolucre, glume, lemma, polygamy, etc. One species, S. elymoides . Thatch streak, scabrous; leaves scabrous, glaucous; spike straight, flowers loose, involucral and arist very long, scabrous, divaric; glumes smooth, back uninerve, scabrous glumes on margins. — Missouri.
     

    The next taxon Rafinesque described was Critesion Raf. a generic name still used as a segregate of Hordeum by Weber & Wittmann (2012) and Shaw (2008).

    Literature Cited:
    - Anonymous, n.d..
    - Swezey, Goodwin D., 1891.  

    Swezey (1891) described Elymus elymoides (Raf.) Swezey in his Nebraska Flowering Plants.

    Original Text
    ...
    I have endeavored in every case to give a plant the oldest available specific name it has borne, believing that this is the only rule which can lead ultimately to a settled nomenclature. In some cases this may lead to less euphonius combinations as in the case of Elymus elymoides; and here too these is more justification for the name E. Sitanion since this species constituted Rafinesque's entire genus Sitanion when it bore the name S. elymoides. Yet if it is an Elymus and it its original specific name was elymoides, then it should, as it seems to me, bear the name Elymus elymoides, (Raf.), euphony or no euphony. In this case and in that of Petalostemon compactus, (Sprengl.), I have not been able to learn who, if any one, has used this conbination of generic and specific names before and have accordingly left blank the name following the original author.
    Elymus elymoides, (Raf.) — — (E. Sitanion, Schultes.) Lewellen.

    So while Swezey did not know who first used Elymus elymoides, it turns out it was him!

    An anonymous review says the following about this publication.

    Original Text
    Nebraska Flowering Plants. Goodwin D. Swezey. (Doane College, Natural History Studies No. 1, Pamph. 8vo. pp. 16. Crete, 1891.
    This is a list of Nebraska localities for flowering plants in the herbarium of Doane College, based mainly on collections made by Prof. Swezey in a tour through the western part of the State, and by some of his students; 553 species and varieties are enumerated, of which 76 are here first definitely recorded as occurring within the area. The nomenclature is based on the stability of the oldest specific name, ‘believing that this is the only rule which can lead ultimately to a settled nomenclature.’ Two binomials are here first proposed: Petalostemon compactus (Spreng.) (P. macrostachyus, Torr.) and Elymus elymoides (Raf.), (E. Sitanion, Schultes). The original author is cited in parenthesis. The list is an important supplement to Mr. Webber's Flora.
    N. L. B.

    I suspect that “N. L. B.” is Nathaniel Lord Britton.

    Literature Cited:
    - Small, Jared G., 1899.  

    Sitanion hystrix was published by Jared G. Small in a series of Studies on American Grasses published by the U. S. Department of Agriculture. Small places all of Nuttall's Ægilops, excluding others of Linnaeus, in Sitanion, and adds quite a few specific names one might recognize.

    1. Sitanion jubatum J. G. Smith, sp. nov. (=Elymus multisetus (J.G.Sm.) Burtt Davy)

    2. Sitanion villosum J. G. Smith, sp. nov. (=Elymus multisetus (J.G.Sm.) Burtt Davy)

    3. Sitanion multisetum J. G. Smith, sp. nov. (=Elymus multisetus (J.G.Sm.) Burtt Davy)

    4. Sitanion polyantherix J. G. Smith, new name. Polyantherix hystrix nees, in Ann. Nat. Hist. 1: 284 (1838), not Ægilops hystrix Nutt. (=Elymus multisetus (J.G.Sm.) Burtt Davy)

    etc.

    Yikes!!! It's a mess. (Sitanion hystrix J.G.Sm., Bull. Div. Agrostol. U.S.D.A. 18: 15, pl. 2 (1899). https://www.biodiversitylibrary.org/page/35499501#page/366)

    Literature Cited:
    - Bentham, George, and John Dalton Hooker, 1873.
    - Hackel, Eduard, 1890.
    - Nuttall, Thomas, 1818.
    - Small, Jared G., 1899.  

    Original Text Comments
    INTRODUCTION The text of the Introduction was written by Lamson-Scribner.
    ...  
    Nuttall,1 who first described the species of this genus, referred it to the European Ægilops and named his plant Ægilops hystrix His description was carefully drawn up and his species can be readily recognized. A year later, Rafinesque2 published his genus Sitanion, based upon a single species, which he named Sitanion elymoides. It has been found impossible to determine with certainty which of the species enumerated in the present paper was the one named by Rafinesque; it certainly was not, however, the grass described by Nuttall.  
    Our leading authorities, Bentham and Hooker,3 Hackel,4 and Baillon,5 have all reduced Sitanion to a section of Elymus. The articulate rachis, readily breaking up at maturity, abd the usually bifid or many parted and awned empty glumes are well-defined characters, distinguishing the wpecies from Elymus, and justifying their separation as a distinct genus. To be sure there are species so closely connecting Elymus with Sitanion that it is difficult to determine to which genus they ought to be referred, but the same is true in the case of Elymus and Agropyron; there are intermediates which may with equal propriety be placed either in the one genus or the other.  


    1 Genera North American Plants, 1: 86. 1818
    2 Journ. Phys., 89: 103. 1819.
    3 Genera Plantarum 3: p. 1207.
    4 Die Natürlichen Pflanzenfamilien 2: part 2, p. 88.
    5 Histoire des Plantes, Monographie des Graminées, 258.
     
    ....  

    Literature Cited:
    - Nuttall, Thomas, 1818.
    - Small, Jared G., 1899.

    Other articles:
    • Interstate 80:   at Wamsutter;

    Locations: La Veta. Villa Grove. Wamsutter.  

    Original Text Comments
    12. SITANION HYSTRIX (Nutt.) J. G. Smith, new combination. (Ægilops hystrix Nutt. Gen. N. Am. Pl., 1: 86, 1818.) Pl. II.  
    Culms 1 to 3 dm. high, slender, erect or ascending, scabrous above, clothed at the base with papery leaf-sheaths. Innovations very leafy, one third to two-thirds the length of the culms. Sheaths striate, strigose-pubescent, open at the throat, closely envoloping the internodes. Ligule almost obsolete. Blades narrowly linear, flat ot at length involute, strigose-pubescent throughout, prominently 9-nerved, scabrous along the margins, erect or ascending; those of the innovations 7 to 12 cm. long, 1 to 2 mm. wide; culm leaves about as long, 2 to 4 mm. wide. Spike 5 to 7 cm. long, erect or subflexuous, exserted, or its basal portion inclosed in the uppermost leaf-sheath, closely flowered. Spikelets 3- to 4-flowered, complessed. Empty glumes bifid, from bear the base and unequally 2-awned; the strongly scabrous, glaucous, divergent awns, 3 to 4 cm. long. Flowering glume 7 to 8 mm. long, linear-lanceolate, minutely pubescent, 3-awned, the miffle awn rather slender, recurved, about 3 cm. long. Palea as long as or longer than the flowering glume, scabrous, tipped with two slender awns, 2 to 3 mm. long. Internodes of the rachis glaucous, linear, not at all dilated above, about 5 mm. long.  
    A common, worthless bunch grass on shale hills and among the sagebrush on the high plains from western Colorado to eastern Washington.  
    SPECIMENS EXAMINED: Wyoming: P. A. Rydberg, No. 2028, Wamsuter, July 24, 1895; C. L. Shear, No. 280½, Wamsutter, June 24, 1895; No. 283, Green River, June 25, 1895; Thomas A. Williams, No. 2437, dry rocky hillsides, Evanston, July 10, 1897; No. 2379, dry sagebrsh hills, Green River, July 9, 1897; Aven Nelson, No. 3058, Green River Hills, May 31, 1897; No. 3669, Wamsutter, July 10, 1897; No. 3784, North Vermilion Creek, July 20, 1897.  
    Washington: C. V. Piper, No. 2579, on sagebrish land, Ellensburg, July 9, 1897. A. B. Leckenby, Walla Walla, July 12, 1898.  
    Colorado: John Wolfe, No. 623, 1873; C. Thomas, 1869; and F. E. Clements, No. 60, Walsenburg, July 10, 1896.

    John Wolfe, No. 623, 1873. NY1655258, Lieut. G. M. Wheeler, Corps of Engineers, U. S. Army. Explorations and Surveys West of the 100th Meridian. Locality, Denver, Colorado. Gift from Wesleyan Univ. Deposited NY in 1981. See Brittonia 34(4). 1982. NY1673562, Lieut. G. M. Wheeler, Corps of Engineers, U. S. Army. Explorations and Surveys West of the 100th Meridian. Locality, Denver, Colorado. YU111158. Lieut. G. M. Wheeler, Corps of Engineers, U. S. Army. Explorations and Surveys West of the 100th Meridian. Locality, Denver, Colorado. Sheffield Scientific School. Daniel C. Eaton Collection.

    C. Thomas, 1869, collection not found online.

    F. E. Clements, No. 60, Walsenburg, July 10, 1896.This collection was not found online. However, two other collections of the taxon were found collected later in the same year, one at La Vela (sic), Huerfano County, and one at Villa Grove, Saguache County.

    There are in the herbarium of the Philadelphia Academy of Science two of Nuttall's specimens of Sitanion. One of these, labeled “Chretomeris trichoides, R. Mts. Platte,” is exactly identical with No. 3784, A. Nelson, and No. 283, C. L. Shear. both collected in the Red Desert of Wyoming. The other, labeled “Elymus difformis, R. Mts. Platte,” in nearly identical with No. 2028, Rydberg, from Wamsutter, Wyo. If these specimens are those from which Nuttall's description of Ægilops hystrix was drawn. and they agree better with his description than any specimen from the “arid plains of the Missouri” so far examined, then there was undoubtedly a mistake made in referrinf the habitat of this to that locality. I doubt that the two Nuttall specimens of Sitanion in the herbarium of the Philadelphia Academy of Science are the specimens from which Nuttall described Ægilops hystrix. It has to do with timing. The specimens location is “R. Mts. Platte” Yet when Nuttall (1818) published Æ. hystrix he had not yet been to the Rocky Mountains along the Platte River. The earliest specimens from that region would have been the Wyeth specimens from 1833, a date long after Nuttall published Æ. hystrix.
    I am assured by Dr. E. L. Greene that it is highly improbable that Rafinesque drew his description of S. elymoides from Nuttall's plant, and it is certain that Rafinesque's description (Journ. Phys. 89: 1819) differs in important particulars from that of Ægilops hystrix, Nuttall. I am, however, unable definitely to identify any Sitanion with which I am familiar as the true S. elymoides, Raf. The locality, “Missouri,” of 1819, was then applied to what now constitutes several large States in which a dozen of more separate species occur. Rafinesque apparently left no type, and the original description is too fragmentary to enable one to more than guess at the identity of the plant which he described.  

    Literature Cited:
    - Harrington, H. D., 1964, 2nd ed..  

    Harrington (1964, 2ed.) treated this grass as Sitanion hystrix (Nutt.) J. G. Smith

    Literature Cited:
    - Wingate, Janet L., 1994.  

    Wingate (1994) does not accept infraspecific names in E. elymoides.

    Literature Cited:
    - Shaw, Robert B., 2008.  

    Shaw (2008) accepts subsp. brevifolius and therefore subsp. elymoides.

    Literature Cited:
    - Mason-Gamer, Roberta J., Melissa M. Burns, and Marianna Naum, 2010.

    Other articles:
    • Glossary:  allotetraploid;  

    Mason-Gamer, et al. (2010) …

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittmann (2012) do not accept infraspecific names in E. elymoides.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015) does not accept infraspecific names in E. elymoides.

     

    Other articles:
    • Golden Checklist Flora:  Elymus glaucus;  

    Elymus glaucus Buckley. “Blue Wild Rye”

     

     

    Other articles:
    • Golden Checklist Flora:  Elymus trachycaulus;  

    Elymus trachycaulus (Link) Gould ex Shinners. “Slender Wheatgrass”

     

    Literature Cited:
    - Link, Heinrich Friedrich, 1827-1833.  

    Link (1833, v. 2., p. 189) published Triticum trachycaulum grown at the Berlin garden from seeds sent by Dr. Richardson. Richardson is mentioned several times as having send seeds to Link, and Link named one species after him, Stipa richardson (=Achnatherum richardsonii)

    Original Text
    28. Tr. trachycaulum. * Folia plana rigidiuscula striata scaberrima. Spica longiuscula , spiculae 5—7 florae. Valvae septemnerviae acutatae. Valvula ext. aristato-acutata. — Semina ex itinere in Americam borealem occidentalem attulit clar. Dr. Richardson nobisque dedit. ♃. T. Gramen ad 4 pedes in Horto allum. Caulis superne pilis brevibus rigidis asperrimus. Vaginae scabrae striatae, ligula vix ulla sed auricula ad oram vaginae; lamina ped. circiter longa sulcata asperrima 3 lin. lata. Spica ad ped. longa ; spiculae 8 liu. longae ; valvae valvulis parum breviores, valvulae exter. laeves superne in nervis aristaque asperne, arista lin. longa.

    Literature Cited:
    - Shinners, Lloyd H., 1954.  

    Shinners (1954, p. 28) published Elymus trachycaulus either from a manuscript by Gould, or knowing Gould was intending to publish it.

    Original Text
    Elymus trachycaulus (Link) Gould, ined. Triticum trachycaulum Link, Enum. Pl. Hort. Reg. Berol. Altera 2: 189. 1833. Elymus pauciflorus (Schweinitz) Gould, 1947; not Lamarck, 1791. Known in Texas from the Panhandle. The orthography follows that of Link, using second declension endings instead of the more usual third declension form — caulis.

    Frank W.Gould (1913-1981) was an American agrostologist who earned his PhD from University of California at Berkeley, and served most of his career at the S. M. Tracy Herbarium at Texas A & M University. He was the author of 80 definitive treatments on grasses, four grass manuals, and the well known textbook Grass Systematics.

     

    Other articles:
    • Golden Checklist Flora:  Elymus virginicus L.;  

    Elymus virginicus L. “Virginia Wildrye”

     

     

    Other articles:
    • Golden Checklist Flora:  Eragrostis cilianensis;  

    Eragrostis cilianensis (All.) Vignolo ex Janch. “Stinkgrass”

     
     

    First published as Poa cilianensis by Allioni (1785). The description is in Latin, and the illustration is not particularly informative.

     

    Other articles:
    • Golden Checklist Flora:  Eragrostis pectinacea;  

    Eragrostis pectinacea (Michx.) Nees ex Steud. “Tufted Lovegrass”

    There is disagreement whether to list authorship of this name as “(Michx.) Nees ex Steud.” or simply as “(Michx.) Nees.”

    Literature Cited:
    - Michaux, Andre, 1803.  

    Michaux (1803) published the grass as Poa pectinacea as found in fields of Illinois. It might be interesting to determine the relationship between the Illinois of today, and the Illinois of Michaux's time.

    Literature Cited:
    - Nees von Esenbeck, Christia Gottfried Daniel, 1841.  

    It appears to me that Nees (1841, p. 374) in his Flora of Southern Africa mentions a new name in passing wondering whether his Eragrostis homomalla N. ab E. could be distinguished from E. pectinaceam Michx. In the process Nees assumed that Michaux's Poa pectinaceum was in fact Eragrostis pectinacea.

    Original Text
    ... Differt ab omnibus cognitis huius affinitatis ramis paniculae alterius lateris patentibus quin etiam refractis, alterius contra erectis aut adpressis, quo charactere accedente facilius iam distinguitur as Er. pectinacea Michx., veree distincta specie, neque cum Er. pilosa coniungenda, cui ramuli paniculae rigidiores quidem sunt, at vero longius a basi divisi, et axillae praeterea pilosi.
    Adnot. An huius loci Er. verticillata Link. Hort. Ber. I. p. 189. (excl. syn.) an potius ad Er. pectinaceam spectans ?
    This is a long way from validly publishing a name, at least according to current standards, yet some authorities give Nees authorship.

    Literature Cited:
    - Steudel, Ernst Gottlieb, 1855.  

    The first time Eragrostis pectinacea appears in print as a nominated name was in Steudel (1855, v. 1, p. 272) in the group of north American species (Species Americae septentrionalis.)

    Original Text
    132. E. PECTINACEA. Michx. (Fl. Am. I. 69. sub: Poa.) ... [Latin diagnosis omitted] ... P. pilosa Muhlbrg. [Circle with dot – probably means plant is an annual.] Am. sptr.
    Steudel does in a way acknowledge Michaux's basionym of Poa pectinacea.

     

    Other articles:
    • Golden Checklist Flora:  Eriocoma;  

    Eriocoma Nutt. in the Golden s.l. Flora

    Nuttall (1818, v. 1, p. 40) …

    63. * ERlOCOMA.† (Silk-grass.)
    Calix 2-valved, 1 -flowered; valves gibbous and coarctate above, longer than the corolla, both 3-nerved and cuspidate. Corolla 2-valved, roundish; valves coriaceous, vested with a silky wool, the outer valve terminated by a short triquetrous deciduous awn. Anthers bearded. Seed large, somewhat spherical.
    Flowers dichotomously paniculate, peduncles flexuose, capillary, and clavulate. Leaves very long, involute and subulate, nodes of the culm distant, entirely sheathed.

    † From εξιον, wool, and χομη, a head of hair. A grass producing a fastigiate tuft if silky hair upon the glume of the corolla.

     

    Other articles:
    • Golden Checklist Flora:  Eriocoma hymenoides;  

    Eriocoma hymenoides (Roem. & Schult.) Rydb. “Indian Rice Grass”

     

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814, v. 2, sup., p. 728) was the first to publish a name, using Stipa membranacea stating that he saw the grass in Bradbury's herbarium. However, that name was illegitimate because it was previously used by Linnaeus (1753) for a grass growing in “Hifpania” (Spain).

    Literature Cited:
    - Roemer, Johann Jacob, and Josef August Schultes, 1817-1830.  

    Roemer & Schultes (1817, vol. 2, p. 339) published the first valid name for this grass. Their description is identical to that of Pursh (1814), so I think it is safe to assume that the type of Bradbury was used.

    Original Text Translation and Comments
    29. S. hymenoides ; foliis convoluto-filiformibus glabris, paniculâ laxâ, pedicellis flexuosis, calycibus membranaceis trivervibus longe acuminatis, corollis calyce brevioribus sericeo-villosis, villis corollam superantibus, aristâ nudâ rectâ calyce paulo longiore. S. membranacea Pursh Fl. Amer. Sept. II p. 728. 29. S(tipa) hymenoides, the leaves twisted-filiform, glabrous, the panicle loose, the pedicels flexuouse, calyx 3-nerved gradually acuminate, membranous, corolla shorter than calyx sericeous-villous, awn nude [glabrous?] straight, slightly longer than calyx.
    Altitudo 18 pollicum: spicae ovales, pilis longis sericeis tectae. Ad littora fluvii Missouri.   About 18 inches tall; heads ovate, long silky hairs when young. On the shores of the Missouri river.

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, p. 40) published a new genus Eriocoma and, apparently not aware of Roemer & Schultes (1817), above, published a valid name for Pursh's Stipa membranacea. This, if course, was s superfluous name.

    Original Text Translation and Comments
    63. * ERIOCOMA. † (Silk-grass.)  
    Calix 2-valved, 1-flowered; valves gibbous and coarctate above, longer than the corolla, both 3-nerved and cuspidate. Corolla 2-valved, roundish; valves coriaceous, vested with a silky wool, the outer valve terminated by a short triquetrous deciduous awn. Anthers bearded. Seed large, somewhat spherical. triquetrous = having three corners or salient angles or edges specifically
    Flowers dichotomously paniculate, peduncles flexuose, capillary, and clavulate. Leaves very long, involute and subulate, nodes of the culm distant, entirely sheathed.  
    Stipa membranacea. Pursh, vol. ii. In Supplement. P. 729 nom. Illeg.
    1. Cuspidata, C.  
    Description. Root perennial; culm 2 to 3 feet high, simple; panicle spreading, dichotomous, flowers by pairs, peduncles capillary flexuosa, clavulate at the summit. Leaves very long, filiform and convolute, a little asperate on the margin, (often more than a foot in length); vagina half a foot, entirely sheathing the stem and the panicle before evolution; ligula entire, conspicuous. Calix 2-valved, 1-flowered, valves membranaceous, about twice the lendth of the corolla, ventricose and gibbous, above the corolla contracted; both 3-nerves, the lateral nerves only about one third the length of the glume, the central nerve ending in a cusp or short awn margined by the glume at its base, nerves a little pubescent. Corolla 2-valved, short, nearly oval, in fruit almost spherical, valves coriaceous, vested with an exserted silky villus, extending beyond the corolla, the dorsal valve terminated by a triquetrous pungent deciduous awn scarcely the length of the calix, at first perfectly erect, afterwards a little bent; Stamina 3, scarcely exserted beyond the valves of the corolla; anthers small, brown, bifid at both extremities, above terminated by small pubescent tufts. Style 1. Stigmas 2, short, hirsutely villous. Ovarium sheathed by a 3-leaved perisporium(or nectary). Seed nearly spherical.  
    This genus is nery nearly allied to Oryzopsis, but at the same time sufficiently distinct both in habit and character; having a culm with remarkable long sheathing and almost filiform subulate leaves, a dichotomous spreading panicle, a ventricose, coarctate, awned calix twice the length of the corolla, which last is furnished with a deciduous awn, and a long silky villus.  
    Habitat. On the grassy plains of the Missouri, from the Arikaree village to the Northern Andes? Flowers in June and July.  

    † From εξιον, wool, and χομη, a head of hair. A grass producing a fastigiate tuft of silky hair, upon the glume of the corolla.

    I may not have the Greek characters correct, some of them were very hard to see in the BHL image.

    Literature Cited:
    - Piper, Charles V., 1906.  

    Frederick V. Coville, writing in the Preface to Piper (1906) Flora of the State of Washington states Piper's obligation “ ... to Mr. P L. Ricker and Mr. W. F. Wight, of the Department of Agriculture, for assistance in bibliography; ...”

    Piper (1906, v. 11, p. 109) published Oryzopsis hymenoides crediting Ricker as the author. [Ricker, Percy Leroy (1878-1973)]

    4. Oryzopsis hymenoides (Roem. & Schult.) Ricker.
    Stipa hymenoides Roem. & Schult. Syst. 2: 339. 1817.
    Eriocoma cuspidata Nutt. Gen. 1: 40. 1818.
    Stipa membranacea Pursh, Fl. 2: 728. 1814, not L. 1753.
    Oryzopsis membranacea Vasey, U.S. Dept. Agr. Div. Bot. Bull. 12: pl. 10. 1891.
    Eriocoma membranacea Beal, Grasses N. Am. 2: 232. 1896.
    Oryzopsis cuspidata Benth; Vasey, U.S. Dept. Agr. Spec. Rep. 63: 23. 1883.
    Type Locauity: “On the banks of the Missouri.”
    Rance: British Columbia to California, east to Alberta and New Mexico.
    SPECIMENS EXAMINED: Douglas County, Spillman, May, 1896; Sandberg & Leiberg, 281; Ellensburg, Piper, May, 1897; North Yakima, Henderson, May, 1892; Mrs. Steinweg in 1894; Pasco, Piper 2964; Hindshaw 33; Sunnyside, Cotton 416; Rattlesnake Mountains, Cotton 421; Walla Walla, Lyall, June, 1860; Sprague, Sandberg & Leiberg, June, 1893; Kittitas County, Vasey 85; Wallula, Cotton 1030, 1048.
    ZONAL DISTRIBUTION: Upper Sonoran.

    Literature Cited:
    - Rydberg, Per Axel, 1912.  

    Rydberg (1912, p. 102) formed the new combination of Eriocoma hymenoides (R. & S.) Rydb., as strictly a nomenclatural correction.

    Literature Cited:
    - Barkworth, Mary E., 1993.  

    Barkworth (1993) … based on morphological and anatomical evidence.

    Literature Cited:
    - Peterson, Paul M., Konstantin Romaschenko, Robert J. Soreng, and Jesus Valdés Reyna, 2019.  

    Peterson, et al., 2019, reorganized a lot of Stipeae, but primarily broke up Achnatherum, placing the New World members in Eriocoma, and retaining Achnatherum as a strictly Euraisian genus.

     

    Other articles:
    • Golden Checklist Flora:  Eriocoma robusta;  

    Eriocoma robustum (Vasey) Romasch. “Sleepygrass”

     

    Literature Cited:
    - Coulter, John M., 1890.

    Locations: Chinati Mountains.  

    John Coulter (1890, p. 56) first published robusta as a variety of Stipa viridula (=Nassella viridula). Coulter gave Vasey authorship of the name, though it does not appear Vasey ever published it himself. Vasey could have identified the grass and written the manuscript from which Coulter published the name. However, in that case, one would expect the author to be “Vasey ex Coulter.”

    Original Text
    714. Stipa viridula Trin., var. robusta Vasey, n. var. Culms densesly tufted, 12 to 18dm high, stout, leafy : lower sheaths loose and broad, longer than the internodes; blades flat and wide or involute above, often 6dm long, scabrous: panicle dense and large, erect, 25 to 40cm long: empty glumes 10mm long, three to five nerved, callus short, densely hairy. — Chenate Mountains (Presidio county). Ranges from Colorado to Mexico.

     

    Other articles:
    • Golden Checklist Flora:  Eriocoma scribneri;  

    Eriocoma scribneri (Vasey) Romasch. “Scribner Needle Grass”

     

    Literature Cited:
    - Vasey, George, 1884.

    Locations: Santa Fe.  

    George Vasey (1884, p. 125) described Scribner Needle Grass from a collection made near Santa Fe, New Mexico. He doesn't say who made the collection or when. However, there are two vouchers in SEINet with type-status; one with an image at NY and labeled a co-type, and a record at MO labeled an isotype.

    Original Text
    New Grasses.
    By George Vasey
    Stipa Scribneri. — Culms 2-3 ft. high, stout, erect; lower leaves half as long as the culm, smooth, flat below, becoming involute at the long acuminate point; upper sheath enclosing the base of the panicle, which is narrow, erect, and 6-8 inches long, the branches in twos or threes and appressed; outer glumes unequal, lower one 6-7 lines, upper about 5 lines long, both 3-nerved, acuminate; flowering glume 3-5 lines long, hairy, hairs longer above, and at the apex forming a white crown a line or more long; awn rather slender, 8-9 lines long, not hairy; stipe short, very acute, pubescent; palet less than a line long, obtuse and adherent to the grain.
    Differs from S. viridula particularly in the unequal glumes, the hairy crowned flowering-glumes, the more slender awn, and the very short palet.
    Collected on dry hill-sides at Sante Fe, New Mexico.
    At the time Vasey (1884) named the grass for him, Frank Scribner was the botanist for the Northern Transcontinental Survey. In May 1885 he was appointed an assistant in the USDA Division of Botany, and in 1894 became the leader of the new USDA Division of Agrostology.

     

    Other articles:
    • Golden Checklist Flora:  Glyceria striata;  

    Glyceria striata (Lam.) Hitchc. “Striate Manna Grass”

     

    Literature Cited:
    - Lamarck, Jean-Baptiste Pirre Antoine de Monet de, 1791.  

    Lamarck (1791, 1(1), p. 183) published Poa striata from grasses collected in Virginia and Carolina.

    Original Text
    984. POA ftriata.
    P. panicula diffufa capillari, fpiculis glabris fubquinqueflorus, corollis exquifite ftriatis.
    E Virginia, Carol. Cal. brevis. F. glabra

    Literature Cited:
    - Hitchcock, Albert Spear, 1928.  

    Hitchcock (1928, p. 157) published Glyseria striata without explanation or comment.

    Original Text
    Glyceria striata (Lam.) Hitchc.

    Poa striata Lam. Tabl. Encycl. 1:183. 1791.
    Poa nervata Willd. Sp. Pl. 1:389. 1797.
    Glyceria nervata Trin. Mem. Acad. St. Petersb. VI. Math. Phys. Nat. 1:365. 1830.
    Panicularia nervata Kuntze, Rev. Gen. Pl. 1:783. 1891.

     

    Other articles:
    • Golden Checklist Flora:  Hesperostipa comata;  

    Hesperostipa comata (Trin. & Rupr.) Barkworth. “Needle and Thread”

     

    Literature Cited:
    - Michaux, Andre, 1803.  

    Credit probably goes to Andre Michaux (1803) for first recognizing H. comata who noted the grass lives “ … in the rocky mountains from the Hudson to Canada.” Unfortunately, Michaux applied Stipa juncea to this grass, a name that Linnaeus had already applied to a grass occurring in Switzerland and France.

    Literature Cited:
    - Pursh, Frederick, 1814.  

    Pursh (1814) also applied S. juncea to a Lewis & Clark collection made July 8, 1806, made “ … Valleys of the Missouri in the Rocky Mountains.”

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818) also applied S. juncea to his collections on the grassy plains of the Missouri.

    Literature Cited:
    - Trinius, Carl Bernhard, and F. J. Ruprecht, 1842.  

    Trinius and Ruprecht (1842, p. 45) ...

    Original Text
    51. Stipa comata n. Paniculae pl. min. implexo-contractae radiis subternis, aliis fere a basi — , aliis superius floriferis; glumis subulatis, subaequalibus, valvula inferiore 5 lineali undique brevepilosa subduplo longioribus; arista subpersistente, torlili, plicata et varie inflexa, subsexpollicari; anthcris barbatis.
    Stipa juncea Nuttall (non L.) Gener. 1. (1818) p. 58? ad ripas Missouri et in Virginia abundans itinerantibus molestissima. «Arista laevis, gracilis, vix contorta, semipedalis. »
    Stipa capillata Hooker! Fl. Bor. Amer. 2. (1840) p. 237.
    Carlton House Fort ad fl. Saskatchawan (Drumond) ; ad ripas et in planitie fl. Columbia prope «Missouri Portage» (Douglas).
    Simillima Stipae capillatae, sed flosculo undique et ad apicem usque barbatulo, setulis aristarum hinc inde longioribus, denique antheris distincte barbatis diversa. Reliqua ut in St. capillata; specimen suppetens pedale.

    Stipa juncea Nuttall (1818), which he described as a variety of Stipa juncea L., is an illegitimate name. The name was unavailable, having been previously used by Linnaeus for a grass found in Switzerland and France.

    Literature Cited:
    - Barkworth, Mary E., 1993.  

    Barkworth (1993) described Hesperostipa as a North American endemic that is distinct from the Eurasian Stipa s. s. and more closely allied to the South American genera of Piptochaetium and Nassella.

     

    Other articles:
    • Golden Checklist Flora:  Hordeum brachyantherum;  

    Hordeum brachyantherum Nevski. “Meadow Barley”

    The name was published in 1936 in Sergei Nevski, a Russian botanist who worked at the Main Botanical Garden in Leningrad. Most collections before that time were originally determined Hordeum nodosum L.

    Most plants are tetraploid, though there is a diploid in California. A hexaploid has also been found.

     

    Other articles:
    • Golden Checklist Flora:  Hordeum jubatum;  

    Hordeum jubatum L. “Foxtail Barley”

     

     

    Other articles:
    • Golden Checklist Flora:  Hordeum pusillum;  Nuttall, 1818, publication details;
    Full Size ImageColl. No. 1453, Hordeum pusillum  

    Hordeum pusillum Nutt. Little Barley.

     

    Literature Cited:
    - Nuttall, Thomas, 1818.  

    Nuttall (1818, v. 1, p. 87) ...

    Nuttall's Text My interpretation
    119. HORDEUM. L. (Barley)  
    Calyces lateral, 2-valved, mostly 1-flowered, aggregted by threes, so as to resemble a setaceous 6-leaved involucrum; the central flower sessile, the lateral ones stipitate, usually sterile. Corolla 2-valved, acute; exterior valve awned.  
    Very nearly allied both by habit and character to the preceding genus. Flowers spiked, imbricated mostly in 2 rows; calycine involucrum setaceous, 6 leaved, division approaching by pairs. In the H. hexastichon, the flowers are imbricated in 6 ranks, because all the flowers are hermaphrodite; probably a mere effect of cultivation. [The preceding genus was ElymusEd.]
    Species. 1. H. vulgare. Cultivated … 2. * pusillum Lateral masculine or nertral flowers awnless, acute; four internal calicina glumes, coriaceous and dilated, those of the hermaphrodite sublanceolate; internal valve of the lateral masculine flower, subsemi-ovate. [The description of H. vulgare is skipped. – Ed.]
    Culm 4 to 6 inches, decumbent, or somewhat genoculate at the base. Leaves rather glaucous, a little pubescent on the under surface, striate, about one and a half inches long, and almost obtuse; uppermost sheath tumid and very smooth, embracing the spike. Spike linear; about one and a half inches long. Glumes by threes, distichally imbricated. Lateral imperfect flowers awnless, acute; central sessile flower awned, the awn almost exactly the length of that of the subtending calix; awns scabrous. Calix smooth, nerveless, exterior valve in the outer flowers setaceous from its base, the inner valves obliguely dilated, and rigidly coriaceous, all awned, the inner divisions of the lateral flowers, appearing nearly semi-ovate, the central ones sublanceolate. Corolla nerveless, the inner valve furnished with a short awn, arising from its base. Nearly allied, apparently, the the H. maritimum.  
    On the arid and saline plains of the Missouri.  
    3. jubatum. On the calcareous islands of Lake Huron and Michigan, also on the banks of the Missouri.  
    The genus Hordeum exists chiefly in Europe, extending into Northern Africa, and Tartary in Asia. The 2 species above described are natives of North America, and the F. jubatum is also common to Smyrna. [One would assume that Nuttall was referring to Smyrna in what we now call Turkey. However, there are towns named Smyrna in Georgia, Tennessee, North Carolina, Delaware, Michigan, and Maine, and Nuttall may have been in or near some of those towns in his travels. – Ed.]

       

    Koeleria Pers. “Junegrass”

    Koeleria is a common and widespread genus of plants in the grass family, found on all continents except Antarctica and on various oceanic islands. It includes species known generally as Junegrasses. The genus was named after German botanist Georg Ludwig Koeler (1765–1807).

     

    Other articles:
    • Golden Checklist Flora:  Koeleria macrantha;  

    Koeleria macrantha (Ledeb.) Schult. “Prairie Junegrass”

     

    Literature Cited:
    - Ledebour, Carl (Karl) Friedrich von, 1812.  

    Ledebour (1812, p. 515) published the basionym of our grass, Aira macrantha Ledeb. as one among “Sixty New Plants Indigenous in Imperial Russia.”

    Original Text Comments
    2. Aira macrantha.  
    A. panicula oblonga subspicata interrupta mutica, glumis acuminatis, flosculis calycem excedentibus, foliis brevissimis glabiis, ligula elongata bilida margine lacera.  
    Culmi caespitosi , sesquipalmares , florentes ultra medium aphylli.  
    Folia brevissima (vix unciam longa) , lanceolata , plana, glabra.  
    Ligula elongata, ultra mediam bifida, margine lacera.  
    Panicula subspicata, oblonga, interrupta, rhachis scabra, trîangularis; rami brevissimi, tri - vel quadriflori.  
    Spiculae ovatae, biflorae.  
    Flosculi calycem excedentes ; glumae omnes acnminatae, compressae, ad carinam serrtilato - scabrae.  
    Hab. in jugo montium Jablonnoi - Chrebet. ♃ Yablonovyy Khrebet, 350-600 km. east of and roughly paralleling Lake Baikal, north of Mongolia.
    Obs. Differt haec ab A. cristata folîis omnibus latioribus glabris, ligula elongata ; ab A. glauca flosculis calycem excedentibus ; ab utraque autem caule humiliori a floribus majoribus diversa.  

     

    Other articles:
    • Golden Checklist Flora:  Leymus triticoides;  

    Leymus triticoides (Buckley) Pilger. “Beardless Wildrye”

     

       

    Lolium arundinaceum (Schreb.) Darbysh. “Tall Fescue”

     

    Literature Cited:
    - Gmelin, Johan Georg, 1747-1769.
    Full Size ImageGmelin's (1747) description of Tall Fescue.  

     

    Literature Cited:
    - Gmelin, Johan Georg, 1747-1769.
    - Scheuzer, Johann Jacob, 1719.
    - Schreber, Johann Christian Daniel von, 1771.  

    There seems to be general agreement (by extensive Google search) that what is called “Tall Fescue” is the grass known by its basionym of Festuca arundinacea Schreb.

    Original TextInterpretation and Comments
    1000. FESTVCA (arundinacea) panicula ſpicata ſtricta, ſpiculis oblongis erectis paucifloris, ariſtatis, calycibus anguſtatis. Gmel. Sibir. 1. p. 111. 1000. Festuca arundinacea panicle spike strict, spikes oblong erect awn, calyx angular. Gmelin, Flora Sibirica sive Historia Plantarum Siberiae. 1. p. 111.
    Gramen arundinaceum, locuſtis viridi-ſpadiceis loliaceis brevius ariſtatis. Scheuchz. Agr. p. 266. t. 5. f. 18. Grass reed-like, … Scheuzer, Agrostographia sive Graminum, … p. 266. t. 5. f. 18.
    In prato acclivi hinter dem Biniz, loco humido. In the steep field beyond the Biniz [East Germany], in a damp location.

    • “arundinacea” would mean cane-like from arundo (“cane”) +? -aceus.
    • “loliaceis” could mean chaffy, but Lolium itself is a name given by Virgil to a troublesome weed.
     

    There is less agreement as to its current accepted name, whether it should be Festuca arundinacea Schreb., Lolium arundinaceum (Schreb.) Darbysh., or Schedonorus arundinaceus (Schreb.) Dumort.

    Literature Cited:
    - Ackerfield, Jennifer, 2015.
    - Baldwin, Bruce G., Douglas H. Goldman, David J. Keil, Robert Patterson, and Thomas J. Rosatti, 2012.
    - Harrington, H. D., 1954.
    - Shaw, Robert B., 2008.
    - Weber, William A., and Ronald C. Wittmann, 2012.
    - Wingate, Janet L., 1994.  

    Starting close to home, most Colorado authors use Schedonorus arundinaceus (Schreb.) Dumort., including Ackerfield (2015), Weber & Wittman (2015), and Shaw (2008).

    Wingate (1994) uses Festuca arundinaceus Schreb., although it is a older reference. Harrington (1964, 2nd ed.) used Festuca elatior var. arundinaceae (Screb.) Celak..

    By way of comparison, the Jepson Manual of California (Baldwin, et al., 2012) retains Festuca arundinacea Schreb.

     

    On a regional level, the Southwest Biodiversity Network (SEINet) taxon tree has kind of a split personality with regard to “Tall Fescue.” The current (3 October 2019) taxon tree accepts both Lolium arundinaceum (Schreb.) S.J. Darbyshire and Schedonorus asundinaceus (Schreb.) Dumort.

    Poaceae
           [Gramineae]
             Lolium 
                   [Lolium x aschersoniana]
                     Lolium arundinaceum  
                           [Festuca arundinacea]
                           [Festuca elatior subsp. Arundinacea]
                           [Festuca elatior var. arundinacea]
             Schedonorus 
                     Schedonorus arundinaceus 
                           [Avena secunda]
                           [Bromus arundinaceus]
                           [Bromus elatior]
                           [Festuca elatior f. aristata]
                           [Festuca elatior f. elatior]
                           [Festuca fenas]
                           [Festuca mediterranea]
                           [Festuca orientalis]
                           [Festuca phoenix]
                           [Festuca uechtritziana]
                           [Poa elatior]
                           [Poa kunthii]
                           [Poa phoenix]
                           [Poa uliginosa]
                           [Schedonorus elatior]
                           [Schedonorus phoenix]
                           [Tragus elatior]
    

    One consequence is that both names have to be entered into a query when searching for collections of “tall fescue.”


    Full Size ImageCollections of “tall fescue” entered into SEINet  

    The map at left (3 Oct 2019) shows the locations of Colorado collections of “tall fescue” that have coordinates entered into SEINet. Within Jefferson County, collections have primarily been made at Chatfield Farms, Hildebrand Park and Deer Creek Canyon Park. There are no collections of this grass from Golden s.l.

    Literature Cited:
    - Henson, James F., 2001.  

    USDA Plants accepts Schedonorus arundinaceus (Schreb.) Dumort. as a conserved name. Festuca arundinacea Schreb and Lolium arundinaceum (Schreb.) S.J. Darbyshire are treated as synonyms. Perhaps somewhat ironic is the USDA NRCS plant guide for “Tall Fescue” (Henson, 2001) which accepts the name of Lolium arundinaceum (Schreb.) S.J. Darbyshire.

    Literature Cited:
    - Soreng, Robert J., Edward E. Terrell, John Wiersma, & S. J. Darbyshore, 2001.  

    Conservation of Schedonorus arundinaceus (Schreb.) Dumort. was proposed by Soreng, et al. (2001). S. arundinaceus Roem. & Schult. Syst. Veg. 2: 700. 1817 was rejected because it is not the basionym of any currently accepted taxon. Rather is it a synonym of Scolochloa festucacea (Willd.) Link.

    Literature Cited:
    - Fribourg, H. A., D. B. Hannaway, and C. P. West, 2009.  

    Tall Fescue Online monograph. (https://forages.oregonstate.edu/tallfescuemonograph/turf/invasive_species)

    Hexaploid tall fescue [Lolium arundinaceum (Schreb.) Darbysh. = Schedonorus arundinaceus (Schreb.) Dumort., formerly Festuca arundinacea Schreb. Var. arundinacea] is an agronomically vital member of the grass family that has been characterized based on morphological characteristics, interfertility relationships, and, more recently, genetic criteria such as gene sequences. Efforts to improve tall fescue for forage or turf and to gain insight into processes of grass evolution hinge on an accurate depiction of the interrelationships that exist between this species and other Lolium and Festuca grasses. The evolution of hexaploid tall fescue, and a majority of grasses, has involved interspecific hybridization that obscures species boundaries but, more importantly, serves as a rapid means of combining distinct genomes into novel progeny with enhanced evolutionary potential. We discuss here the systematics of the Festuca-Lolium complex within the context of such processes and highlight the dynamic and often confounding evolutionary history that characterizes tall fescue and its relatives (Fribourg, et al., 2009).

    Festuca arundinacea Schreb. Tall fescue (https://www.invasiveplantatlas.org/subject.html?sub=3037)

    Forage Identification: Tall fescue. Department of Plant Sciences. Tall fescue (Festuca arundinacea (Schreb.) Darbysh) (http://www.uwyo.edu/plantsciences/uwplant/forages/grasses/tall-fescue.html)

    Phylogeny of tall fescue and related species using RFLPs. Theor Appl Genet. 1994 Aug;88(6-7):685-90. doi: 10.1007/BF01253971. (https://www.ncbi.nlm.nih.gov/pubmed/24186163)

    Melanie L Hand, Noel OI Cogan, Alan V Stewart & John W Forster Evolutionary history of tall fescue morphotypes inferred from molecular phylogenetics of the Lolium-Festuca species complex BMC Evolutionary Biologyvolume 10, Article number: 303 (2010) (https://bmcevolbiol.biomedcentral.com/articles/10.1186/1471-2148-10-303)

     

    Other articles:
    • Golden Checklist Flora:  Muhlenbergia andina;
    • Field Notes:  Coll. No. 2392, 24 Jul 2020;
    Full Size ImageColl. No. 2392, Muhlenbergia andina.
    Full Size ImageRhizome of Coll. No. 2392, Muhlenbergia andina.  

    Muhlenbergia andina (Nutt.) Hitchc. “Foxtail Muhly”

     

    Literature Cited:
    - Nuttall, Thomas, 1848b.
    - Spitzzeri, Paul R, 1998.

    Locations: Colorado River.  

    Nuttall (1847, p. 187) described the grass from a collection by William Gambel.

    Original Text
    CALAMAGROSTIS.
    § *Trichagrostis. — Spikelets one-flowered, the flower sessile, with long hairs at the base. Glumes two, subequal, membranaceous, acute, longer than the flower, the lower with a short terminal awn. Paleae two, very acute, the lower carinate, ending in an exserted capillary awn ; the upper one-nerved, acuminate. Caryopsis free, cylindric-oblong, much shorter than the glume.
    C. *andina. A simple stemmed small grass, with a compressed culm, about two or three feet high ; ligules minute ; flowers in a short, paniculate, narrow spike, about three or four inches long, half an inch wide, with a habit very distinct from the general character of the genus ; glumes very long and slender, each with about a single nerve ; very narrow and membranaceous ; one of them distinctly awned, the other acuminate ; pappus copious, longer than the small flower ; no lateral rudiment of another flower.
    Hab. In Upper California, on the Colorado of the West.
    Gambel traveled to Los Angeles with the Rowland-Workman party, and left with them for California on September 1, 1941 (Graustein, p. 350; see also https://en.wikipedia.org/wiki/John_A._Rowland). Assuming they took the northern route of the Old Spanish Trail, Gambel would have passed through territory known where Muhlenbergia andina is known to occur in proximity to the Colorado River near Moab, Grand County, Utah. If they traveled the southern route, Gambel might have seen it near Tsegi Canyon or Betakin Canyon. The route traveled might be stated in Spitzzeri (1998). Unfortunately I have been able to break through the paywalls of JSTOR or UC Press, nor obtain a copy of the article through ResearchGate.

    Literature Cited:
    - Hitchcock, A. S., 1920.  

    Hitchcock (1920) moved C. andina to Muhlenbergia.

    Original Text
    Vaseya Thurb., in Gray, Proc, Acad. Phila. 1863: 79. 1863. The type is V. comata Thurb., the only species described. This is Muhlenbergia andina (Nutt.) Hitchc. (Calamagrostis andina Nutt.).

    Literature Cited:
    - SEINet, 2019+.

    Other articles:
    • US Highway 160:  Tsegi;

    Locations: Tsegi Canyon.  

    A GIS analysis of georeferenced collections of Muhlenbergia andina (Nutt.) Hitchc. (SEINet, 2021, n=299), within 100 km of the Colorado River, and within 3 km of a trace of the Old Spanish Trail, show that the most likely location of the Gambel collection is in Tsegi Canyon and vicinity, Navajo County, Arizona. There is a tiny chance that the collection was near Moab, but only if the Rowland-Workman party followed the northern route of the Old Trail. There is also a small chance that the collection was made in Piute Canyon or along Piute Creek, 23 km. northwest of Tsegi Canyon. However, the largest number of collections are found in or near Tsegi Canyon. There are three collections with four vouchers in this area. These are S. Holiday, #376, 9/9/1995 ASC63457; Wetherill, s.n., 8/22/1935, MNA B.3488; and M. J. A. Wetherill, #538, 8/22/1935, ARIZ91714 & ARIZ91715 There is a fourth collection just outside the Tsegi Canyon area: D. Roth, #1308, 8/30/2001, ASC70375. And finally, the collection in Piute Canyon is: Kenneth D. Heil and Arnold Clifford, #17190, 8/14/2001, SJNM330.

     

    Other articles:
    • Golden Checklist Flora:  Muhlenbergia asperifolia;  

    Muhlenbergia asperifolia (Nees & Meyen ex Trin.) Parodi. “Alkali Muhley”

     

    Literature Cited:
    - Trinius, C. B., 1840.
    Full Size ImageDried collection of Alkali Muhley, my collection no. 490.  

    Trinius (1845, 6e serie, t. 2, p. 95) published Vilfa asperifolia as described by Nees and Meyen. asperifolia would mean “rough leaves.” The paper was either read or communicated on 5 June 1840, but not published until July 1845.

    Original Text
    † 61) Vilfa asperifolia N. et Meyen! 6 — 10-po!Iicaris. Juba patente , subpauciflora , 4 — 5 - pollicari ; Radiis inferioribus subtripollicaribus, basi paullo nudis, compositis; Spiculis vix linealibus; Glumis parum l. paullo inaequalibus , valvulis pl. min. ↷ (l. injcriore fere ½) brevioribus, dorso hispidulis; Valvula inferiore submucronata , superiore obtusiuscula. Foliis planis, asperis,fere lineam lotis, culmeis 1 ½-pollicaribus, ramorum pollicaribus et brevioribus.
    Chile; Rio Mayno: Copiapo.
    Radix .... Culmus diffuso-ramosissimus, decumbens, glaber. Nodi plures, glabri. Vaginae nodis longiores, glabrae, saepe hiantes. Ligula membranacea, brevis. Folia patentia, aspera l. asperiuscula, in ramis subdisticha, juniora angustissima, inferiora pollicaria, superiora semipollicaria, in culmo pl. min. 1 ½- pollicaria, fere lineam lata, acuta l. acnminata , glauco- viridia. Juba (junior) basi folio subinvolucrata. Radii strictiusculi, solitarii, paullo distantes, superne scabri, compositi l. subcompositi. Pedicelli spicula multo (3 — 7-ies) longiores, hispidi. Spiculae aeneae, acutae, haud raoe biflorae. Glumae 1-nerves, acutae, dorso hispidulae, superior ipso apice sub lente strigillosa. Valvula inferior 1—l. sub-5-nervis, ex apice obtusiusculo mucronulata: mucrone hispidulo; superior obtusiuscula, 2-nervis. Antherae lineares.

    Carl Bernhard von Trinius (6 March 1778, Eisleben – 12 March 1844, St. Petersburg) was a German-born botanist and physician. As a botanist, Trinius was a specialist in grasses and described many species in his career, including Agrostis pallens, Cenchrus agrimonioides and Festuca subulata. The genus Trinia and species Trinia glauca are named after him. The so-called "Herbarium Trinii" (a collection of roughly 4000-5000 plants) was bequeathed to the botanical museum in St. Petersburg (WikiPedia, 2021).

    Christian Gottfried Daniel Nees von Esenbeck (14 February 1776 – 16 March 1858) was a prolific German botanist, physician, zoologist, and natural philosopher. He was a contemporary of Goethe and was born within the lifetime of Linnaeus. He described approximately 7,000 plant species (almost as many as Linnaeus himself). His last official act as president of the German Academy of Natural Scientists Leopoldina was to admit Charles Darwin as a member. He was the author of numerous monographs on botany and zoology. His best-known works deal with fungi.

    Franz Julius Ferdinand Meyen (28 June 1804 – 2 September 1840) was a Prussian physician and botanist. Meyen was born in Tilsit, East Prussia. In 1830 he wrote Phytotomie, the first major study of plant anatomy. Between 1830 and 1832, he took part in an expedition to South America on board the Prinzess Luise, visiting Peru and Bolivia, describing species then new to science such as the Humboldt penguin.

     

    Parodi (1928) is given credit for Muhlenbergia asperifolia in Revista de la Facultad de Agronomia y Veterinaria. Buenos Aires vol. 6: 117, p. 117. I have been unable to find this volume on the Internet.

     

    Other articles:
    • Golden Checklist Flora:  Muhlenbergia montana;  

    Muhlenbergia montana (Nutt.) Hitchc. “Mountain Muhly”

     

    Literature Cited:
    - Nuttall, Thomas, 1848b.  

    Nuttall (1848, p. 187) published Calycodon montanus from a collection by Gambel near Santa Fe, New Mexico.

    Nuttall's Text Comments and Interpretation
    *CALYCODON
    Spikelets one-flowered, the flower sessile, bearded at the base. Glumes two, unequal, shorter than the flower, membranaceous, the lower truncate, acutely three-toothed; the lower smaller, one-toothed. Paleae two, the lower sublanceolate, carinate, terminaing in a longish scabrous awn; at length indurated, with a silky pilose margin; the upper palea lanceolate, one-nerved, indurated and involute. Anthers three. Stigmas two, plumose. — A scabrous leaved grass, with a simple inarticulated culm, terminated by a loose, narrow, somewhat spiked panicle. So called in allusion to the remarkable toothing of the calyx.
    C. *montanum. Leaves short and narrow, somewhat scabrous; ligules membranaceous, elongated; panicles four or five inches long, narrow, with the branches appressed; flowers clustered on the branches, three or four together, some nearly sessile and others pedicellate; glumes variable, membranaceous and eroded at the summit, the lower, three-nerved, with three wither short, or rather long and acute teeth, sometimes with a fourth membranous tooth; the upper glume also eroded, and ending in a single tooth from the nerve; the lower palea lanceolate, carinate, scabrous, and indurated, terminated by a long, slender, scabrous awn; the inner margin silky, with soft shining hair, of which there are two tufts at the base of the paleae; the inner paleae also indurated and herbaceous in the centre, involving the germ and the stamens.
    A perennial grass, with a simple, unnjointed culm, about eighteen inches high. Somewhat allied to Muhlenbergia, (when restrained to its proper limits,) but perfectly distinct by its very remarkable glumes. The ripe seed we have not seen.
    Hab. In the Rocky Mountains, near Santa Fe, Mexico. Flowering in August.  

    Literature Cited:
    - Hitchcock, A. S., 1920.  

    Hitchcock (1920) placed Nuttall's C. montanum into Muhlenbergia.

    Original Text
    Calycodon Nutt. Journ. Acad, Phila. II 1: 186. 1848 The type is C. montanum (Muhlenbergia montana Hitchc.), the only species described.

     

    Other articles:
    • Golden Checklist Flora:  Muhlenbergia racemosa;  

    Muhlenbergia racemosa (Michx.) Britton, Stern & Poggenb. “Marsh Muhly”

     

    Literature Cited:
    - Michaux, Andre, 1803.  

    Michaux (1803, p. 53) in Flora Boreali-americana placed racemosa in Agrostis.

    Original Text
    [Agrostis] racemosa. A. culmis erectis : foliis anguste longeque linearibus, erectis : racemo oblongo , e spicis densifloris subinterrupto : glumae exterioris valvis in mucronem aristiformem desinentibus, gluma interiore mutica longioribus.
    Obs. Affinis A. lateriflorae.
    Hab. in ripis sabulosis inundatis fluminis Mississipi.

    Literature Cited:
    - Britton, Nathaniel Lord, Emerson Ellick Sterns, and Justus Ferdinand Poggenburg, 1888.  

    Britton, et al. (1888) simply published Muhlenbergia racemosa without comment, as follows:

    Original Text
    Muhlenbergia racemosa, (Michx.) (M. glomerata, Trin.)

     

    Other articles:
    • Golden Checklist Flora:  Muhlenbergia wrightii;  

    Muhlenbergia wrightii Vasey ex J.M. Coult. “Spike Muhly”

    There are five vouchers of Wright #1986, labeled Muhlenbergia wrightii: two at GH, one at MO, and two at NY. The two vouchers at GH are lebeled isotypes. Several are labeled isotypes. The voucher at MO is labeled an isotype. NY381414 is annotated “Type number of M. wrightii Vasey, Determined by Richard W. Pohl 1967.” This voucher was from the Grass Herbarium of Mr. George V. Nash, purchased 1911. The other voucher at NY (NY381413) is from the Torrey Herbarium and bears the note, “Probably a new species, but I do not feel well enough satisfied with it to describe it. J. T.”

    GH24000 does not give a location. GH24001 is annotated, “Field no 511, &slquo;511, grass, Valley of Sonora, along branches, in small compact bunches, Sept 14, 1851&srquo; east of Huachuca Mt. south of Benson. Cochise Co, Ariz!” NY381413 gives the location as New Mexico, as does NY381414.

    Literature Cited:
    - Coulter, John Merle, 1885.  

    Coulter (1885, p. 409) first published M. wrightii noting the author was “Vasey ined.”

    2. M. Wrightii, Vasey ined. Stems erect, 9 inches to a foot high or more: leaves involute, rather rigid and pungently pointed, scabrous, pale; sheaths much shorter than the internodes: panicle spike-like, 1 to 3 inches long, the two or three lowest clusters of spikelets somewhat distant : the glumes and palets scabrous, especially on the midribs ; lower glume the shorter, ⅓ to ½ the length of the flowering glume, mucronate pointed ; upper glume longer, 1-nerved and short-awned : flowering glume l-nerved, tipped by a stout rough awn about ⅓ the length of the palet. — Colorado and New Mexico.

     

    Other articles:
    • Golden Checklist Flora:  Munroa squarrosa;  

    Munroa squarrosa (Nutt.) Torr. False Buffalograss.

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Glossary:  squarrose;  

    Nuttall (1818, v. 1, p. 49) …

    75. CRYPSIS. Lamarck. (Thorn-Grass.)
    Calix 2-valved, oblong, 1-flowered, Corolla 2-valved, longer than the calix. Stamina 2 or 3. (Spike surrounded at the base by the sheath of the leaf; or the flowers collected into a leafy capitulum.)
    Culm decumbent or procumbent, extremely branched; leaves rigid and pungent; flowers collected in squarrose heads, or short and dense irregularly involucrate, lobed spikes.
    SPECIES. 1. C. * squarrosa. Stem decumbent, much branched; leaves short, all rigid, and sharply pungent; capituli squarrose, few flowered; dorsal valve of the corolla coriaceous, somewhat cleft at the point, with a shortish subulate central cusp.
    On arid plains near the “Grand Detour” of the Missouri, almost exclusively covering thousands of acres, and as pungent as thorns. ☉ Not more than 3 or 4 inches high; the flowers not collected into heads, as in the European species, but merely in squarrose terminal fascicles; the outer glume of the corolla is likewise cleft so as to present 3 short coriaceous subulate points.

    Literature Cited:
    - Torrey, John, and Asa Gray, 1857a.  

    Torrey (1859, p. 102) report on the botany of the Whipple expedition....

    Original Text Comments
    MONROA. Nov. Gen.  
    ... [Latin genus diagnosis not reproduced.] ...  
    Monroa squarrosa. Crypsis squarrosa, Nutt. Gen. 1. p. 49. Hills and ravines, Anton Chico, New Mexico ; September. “On the arid plains of the Upper Missouri, near the Grand Detour, it covers, almost exclusively, thousands of acres.&rdquo — Nutt. This grass is very distinct from Crypsis, and belongs, as we think, to the tribe Hordeaceae. In the notice of Dr. James' plants, collected in Long's 1st expedition, (Ann. Lyc, Nat. Hist. N. York, 2, p. 254,) it was intimated that is was probably a distinct genus. ... [Description not reproduced.] ... We dedicate this singular genus to Major Monro, of the East India Company's service, who has made the grasses an especial study.  

     

    Other articles:
    • Golden Checklist Flora:  Nassella viridula;  

    Nassella viridula (Trin.) Barkworth. “Green Needlegrass”

     

    Literature Cited:
    - Trinius, C. B., 1836.  

    Trinius (1836, p. 39) published Stripa viridula Trin, describing the habitat as “V. spp. Am. bor.?”

    Literature Cited:
    - Barkworth, Mary E., 1990.

    Other articles:
    • Glossary:  ecostate;  

    Barkworth (1990, p. 597) revised the circumscription of Nassella Desv. to include all species of Stipeae with strongly overlapping lemma margins; lemma apices that are fused into a crown; paleae that are highly reduced, ecostate and glabrous; long epiblasts; and lemma epidermes with very short fundamental cells having silicified cell walls. The expanded genus includes 79 species, almost all of which are South American. Our most common Nassella in Colorado is N. viridula (Trin.) Barkworth or “Green Needlegrass” N. tenuissima (Trin.) Barkworth is reported from Larimer County, and may be escaped from cultivation.

     

    Other articles:
    • Golden Checklist Flora:  Panicum capillare;  

    Panicum capillare L. “Witchgrass”

    The name was applied by Linnaeus (1753) from known locations of Virginia and Jamaica.

     

    Literature Cited:
    - Rydberg, Per Axel, 1900a.

    Other articles:
    • Golden Checklist Flora:  Pascopyrum smithii;  

    Pascopyrum smithii (Rydb.) Á. Löve. “Western Wheatgrass”

    Agropyron spicatum Scribn. & Smith nom. nov., https://www.biodiversitylibrary.org/item/110083#page/253/mode/1up

    First published as Agropyron smithii Rydb., 1900, as a segregate from A. spicatum, which by Rydberg's comments may have been somewhat muddled.

    Moved to Pascopyrum smithii (Rydb.) Á.Löve, Taxon 29: 168, 547 (1980).

    Literature Cited:
    - Löve, Áskell, 1980a.
    - Löve, Áskell, 1980b.  

    Love (1980a, p. 168) published his monotypic Pascopyrum ...

    Original Text
    Pascopyrum Love, gen. nov. (based in the description of Agropyron spicatum Scribn. & Merr. 1897, U. S. Dept. Agr., Div. Agrostol. Bull. 4:33 [“type *** Geyer, upper Missouri”], not the basionym Festuca spicata Pursh; generic type P. smithii).
    Pascopyrum smithii (Rydb.) Love, comb. nov. (based on Agropyron smithii Rudberg, 1900, N. Y. Bot. Gard. Mem. 1:64). 2n = 56. Canada: Manitoba, Sandilands, roadside. L. Cyt.T6159.

    Love (1980b, p. 547) corrected his oversight and published a Latin diagnosis ...

    Original Text
    Addendum

    The author would like to take this opportunity to correct an unfortunate misunderstanding when he, in Taxon 29(1), 1980, p. 168, failed to include the following Latin description of the new monotypic genus Pascopyrum Love: [... Latin description omitted ...] Typus generis: Pascopyrum smithii (Rydb.) Love, cf. Taxon, l.c., p. 168.

     

    Other articles:
    • Golden Checklist Flora:  Phleum pratense;  

    Phleum pratense L. “Timothy”

     

     

    Other articles:
    • Golden Checklist Flora:  Poa annua;  

    Poa annua L. “Annual Blue Grass”

     

     

    Literature Cited:
    - Vasey, George R., 1883.

    Other articles:
    • Golden Checklist Flora:  Poa arida;  

    Poa arida Vasey. “Plains Bluegrass”

    George Vasey (1883) described P. arida from a collection he made at Socorro, New Mexico, in 1881.

    Original Text
    Type specimen collected by G. R. Vasey at Socorro, New Mexico, in 1881; other specimens collected in Utah, Colorado, Kansas, and northward to British America. This is P. andina Nutt., in Herb. Phila. Acad., fide F. Lamson-Scribner, but Nuttall's manuscript name is preoccupied by a Chilian species, P. andina Trin. It may be distinguished from Poa fendleriana (Steud.) Vasey by the smaller spikelets, smaller florets, and the pubescence between the nerves of the floral glumes.

     

    Other articles:
    • Golden Checklist Flora:  Poa fendleriana;  

    Poa fendleriana (Steud.) Vasey “Muttongrass”

     

    Literature Cited:
    - Steudel, Ernst Gottlieb, 1855.  

    Steudel (1855, pars 1, pg. 278)

    Original Text
    213. E. FENDLERIANA. Steud. Radice fibrosa horizontaliter descendente ; culmo erecto simplici vaginis scabris tecto, uti tota planta pallide glaucescente ; ligula ovata obtusa brevi fimbriata; foliis angustis culmo (1—1½-pedali) brevioribus scabriusculis planis vel subconvolutis ; paniculae erectae (3-pollicaris) rhachi radiisque scaberrimis, his rigidulis alternatim 2—3-nis pauci- (1—4-) spiculatis ; spiculis ovato-oblongis 5—9-floris ; glumis tenuibus ovato lanceolatis margine late membranaceo-pellucidis carina viridibus acuminatis uninerviis ; valvula inferiore carinata herbacea (plus minus rubro-colorata) margine hyalino-membranacea ad carinam basi pilosa apicem versus aculeolata scabra praeter carinam vix nervosa ; inferiore hyalina margine herbacea et ciliata. Praecedenti videtur affinis. Fendler hrbr. nr. 932. Mexico. The preceeding was Eragrostis uninervia now placed in Diplachne

    Steudel does not mention that the grass is dioecious.

    Literature Cited:
    - Vasey, George R., 1893a.
    - Vasey, George R., 1893b.  

    Vasey (1893a, 2(2), no. 74) published Poa fendleriana. However, I have been unable to find an online version of Volume 2, Part 2.

    I assume that Vasey (1893b, v. 2, p. 297) is nearly identical in content, though it is not the original publication of Poa fendleriana.

    While the grass is named for Fendler, who only collected in New Mexico, and Steudel (1855) refers specifically to a Fendler collection, Vasey gives the geographic distribution as Southern California, Arizona, and Nevada.

    Original Text
    No. 74
    POA FENDLERIANA (Steud.) Vasey; Eragrostis Fendleriana Steud. Syn. Gram. 179 (1855).
    … [Diagnosis omitted] ...
    Southern California, Arizona, and Nevada. This species was named without description Sclerochloa Californica Munro in Benth. Pl. Hartw. 342 (1857), and was described as Atropis Californica Munro in Wats. Bot. Gal. ii. 309 (1880). In Vasey, Cat. Grasses of U. S. 81 (1885) it was again changed to Poa Californica, and by this name it is probably most widely known. Besides these proper synonyms, specimens have been wrongly named P. andina Nutt. and P. tenuifolia Nutt. It is apparently completely dioecious, and this fact is probably one cause for the multiplicity of names and the confusion regarding the species. The staminate panicle is more open and has fewer spikelets than the pistillate, and the glumes are narrower, thinner, less pubescent, and slightly smaller. The spikelets of the staminate plant are flat, with divergent florets open at maturity; while those of the pistillate plant are more turgid, with merely carinate margins, the florets remaining erect and closed.

     

    Other articles:
    • Golden Checklist Flora:  Poa palustris;  

    Poa palustris L. “Fowl Bluegrass”

     

     

    Other articles:
    • Golden Checklist Flora:  Poa secunda;  

    Poa secunda J. Presl. “Sandberg Bluegrass”

     

     

    Other articles:
    • Golden Checklist Flora:  Poa tracyi;  

    Poa tracyi Vasey “Tracy's Bluegrass”

     

    Literature Cited:
    - Vasey, George R., 1888.

    Locations: Raton.  

    Vasey (1888, p. 48) described Poa tracyi collection made by Prof. S. M. Tracy near Raton, New Mexico.

    Original Text Comments
    Prof. S. M. Tracy collected in New Mexico, Arizona, S. California, Nevada, Utah and Western Colorado, over 200 species of grasses, many of them interesting, particularly the Oryzopsis Webberi, collected at Reno, Nevada, the first specimens which have been collected since those by Mr. Lemmon, on which the species was founded by Dr. Thurber, as Eriocoma Webberi. Currently known in Colorado as Achnatherum webberi (Thurb.) Barkworth, though it appears we will soon revert to calling it Eriocoma webberi Thurb. It is not known from Jefferson County, there being only one collection from Grand Junction.
    The following appear to be new : Poa Tracyi, from mountain sides near Raton, New Mexico. It is of the flexuosa group, 2 to 3 feet high, with short leaves, panicle 4 to 6 inches long, spikelets large, flowing glumes strongly five-nerved, scabrous-pubescent, hairy at the base ; Diplachne Tracyi, near D. fascicularis, 1 3^ to 2 feet high, erect, and narrow, leaves equaling the culm, panicle long, branches appressed, spikelets seven to nine-flowered, flowering glumes with two acute lobes at the apex and a short awn between them. In clumps growing in ditches at Reno, Nevada.  

     

    Other articles:
    • Golden Checklist Flora:  Pseudoroegneria spicata;  

    Pseudoroegneria spicata (Pursh) A. Löve. “Bluebunch Wheat Grass”

     

    Literature Cited:
    - Moulton, Gary E., 1999.
    - Pursh, Frederick, 1814.

    Other articles:
    • Golden Checklist Flora:  Pursh, 1814, publication details;  

    (Pursh, 1814-1816, v. 1, p. 83) published Festuca spicata from a dried specimen he saw in the Lewis & Clark herbarium.

    Original Text
    96. FESTUCA. Gen. pl. 119.  
    1. F. spiculis alternis sessilibus erectis subquinquefloris, floribus subulatis glabriusculis, aristis longis scbaris, foliis linearibus culmoque glabris. spicata.
    On the waters of the Missouri and Columbia rivers. June v. s. in Herb. Lewis.  

    Moulton (1999, no 142) confirms that there is a collection of the grass in the Lewis & Clark herbarium. The oldest label states the collection was made on the plains of the Columbia, June 10th, 1806. An annotation label states:

    =Agropyron spicatum (Pursh) S & S
    Type Festuca spicata
    Determined by A. S. Hitchcock
     

    Pseudoroegneria (Nevski) A. Love and P. spicata (Pursh) A. Love were published in “IOPB Chromosome Number Reports LXVI,” Taxon 29: 168 (1980). The article is in JSTOR.org but unfortunately hidden behind a paywall.

     

    Other articles:
    • Golden Checklist Flora:  Schedonnardus paniculatus;  

    Schedonnardus paniculatus (Nutt.) Trel. “Tumblegrass”

     

    Literature Cited:
    - Nuttall, Thomas, 1818.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1818, publication details;  

    Nuttall (1818, v. 1, p. 81) …

    Original Text
    113. LEPTURUS. R. Brown. (Rotbollia, species, Willd.)
    Flowers polygamous, spiked. Rachis articulated, filiform; articulations single-flowered. — Calix fixed, or growing to the rachis, 1 or 2-valved, the valve simple, or biparted.
    With the precise characters of this genus, as described by Mr. R. Brown, I am unacquainted, but satisfied with the propriety of separating plants of such dissimilar habits, as have been hitherto referred to Rotbollia, I have ventured to give it, however imperfectly.
    Species. 1. L. * paniculatus. Rachis incurved, compounded, acutely triangular, branches and summit flower-bearing; spikes on one side, subulate, compressed, unilaterals; calix 2-valved, acuminate, 1-flowered; flowers all hermaphrodite, 2-valved.
    Obs. Annual. Culm scarcely a foot high, roundish, compressed, leaves short, rigid, sheathing the base of the panicle; panicle of naked rachis, slender, rigid, angular, bearing 6 to 10, compressed, subulate, spikes on one side, not soluble or fragile at the articulations, each 1 or 2 inches long; flowers remote, on one side of the rachis. Calix rigidly fixed, of 2 unequal parallel valves closing the scrobiculum; flower 2-valved, the exterior valve resembling the calix, the interion membranaceous.
    On dry saline plains, near Fort Mandan, on the Missouri. Flowering in June.

    Literature Cited:
    - Branner, John C., and F. V. Coville, 1888.
    - Coville, F. V., 1888.  

    Original Text
    BIBLIOGRAPHY AND EXPLORATIONS.*
    The work of Nuttall. — The botanical work done in Arkansas up to 1887, when the work of the present Geological Survey began, was confined to a few individuals and a few exploring parties. The earliest explorer was Thomas Nuttall, one of the prominent early naturalists. He spent several years in the territory, as it rhen was, most of his time being occupied in botanical observations. The results of these observations were not published as a report, but at odd times and in different places. They are all long out of print.
    “Journal of Travels Into Arkansas Territory, 1819 : By Thomas Nuttall, p. 236, Phila., 1821.” This book has not been obtained by the writer, and no sketch of its contents can be given.
    “A Description of Some New Species of Plants Recently Introduced Into the Gardens of Philadelphia, from the Arkansas Territory : By THomas Nuttall. Journ Phil. Acad. Sci., Vol. II, pp. 114-123 ; Phila., 1821.” Twelve species of Arkansas plants are described.
    “Description of Two New Genera of the Natural Order Cruciferae : By Thomas Nuittall. Ibid., Vol. V, pp. 132-135.” The plants are Selenia aurea and Streptanthus maculatus ; the latter from Red River, the former from the Arkansas.
    “Collections Toward a Flora of the Territory of Arkansas : By THomas Nuttall. Transactions of the American Philosophical Society, Vol. II (New Series), pp. 139-203. Philadelphia, 1837." There are included two fungi, and several Pteridophyta, the remainder being flowering plants, some newly described. The paper appears to be a list of all the plants, so far as it goes, found in Arkansas up to the time of publication. But the Polypetalae and several orders of the Gamopetalae (one of the Compositae) are omitted.
    ...
    Nomenclature. — The oldest specific or varietal names are used in this list (not going back of Linnaeus' Species, 1753) under whatever genera they may originally have been employed. On this principle Professor William Trelease, Director of the Missouri Botanical Gardens at St. Louis, has kindly revised the list. It is but just to Professor Trelease to add, however, that the time that could be allowed for this revision was too short to admit of the necessary investigations in all cases, so that a few plants appear under questionable names current in Gray's Manual or other works in general use.
    ...
    GRAMINEAE
    ...
    Schedonnardus, Steud.
        paniculatus (Nutt.) (S. texanus, Steud.); Nuttall, Lesquereux.
    ...

    Remarks [from Kew]: Branner & Coville, who authored the article, acknowledged Trelease for revising their list (of plant names), but did not ascribe any nomenclatural novelty to Trelease.

    Literature Cited:
    - Peterson, Paul M., Konstatin Romaschenko, and Gabriel Johnson, 2010.  

    Original Text
    The moderately supported clade of M. sect. Pseudosporobolus ( Fig. 3 ) includes a diverse assemblage of species, such as Schedonnardus paniculatus, that has panicles with long primary branches that do not rebranch, hence containing nearly sessile spikelets; …
    Taxonomy — Because our molecular analysis renders Muhlenbergia paraphyletic, we propose incorporating Aegopogon, Bealia, Blepharoneuron, Chaboissaea, Lycurus, Pereilema, Red?eldia, Schaffnerella, and Schedonnardus within Muhlenbergia. Muhlenbergia is the oldest name. Expansion of the circumscription to include these nine genera within Muhlenbergia requires the least amount of nomenclatural changes and still allows us to recognize a strongly supported monophyletic and morphologically cohesive unit.

     

    Other articles:
    • Golden Checklist Flora:  Schizachyrium scoparium;  

    Schizachyrium scoparium (Michx.) Nash “Little Bluestem”

     

    Literature Cited:
    - Michaux, Andre, 1803.  

    Michaux (1803, p. 57) described Andropogon scoparium from collections or observations in dry Carolina forests.

    Original Text
    ** Spiculis non ita villosis.
    scoparium. A. vaginis villosis : ramis ramosis , elongatis , purpurascentibus , in longam paniculam strictis : spicis simplicibus ; floribus distincte alternis , triandris ; involucellis brevissimis : flosculi additorii rudimento neutro, aristato.
    Hab. in aridis sylvarum Carolinae.

    Literature Cited:
    - Small, John Kunkel, and Per Axel Rydberg, 1913.  

    Small & Rydberg (1913, p. 59) was primarily an effort by John Small, though Per Axel Rydberg contributed the key to the orders and perhaps other parts as well. The section on Poaceae was contributed by Mr. George V. Nash.

    Original Text
    4. Schizachyrium scoparium (Michx.) Nash. Green or purplish, sometimes glaucous. Stems tufted, 4.5-15 dm. tall, the branches in l's-4's : leaf-sheaths usually glabrous, sometimes more or less hirsute ; blades 5 dm. long or less, 8 mm. wide or less, commonly roughened, and frequently somewhat hirsute above near the base : racemes single, 3-6 cm. long, the hairs on the rachis-internodes and pedicels grayish white : sessile spikelet 5-7 mm. long, usually about twice as long as the internode, the first scale generally more or less tuberculate-roughened, the awn of the fourth scale geniculate, 8-15 mm. long, closely spiral at the base, the column exserted ; pedicellate spikelet 2-4.5 mm. long, the first scale usually glabrous, tipped with an awn generally 1 mm. long or less, the pedicel considerally shorter than the sessile spikelet. [Andropogon scoparius Michx.]
    In dry sandy soil, New Brunswick to Alberta, Florida and Texas. Summer and Fall. Broom Grass.

    Note that the common name associated with this grass is “Broom Grass” rather than Little Bluestem that we use here in Colorado.

     

    Other articles:
    • Golden Checklist Flora:  Secale cereale;  

    Secale cereale L. Cereal Rye.

     

    Literature Cited:
    - Shang, Hai-Ying, Yu-Ming Wei, Xiao-Rong Wang, and You-Liang Zheng, 2006.  

    It is thought that cultivated rye originated in the Mount Ararat and Lake Van area of eastern Turkey, linguistic evidence suggesting that the introduction of cultivated rye to southern and western Europe and Central Asia were independent of each other (Sencer and Hawkes 1980). Khush (1962) concluded that cultivated rye probably entered Europe by two routes, one being through the northern Caucuses and the other through central Asia. Bushuk (1976) proposed that cultivated rye was probably distributed from south-western Asia to Russia, and thence into Poland and Germany from where it gradually spread throughout most of Europe and eventually to North America and western South America. Rye was introduced into China from Turkey and later the species was into Japan. Ma et al. (2004) found that American cultivars were more closely related to Chinese cultivars than to European cultivars and that temporal isolation had influenced the genetic diversity of rye more than geographical isolation.

    In their book concerning the origin of isolating mechanisms in flowing plants, Max et al. (1978) observed that geographical, ecological and reproductive isolation should be taken into account when studying plant evolution. In our study we analyzed the genetic similarities of cultivated rye accessions from Asia, Europe, North America and South America, but could not make any deductions regarding the domestication process of cultivated rye, indicating that further studies are needed to detect the phylogenetic relationships and evolution process of cultivated rye.


    Bushuk W (1976) Rye: Production, Chemistry, and Technology. In: Walter Bushuk (ed). Am Ass Cereal Chemists Inc, St Paul, pp 1-11.
    Khush GS (1962) Cytogenetic and evolutionary studies in Secale. II. Interrelationships of the wild species. Evolution 16:484-496.
    Ma R, Yli-Mattila T and Pulli S (2004) Phylogenetic relationships among genotypes of worldwide collection of spring and winter ryes (Secale cereale L.) determined by RAPD-PCR markers. Hereditas 140:210-221.
    Max KH, William CS and Bruce W (1978) The origin of isolating mechanism in flowing plants. In: Max KH (ed) Evolutionary Biology. Plenum Press, New York, pp 185-317.
    Sencer HA and Haekes JG (1980) On the origin of cultivated rye. Biol J Linn Soc 13:299-313.

     

    Full text requested through ResearchGate.

     

    Other articles:
    • Golden Checklist Flora:  Sorghastrum nutans;  

    Sorghastrum nutans (L.) Nash “Yellow Indian Grass”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Linnaeus (1753, v. 2, p. 1045) described Andropogon nutans from published descriptions by Gronovius (1739) and Sloane (n.d.).

    Original Text Interpreted Latin Interpreted English
    3. ANDROPOGON panicula nutante, ariftis tortuofis laevibus, glumis calycinis hirfutis.

    Andropogon folio fuperiore fpathaceo, pedunculis lateralibus oppofitis unifloris: flexuofis. Gron. virg. 133.

    Gramen avenaceum, panicula minus fparfa, glumis alba fericea lanugine obductis. Sloan. jam. 35. hift. 1. p. 43. t. 14. f. 2.

    Habitat in Virginia, Jamaica.

    nutans. 3. ANDROPOGON panicula nutante, aristis tortuosis laevibus, glumis calycinis hirsutis.

    Andropogon folio superiore spathaceo, pedunculis lateralibus oppositis unifloris: flexuosis. Gronovius, J. F., Flora virginica 1. 2. Leiden 1739. oct. 133.

    Gramen avenaceum, panicula minus sparsa, glumis alba sericea lanugine obductis. Sloane Jamaica. 35. hift. 1. p. 43. t. 14. f. 2.

    Habitat in Virginia, Jamaica.

    nutans. 3. ANDROPOGON nodding panicle, bristle twisted smooth, calyx glume hairy.

    Andropogon upper leaves spathulate, lateral peduncle opposite and single-flowered: drooping. Gronovius, J. F., Flora virginica 1. 2. Leiden 1739. oct. 133.

    Grass resembling Avena, panicle less scattered [more dense than Avena], glumes with white soft silky hairs. Sloane Jamaica. 35. hift. 1. p. 43. t. 14. f. 2.

    Habitat in Virginia, Jamaica.

    nutans.

    Jan Frederik Gronovius (also seen as Johann Frederik and Johannes Fredericus) (10 February 1690 in Leiden – 10 July 1762 in Leiden) was a Dutch botanist notable as a patron of Linnaeus. John Clayton, a plant collector in Virginia sent him many specimens, as well as manuscript descriptions, in the 1730s. Without Clayton's knowledge, Gronovius used the material in his Flora Virginica (1739–43, 2nd ed. 1762).

    Sir Hans Sloane, 1st Baronet PRS (16 April 1660 – 11 January 1753), was an Anglo-Irish physician, naturalist, and collector, with a collection of 71,000 items which he bequeathed to the British nation, thus providing the foundation of the British Museum, the British Library, and the Natural History Museum, London. He was elected to the Royal Society at the age of 24. Sloane travelled to the Caribbean in 1687 and documented his travels and findings with extensive publications years later. Sloane was a renowned medical doctor among the aristocracy, and was elected to the Royal College of Physicians at age 27. He is credited with creating drinking chocolate.

       

    Sporobolus R. Br. “dropseed” or “sacaton”

     

    Literature Cited:
    - Colbry, Vera Lyola, 1957.  

     

    Literature Cited:
    - Peterson, Paul M., Konstantin Romaschenko, Yolanda Herrera Arrieta, an Jeffrey M. Saarela, 2014.  

     

     

    Other articles:
    • Golden Checklist Flora:  Sporobolus airoides;  

    Sporobolus airoides (Torr.) Torr. “Alkali Sacaton”

     

     

    Other articles:
    • Golden Checklist Flora:  Sporobolus compositus;  

    Sporobolus compositus (Poir.) Merr. “Composite Dropseed”

     

     

    Other articles:
    • Golden Checklist Flora:  Sporobolus cryptandrus;  

    Sporobolus cryptandrus (Torr.) A. Gray. “Sand Drop-seed”

     

    Literature Cited:
    - Torrey, John, 1824b.  

    Described by John Torrey (1824b, p. 151) from a collection by Edwin James, M.D.

    Original Text
    2. Agrostis cryptandra.*
    Panicle pyramidal, with spreading subalternate branches, hairy at the axils ; flowers racemose, unarmed ; inferior valve of the calyx very short ; superior as long as the sub-equal corolla ; sheaths bearded at the throat.
    DESCRIPTION.
    Culm 3 feet high, simple, smooth, leafy, with distant joints.
    Leaves linear, 2 — 3 lines broad, smooth on both sides. Sheaths smooth, closed, longer than the joints, a little swollen with concealed abortive panicles, hairy on the margin, densely bearded at the throat. Stipule a mere bearded ring.
    Panicle large, pyramidal, bursting from the upper sheath, which partly conceals its base ; branches spreading, mostly alter- nate, nearly simple, bearded at the base, which is a little thickened. Flowers disposed in a dense racemose manner, purplish.
    Calyx two-glumed, smooth ; inferior glume scarcely one-third as long as the superior, very acute ; superior linear-lanceo- late, acute, one-nerved.
    Corolla two-valved, nearly equal, about as long as the superior glume, slightly scabrous j valves one-nerved.
    Stamens three ; anthers yellowish, exserted.
    Styles two ; stigmas white, plumose.
    Seed ovate, brown, smooth.
    Nectaries collateral, lanceolate, entire.
    Hab. On the Canadian river.

    Literature Cited:
    - Gray, Asa, 1848.  

    Gray (1848, p. 576) …

    Original Text
    3. S. cryptandrus. Leaves flat, pale (2" wide) ; the pyramidal panicle bursting from the upper sheath which usually incloses its base, its spreading branches hairy in the axils ; upper glume lanceolate, acute, twice the length of the lower one, as long as the nearly equal paleae ; sheaths strongly bearded at the throat. ♃. (Agr. and Vilfa cryptandra, Torr.) — Sandy soil, Buffalo, New York, and westward. Ipswich, Massachusetts, Oakes. Aug. — Culm 2° -3° high. Panicle lead-color : spikelets small.

    Literature Cited:
    - Torok, P., et al., 2021.  

    Torok, et al. 2021, reviewed the current distribution and characteristics of Sporobolus cryptandrus (sand dropseed), an invasive C4 grass species of North American origin recently discovered in Hungary and provided information on (i) its current distribution paying special attention to its invasion in Eurasia; (ii) the characteristics of the invaded habitats in Central Europe; (iii) seed bank formation and germination characteristics, crucial factors in early establishment; and (iv) the effects of its increasing cover on vegetation composition (from the Abstract).

    Literature Cited:
    - Hábenczyus, Alida A., et al., 2022.  

    Habenczyus, et al., 2022 evaluated stands of the invasive grass, Sporobolus cryptandrus, in its native North American range and its non-native European range, and found that the spread of Sporobolus, away from its native range, leads to the impoverishment of host communities and compromises the biomass and floral resource-provisioning capacity of the vegetation to higher trophic levels.

     

    Other articles:
    • Golden Checklist Flora:  Sporobolus heterolepis;  

    Sporobolus heterolepis (A. Gray) A. Gray. “Prairie Drop-seed”

     

    Literature Cited:
    - Gray, Asa, 1835.  

    Gray (1835, p. 233) published Vilfa heterolepis from collections in New York state, in addition to collections from Connecticut, Delaware, and Montreal, Canada.

    34. VILFA HETEROLEPIS, (sp.nov.) ; foliis setaceis ; panicula pyramidata, sparsiflora ; gluma inferiore subuliformi, superiore ovata, cuspidata, subduplo breviori ; valvulis perianthio subaequaUbus, muticis, gluma extima paulo minori.
    Root perennial. Culm 1 — 2 feet in height, smooth. Leaves convolute-setaceous, with the margins hispidly scabrous upward ; the lower ones equalling the culm; the upper ones much shorter. Lower sheaths pilose ; upper ones smooth. Panicle pyramidal, spreading or subcontracted; branches solitary, nearly simple, few and loosely flowered. Glumes purplish ; the outer one reduced to a subula, about one half the length of the inner one, which is strikingly membranaceous in texture, ovate or ovate- oblong, one-nerved, with the nerve produced into a short cusp. Valves of the perianth oblong-lanceolate, rather obtuse, thin and membranaceous, a little shorter than the superior glume. Inferior valve, obscurely one- nerved, slightly apiculate. Superior valve two-nerved, a little shorter than the outer one. Stamens 3. Anthers large, linear, orange-red. Stigmas 2, hairy. Styles very short. Caryopsis subglobose, coriaceous, smooth and shining.
    Hab. On rocks, Watertown, Jefferson county, Dr. Crawe. Flowers Aug. — Sept. I have also specimens collected near New Haven, Connecticut, by Mr. J. D. Dana. In Muhlenberg's herbarium there is a fragment of this grass with a specimen of V. juncea, from the late Dr. Baldwin. The locality is not noted on the label, but it was most probably collected in Delaware. Dr. Torrey has also received specimens from the vicinity of Montreal.

    Literature Cited:
    - Gray, Asa, 1848.  

    Gray (1848b, p. 576) published Sporobolus heterolepis placing his previous Vilfa heterolepis in R. Brown's genus of Sporobolus.

    Original Text Comments
    2. S. heterolepis. Leaves involute-thread-form, rigid, the lowest as long as the culm (1°-2°), which is naked above; panicle very loose; glumes very unequal ; the lower awl-shaped (or bristle-pointed from a broad base) and somewhat shorter, the upper ovate-obling and taper-pointed and longer, than the equal paleæ. ♃ (Vilfa heterolepis, Gray.) — Dry soil, New Haven, Connecticut; Watertown, New York; and Columbus, Ohio. Aug. — Plant exhaling an unpleasant scent (Sullivant), stouter than the last [S. junceus], the spikelets thrise larger. Utricle spherical (1" in diameter), shining, thick and coriaceous! The stated dimensions of the seed has got to be some sort of mistake.

       

    Thinopyrum Love

     

    Literature Cited:
    - Barkworth, Mary E., and Douglas R. Dewey, 1985.

    Other articles:
    • Golden Checklist Flora:  Notes on Thinopyrum intermedium, Barkworth & D. R. Dewey, 1985;  

    Barkworth & Dewey (1985) ...

    Original Text
    7. Thinopyrum A. Love, Taxon 29: 351, 1980, Fig 6.
    As recognized here, Thinopyrum comprises approximately 17 species in Eurasia, of which three have become established in North America. Their removal from Elytrigia is supported by both karyotypic and genomic analyses (Cauderon, 1966; Heneen and Runemark, 1972a,b; Lyubimova, 1970). Love (1984) places the caespitose species, such as T. ponticum, in the genus Lophopyrum but Cauderon and Saigne (1961), Cauderon (1966) and Dvorak (1981) have shown that the genomes of such species are essentially the same as those found in Thinopyrum sensu stricto. We have previously included Thinopyrum in Elytrigia (Barkworth, Dewey, and Atkins, 1983; Dewey, 1982, 1983a, b), but with considerable misgivings.

     

    Other articles:
    • Golden Checklist Flora:  Thinopyrum intermedium;  

    Thinopyrum intermedium (Host) Barkworth & D. R. Dewey “Intermediate Wheatgrass”

     

    Literature Cited:
    - Barkworth, Mary E., and Douglas R. Dewey, 1985.

    Other articles:
    • Golden Checklist Flora:  Notes on Thinopyrum, Barkworth & D. R. Dewey, 1985;  

    Barkworth & Dewey (1985) ...

    Original Text
    Thinopyrum intermedium (Host) Barkw. & D. R. Dewey, comb. nov. Basionym: Triticum intermedium Host, Gram. Austr. 3:23, 1805.

     

    Other articles:
    • Golden Checklist Flora:  Vulpia octoflora;  

    Vulpia octoflora (Walt.) Rydb. “Six Weeks Fescue”

     

    Literature Cited:
    - Walter, Thomas, 1788.  

    Walter (1788, p. 81) published Festuca octoflora

    Original Text
    45. FESTUCA. Cal. 2-valvis. Spicula oblinga teretiufcula. Glumis acuminatis.
    octoflora 2. panicula erecta, fpiculus octofloris acuminatis.

    Literature Cited:
    - Gmelin, Karl Christian, 1805.  

    The genus Vulpia was proposed by Gmelin (1805) for plants common in the German state of Baden (now part of Baden-Wurttemberg), Alsace and on both sides of the Rhine.

    Original Text Translation and Comments
    Hab. Utrinque paffim copiofe in agris fabulofis, arvis et collibus apricis; prope Carlsruhe in arvis et ad vias fabulofas paffim abunde. Habitat. Abundant everywhere in the sunny fields of both sides [of the Rhine River] to the hills; and abundant near Carlsruhe.

    Literature Cited:
    - Rydberg, Per Axel, 1909b.  

    Rydberg (1909) published Vulpia octoflora (Walt.) Rydb. without comment or explanation.

    Literature Cited:
    - Lunell, J., 1915.  

    Lunell (1915, v. 4, n. 6, p. 235) ...

    227. Carex brevior (Dewey) Mackenzie, in ed.
    Carex stramina var. brevior Dewey, Am. Journ. Sci. II: 158. (1826).

     

    Other articles:
    • Golden Checklist Flora:  Carex brevior;  

    Carex brevior (Dewey) Mack. “Shortbeak Sedge”

     

     

    Other articles:
    • Golden Checklist Flora:  Carex deweyana;  

    Carex deweyana Schwein. “Dewey sedge”

     

     

    Other articles:
    • Golden Checklist Flora:  Carex douglasii;  

    Carex douglasii Boott “Douglas Sedge”

     

       

    Carex duriuscula C.A. Mey. “Needleleaf Sedge”

    (Syn: Carex stenophylla Wahlenb., Carex stenophylla Wahlenb. ssp. eleocharis (L. H. Bailey) Hultén)

     

    Other articles:
    • Golden Checklist Flora:  Carex inops heliophila;  

    Carex inops L.H. Bailey ssp. heliophila (Mack.) Crins “Sun Sedge”

     

    Literature Cited:
    - Bailey, L. H., 1886.  

    Carex inops was published by L. H. Bailey (1886) from a collection by L. F. Henderson on sandy grounds on subalpine slopes of Mt. Hood, Oregon, July, 1884. Described as being much like C. pennsylvanica and lacking the ribbed and hardened character of the perigynia of that species and its allies, and the beak is straighter and more deeply cleft.

    There is one voucher record online of an isotype (ORE17017) collected by L. F. Henderson, 24 July 1884, with a locality of “[Clackamas or Hood River], Mt. Hood, alpine.” No image available.

    There is another voucher at Harvard (GH27273) labeled “Type” but dated 1885. And possibly a third undated specimen (NY11164). These appear to be substantially larger than the subspecies we see here in Colorado.

    .

    Literature Cited:
    - Mackenzie, Kenneth K., 1913.  

    Carex heliophila was described by Mackenzie (1913) from a specimen he collected on the open prairie, near Lee's Summit, Jackson County, Missouri, on May 9, 1897. The voucher is now at NY (NY11144).

     

    Crins (1983) reduced C. heliophila to a subspecies of C. inops.

    Carex inops subsp. heliophila (Mack.) Crins, Canad. J. Bot. 61: 1709 (1983).

    Crins, W. J., and P. W. Ball. 1983. The taxonomy of the Carex pensylvanica complex (Cyperaceae) in North America. Canadian Journal of Botany 61(6): 1692-1717. (Carex pensylvanica)

    The Carex pensylvanica complex consists of four North American taxa. Morphological variation patterns within the complex were examined using principal-components analysis and discriminant-functions analysis. These results indicate that two eastern species, C. lucorum Willdenow ex Link, and C. pensylvanica Lamarck, and one western species, C. inops Bailey, should be recognized. The latter species comprises two subspecies, C. inops subsp. inops and C. inops subsp. heliophila (Mackenzie) Crins, comb. nov. Cytological and geographical evidence lend support to this classification. A key and distribution maps for the taxa are provided.

    I have no access to this article. It's $30.00.

     

    Other articles:
    • Golden Checklist Flora:  Carex nebrascensis;  

    Carex nebrascensis Dewey “Nebraska Sedge”

     

     

    Other articles:
    • Golden Checklist Flora:  Carex occidentalis;  

    Carex occidentalis L. H. Bailey “Western Sedge”

     

    Literature Cited:
    - Bailey, L. H., 1889.  

    L. H. Bailey (1889, v. 1, p. 14) ...

    18.—Carex occidentalis.
    C. muricata, Olney, Bot. King's Rep. 362, in part (1871); W. Boott, Bot. Wheeler's Surv. 277 (1878); Bailey, Coulter's Man. 390 (1885).
    C. muricata L., var. Americana, Bailey, Proc. Amer. Acad. Arts and Sci. xxii. 140 (1886).
    Glaucous; leaves narrower than in the last, and relatively longer; spikes more or less scattered into a very slender head an inch or so long, the lowest one or two usually wholly distinct; bracts scale-like, inconspicuous; perigynum larger than in the last, turgid-ovate, abruptly short-beaked, nearly marginless and often smooth; scales muticous. (239c.) Mountains, Montana to Arizona.

     

    Other articles:
    • Golden Checklist Flora:  Carex praegracilis;  

    Carex praegracilis W. Boott “Clustered Field-Sedge”

     

     

    Other articles:
    • Golden Checklist Flora:  Carex stipata;  

    Carex stipata Muhl. ex Willd. “Saw Beaked Sedge”

     

     

    Other articles:
    • Golden Checklist Flora:  Cyperus squarrosus;  

    Cyperus squarrosus L. “Bearded Flatsedge”

     

     

    Other articles:
    • Golden Checklist Flora:  Eleocharis acicularis;  

    Eleocharis acicularis (L.) Roem. & Schult. “Needle Spikerush”

     

     

    Other articles:
    • Golden Checklist Flora:  Eleocharis compressa;  

    Eleocharis compressa Sull. “Flatstem Spikerush”

    (Syn: Eleocharis elliptica Kunth var. compressa (Sull.) Drapalik & Mohlenbr.)

     

    Other articles:
    • Golden Checklist Flora:  Eleocharis macrostachya;  

    Eleocharis macrostachya Britton “Common Spike-Rush”

     

     

    Other articles:
    • Golden Checklist Flora:  Eleocharis palustris;  

    Eleocharis palustris (L.) Roem. & Schult. “Common Spikerush”

     

       

    Eleocharis parvula (Roem. & Schult.) Link ex Bluff, Nees & Schauer “Dwarf Spikerush”

    (Syn: Eleocharis coloradoensis (Britton) Gilly)

     

    Other articles:
    • Golden Checklist Flora:  Schoenoplectus americanus;  

    Schoenoplectus americanus (Pers.) Volkart ex Schinz & R. Keller “Chairmaker's Bullrush”

     

     

    Other articles:
    • Golden Checklist Flora:  Schoenoplectus lacustris;  

    Schoenoplectus lacustris (L.) Palla “Lakeshore Bulrush”

    (Syn: Scirpus lacustris L. )

     

    Other articles:
    • Golden Checklist Flora:  Schoenoplectus pungens;  

    Schoenoplectus pungens (Vahl) Palla “Threesquare”

     

     

    Other articles:
    • Golden Checklist Flora:  Schoenoplectus tabernaemontani;  

    Schoenoplectus tabernaemontani (C.C.Gmel.) Palla “Softstem Bulrush”

    (Syn: Scirpus lacustris L. var. validus (Vahl) Kük., Scirpus validus Vahl)

     

    Other articles:
    • Golden Checklist Flora:  Scirpus pallidus;  

    Scirpus pallidus (Britt.) Fern. “Cloaked Bulrush”

     

    Literature Cited:
    - Britton, Nathaniel Lord, 1889.  

    Britton (1889, v. 9, p. 14) ...

    Scirpus atrovirens, Muhl. var. pallidus, n. var.
    Whole plant pale, including the inflorescence, which is composed of larger heads than in the type, and is more contracted; glumes more squarrose. A well-marked variety, but I am so far unable to detect further differences. The achenium is exactly like that of the eastern plant. Indian Territory (E. Palmer, No. 358, 1868); Mitchell Co., Kansas (M. A. Carleton, 1886); Hitchcock Co., Neb. (H. J. Webber, 1888).

    Literature Cited:
    - Fernald, M. L., 1906.  

    Fernald (1906, v. 8, p. 163) ...

    S. pallidus (Britton), n. comb. Similar: leaves very pale: spikelets pale brown, very numerous in irregular glomerules: scales elliptic=ovate, 2 to 3 mm. long with conspicuous pale midrib prolonged into lond setulose awns, about twice as long as the achenes. — S. atrovirens, var. pallidus Britton, Trans. N. Y. Acad. sci. ix. 14 (1889) — Manitoba to Kansas and the Rocky Mountain.

     

    Other articles:
    • Golden Checklist Flora:  Lemna minor;  

    Lemna minor L. “Common Duckweed”

     

     

    Other articles:
    • Golden Checklist Flora:  Tradescantia occidentalis;  

    Tradescantia occidentalis (Britton) Smyth “Prairie Spiderwort”

     

     

    Other articles:
    • Golden Checklist Flora:  Juncus arcticus;  

    Juncus arcticus (Willd.)Trautv. “Arctic Rush”

     

     

    Other articles:
    • Golden Checklist Flora:  Juncus arcticus var. balticus;  

    Juncus arcticus (Willd.) Trautv. var. balticus (Willd.) Trautv. “Arctic Rush”

    (Syn: Juncus arcticus (Willd.) Trautv. ssp. ater (Rydb.) Hultén, J. ater Rydb., J. balticus Willd., J. balticus Willd. ssp. ater (Rydb.) Snogerup, J. balticus Willd. var. montanus Engelm.)

     

    Other articles:
    • Golden Checklist Flora:  Juncus articulatus;  

    Juncus articulatus L. “Joint-Leaf Rush”

     

     

    Other articles:
    • Golden Checklist Flora:  Juncus bufonius;  

    Juncus bufonius L. “Toad Rush”

     

     

    Other articles:
    • Golden Checklist Flora:  Juncus compressus;  

    Juncus compressus Jacq. “Roundfruit Rush”

     

     

    Other articles:
    • Golden Checklist Flora:  Juncus interior;  

    Juncus interior Wiegand “Inland Rush”

     

    Literature Cited:
    - Wiegand, K. M., 1900.

    Locations: Encampment River.  

    Wiegand (1900, v. 27, p. 516-517) ...

    Original Text Comments
    5. Juncus interior sp. nov.  
    Tall and rather stout (5-10 dm. high), light green : stem erect, nearly terete, coarsely grooved : leaves several, about one third length of culm, blades narrow (1-1.25 mm. wide), flat or involute : sheaths nearly all blade-bearing, large and loose, margin and short rounded auricles membranous, often slightly yellowish : inflorescence large and open, very many flowered, 3-10 cm. long, branches conspicuously ascending : flowers distant and scattered, not secund, pale stramineous : bracts commonly two, exceeding the inflorescence : bracteoles acuminate : perianth 3-4 mm. long, its parts nearly equal, broadly subulate, very acute, scarious margin narrow, extending on the inner to the tip, all appressed or erect : stamens one half the length of the perianth : anthers short-oblong, much shorter than the filaments : styles very short : capsule oblong or rarely ovate-oblong, obtuse, barely apiculate, equaling the perianth ; placentae not quite meeting at the axis except at the ends : seeds oblong, rather small (.35-.50 × .14-.17 mm.), apiculate at both ends, shallowly areolate.  
    Illinois to Wyoming, in dry woods and prairies.  
    Specimes (sic) examined :  
    Illinois: Richmond (Vasey in Gray Herb.; type), between Urbana and Centralia (Vasey), Athens (E. Hall, — June. Bor. Am. Engelm. no. 21); Illinois Dr. Mead, no. 23.  
    Wyoming : Grand Enchantment Creek (A. Nelson, no. 3982, 1897). The correct name is “Grand Encampment Creek” as this can be clearly read on the image of the voucher. The current name is the Encampment River.
    Juncus interior has previously constituted the main part of the so-called J. secundus of the Mississippi valley. Closer observation shows however that it is quite distinct as well in structure as in area of distribution. J. secundus is always a slender plant with rather close sheaths, while the present species is commonly tall and stout with much broader sheaths. The two species also differ in the larger inflorescence of the latter, with larger flowers which are not secund, shorter anthers and more oblong capsule. The bracts also much exceed the inflorescence, while in J. secundus they usually do not.  

     

    Other articles:
    • Golden Checklist Flora:  Juncus torreyi;  

    Juncus torreyi Coville “Torrey's Rush”

     

    Literature Cited:
    - Coville, Frederick Vernon, 1895.  

    Coville (1895, v. 22, p. 303) ...

    * After examing a large number of herbarium specimens and observing both plants in the field for several years, I am convinced that Torrey's Juncus nodosus megacephalus does not intergrade with J. nodosus proper. In view of the earlier Juncus megacephalus of Curtis, Torrey's plant is naned as follows:
    Juncus Torreyi Juncus nodosus var. megacephalus Torr. Fl. N. Y. 2: 326. 1843.
    Juncus megacephalus Wood, Classbook Bot. Ed. 2: 724. 1861. Not Curtis.

     

    Other articles:
    • Golden Checklist Flora:  Yucca glauca;  

    Yucca glauca Nutt. “Soapweed Yucca”

     

    Literature Cited:
    - Nuttall, Thomas, 1813.  

    (Nuttall, 1813, no. 89) ...

    Original Text Comments
    89 *Yucca glauca. ‡ Leaves narrow, and filiferous ; capsules dry, coriaceous, and large as that of Y. gloriosa. — Flower not seen. — Used in N. Mexico as a substitute for hemp. Collected 1600 miles up the Missourie, about lat. 49°. Yucca gloriosa L. is an accepted name for a yucca in the southeast United States.

     

    Other articles:
    • Golden Checklist Flora:  Calochortus gunnisonii;  Watson, 1871, 40th Parallel, publication details;  

    Calochortus gunnisonii S. Watson “Gunnison's Mariposa Lily”

     

    Literature Cited:
    - Torrey, John, and Asa Gray, 1855.

    Other articles:
    • Golden Checklist Flora:  Torrey & Gray, 1855. Pac. R. R. Rep.;  

    Torrey & Gray (1855) in their report on the Botany of Gunnison's expedition, note a Calochortus that they referred to an unknown variety of C. venustus.

    Calochortus venustus, Benth. in Hort. Trans, (n. s.) 1, p. 412, t. 15,/. 2, var ?: sepals erect; petals obovate, bearded and without a spot below the middle, purple at the base. Grows under trees on high mountains. Utah. Stem 2-3-flowered. Leaves grass-like, about two lines wide. Flowers nearly 3 inches in diameter. Sepals lanceolate, striate with purple veins externally. Petals nearly twice as long as the sepals, the upper half white, pale yellow-ish green lower down, where the inside is bearded with longish gland-tipped hairs, which are dark purple at the base. Near the base the hairs are more numerous, and form a transverse tuft; at the very bottom the claw is dark purple. Differs from G. venustus in its much narrower and less bearded petals, and in wanting the red spot above the middle.

    Literature Cited:
    - Watson, Sereno, 1871.

    Other articles:
    • Field Notes:  Coll. No. 2901, 21 Jul 2022;  

    Watson (1871, p. 348) ...

    Calochortus Gunnisoni. (C. venustus, Var. ?, Torr. Bot. Gunnison's Rep., Pac. R. R. Surv. 2. 130.) Habit as in the last ; petals rounded at the apex, white above, yellowish-green below the middle and dark-purple at base, strongly bearded with longish gland-tipped hairs, which are also dark-purple at base, the densely hairy gland transverse and occupying nearly the whole width of the petal; anthers 5" long, rather exceeding the somewhat dilated filaments, oblong-lanceolate, subcordate at base, narrowed above into an awn-like termination or acute ; immature capsule narrowly oblong, attenuate above. — Rocky Mountains of Colorado. Collected also in Utah by Gunnison.

     

    Other articles:
    • Golden Checklist Flora:  Smilax lasioneuron;  

    Smilax lasioneuron Hook. “Blue Ridge Carrionflower”

     

       

    Melanthiaceae

     

    Literature Cited:
    - Angiosperm Phylogeny Group, 2003.  

    The Angiosperm Phylogeny Group II (2003) placed Melanthiaceae at the rank of Family within Liliales.

    Literature Cited:
    - Zomlefer, Wendy B., Walter S. Judd, W. Mark Whitten, Norris H. Williams, 2006.  

    Zomlefer, et al., 2006 published a synopsis of Melanthiaceae. They also recognize Toxicoscordion.

       

    Zigadenus Michx.

     

    Literature Cited:
    - Flora of North America Editorial Committee, eds., 2013.  

    FNANM (Vol. 26, 2003) retains Zigadenus in Liliaceae and places Toxicoscordion in synonomy.

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittmann (2012) write ...

    Original Text Comments
    ... It is interesting to note that, even without any knowledge of the modern evidence for segregation of [Melanthiaceae] and other lily-like families, Rydberg presented essentially the same arrangement in his book on the Rocky Mountain flora, but the taxonomic establishment did not follow him.
    They go on to recognize one species of Toxicoscordion in Colorado, T. venenosum and place T. gramineum and T. paniculatum in synonomy.

    Any Toxicoscordion collected in Colorado and identified with Weber & Wittman (2012) will be T. venenosum. This is somewhat the same issue as with Eriogonum arcuatum and E. flavum.

     

    Other articles:
    • Golden Checklist Flora:  Toxicoscordion paniculatum;  

    Toxicoscordion paniculatum Rydb. “Foothill Death Camas”

     

    Literature Cited:
    - Nuttall, Thomas, 1834a.  

    Nuttall (1834a) ...

    Original Text Comments
    108. Helonias *paniculata. Monoica, scapo inferne folioso paniculato, superne infra apicem fructifero, bracteis membranaceis acuminatis, petalis lanceolato-oblongis, staminibus exsertis, antheris aureis.  
    ♃ Root bulbous? Leaves lanceolate-linear and acute. The scape shortly branched below, the branches subtended by short membrabaceous leaves. The flowers yellowish-white; below the summit of the panicle or raceme arise the fertile flowers upon long peduncles. Styles three, contiguous. Stamens exserted, the anthers bright yellow.  
    Hab. In the Kamas prairie, near Flat-Head river. Flowering in June.  

    Literature Cited:
    - Watson, Sereno, 1871.  

    Watson (1871, p. 343) ...

    Original Text Comments
    Zigadenus paniculatus. (Helonias, Nutt. Amianthium Nuttallii, Gray, Var. β., l. c.) Stem rather stout, 15-30' high, somewhat leafy; leaves sheathing at base, (even the 1-2 uppermost bract-like ones,) broad-linear, 4-8" wide, rough-margined and roughish-puberulent on both sides, folded-carinate and mostly falcate ; eaceme panicled, many-flowered, the lower branches spreading, short, the terminal ones elongated, (3-10';) flowers on slender pedicels, often sterile and short-pedicelled in the lower racemes; bracts membranous, sepals 1-2" long, oblong, abruptly somewhat narrowed at the base, the rather broad claw green and glandular ; ovary-cells about 10-ovuled ; capsule oblong-ovate or oblong, ½-1' long ; seeds 3-5" long, oblong. — an examination of numerous specimens leaves no doubt of the distinctness of these two species. Bulb as in the last, but usually larger, the whole plant stout, growing on dry foothills, and in flower a month earlier. The root of neither species is eaten by the Indians. Oregon and Washington Territory. Frequent on the foot-hills of the Virginia, Trinity, and West Humboldt Mountains, Nevada, and in the Wahsatch ; 5-6,000 feet altitude ; May, June. (1,164.) The “last” is Zigadenus nuttallii A. Gray.

    Literature Cited:
    - Watson, Sereno, 1879.  

    Original Text Comments
    6. Z. paniculatus, Watson. Very similar : usually stout: leaves 3 to 8 lines broad, usually all sheathing : raceme compound : perianth-segments deltoid, acute or acuminate, with a short claw ; gland less deifinitely margined, often reaching nearly to the middle of the blade : fruiting pedicels spreading : capsule ½ to 1 inch : seeds 3 to 5 lines long. — King's Rep. 5. 344. Amianthum Nuttallii, var. β, Gray, Rev. Melanth. 121. Helonias paniculata, Nutt. Journ. Philad. Acad. 7. 57. California (east slope of Sierra Nevada) and Idaho to Utah and the Sackatchewan (sic).

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    In the key to Zigadenus paniculatus, Ackerfield (2015) writes:

    Original Text Comments
    ... Zigadenus paniculatus var. venenosus (S. Watson) Ackerfield occurs to the west of Colorado ...
    Ackerfield (2015) then recognizes Z. paniculatus var. paniculatus and var. gramineus.

       

    Toxicoscordion venenosum Watson “Meadow Death Camas”

    Original Text Comments
    5. Z. venenosus. Stem slender, ½ to 2 feet high : leaves rarely over 2 or 3 lines broad, scabrous, the cauline not sheathing : raceme simple, short, with narrow scarious bracts : perianth free from the ovary ; segments triangular-ovate to elliptical, obtuse or rarely acutish, 2 or 3 lines long, all abruptly contracted to a short glandular claw, the blade rounded or subcordate at base ; gland extending slightly above the claw with a well-defined irregular margin : stamens somewhat adnate to the claw : pedicels suberect in fruit : capsule 4 to 6 lines long : seeds 1½ to 2½ lines long. — California (Monterey and Mariposa Counties) to British Columbia and east to Utah and Idaho. Bulb poisonous. The Coast Range form is usually stouter, with a larger occasionally compound raceme. Hitherto referred to the last species. The “last species” is Z. paniculatus.

    Literature Cited:
    - Watson, Sereno, 1879.
    Full Size ImageInflorescence of Coll. No. 2055, Toxicoscordion paniculatum  

    Original Text Comments
    5. Z. venenosus. Stem slender, ½ to 2 feet high : leaves rarely over 2 or 3 lines broad, scabrous, the cauline not sheathing : raceme simple, short, with narrow scarious bracts : perianth free from the ovary ; segments triangular-ovate to elliptical, obtuse or rarely acutish, 2 or 3 lines long, all abruptly contracted to a short glandular claw, the blade rounded or subcordate at base ; gland extending slightly above the claw with a well-defined irregular margin : stamens somewhat adnate to the claw : pedicels suberect in fruit : capsule 4 to 6 lines long : seeds 1½ to 2½ lines long. — California (Monterey and Mariposa Counties) to British Columbia and east to Utah and Idaho. Bulb poisonous. The Coast Range form is usually stouter, with a larger occasionally compound raceme. Hitherto referred to the last species. The “last species” is Z. paniculatus.

     

    Other articles:
    • Golden Checklist Flora:  Maianthemum stellatum;  

    Maianthemum stellatum (L.) Link. “Starry False Lily of the Valley”

    (Syn: Smilacina stellata (L.) Desf.)

     

    Other articles:
    • Golden Checklist Flora:  Maianthemum racemosum amplexicaule;  

    Maianthemum racemosum (L.) Link ssp. amplexicaule (Nutt.)LaFrankie “Feathery False Lily of the Valley”

     

    Literature Cited:
    - Linne´, Carl von, 1753.  

    Original Text
    ★ * Smilaces T. corollis rotatis.
    5. CONVALLARIA foliis feffilibus, racemo terminali compofito.
    Convallaria racemo conpofito. Roy. lugdb. 26.
    Convallaria foliis alternis, racemo terminale. Hort. cliff. 125. Gron. virg. 38.
    Polygonatum racemofum. Corn. canad. 36. t. 37.
    Polygonatum ramofum & racemofum fpicatum. Morif. hift. 3. p. 537. f. 13. t. 4. f. 9.
    Polygonatum racemofum americanum, ellebori albi foliis ampliffimis. Pluk. alm. 301. t. 311. f. 2.
    Habitat in Virginia, Canada. ♄
    racemofa.
     

    Maianthemum F.H.Wigg., Prim. Fl. Holsat. 14 (1780).

    Literature Cited:
    - Desfontaines, Rene Louiche, 1780.  

    Smilacina Desf. was published by Rene L. Desfontaines as a segregate from Convallaria

    Literature Cited:
    - Nuttall, Thomas, 1834a.

    Other articles:
    • Golden Checklist Flora:  Nuttall, 1834a, publication details;
    • Glossary:  amplexicaul;  

    Nuttall (1834a, p. 58) described Smilacina amplexicaulis from a collection by Nathaniel Wyeth in 1833.

    Original Text
    110. Smilacina (B.) *amplexicaulis. Foliis cordato-ovatis amplexicaulibis subacuminatis, floribus paniculatis, petalis staminibus paulo brevioribus.
    ♃. Stem about a foot high, and as well as the leaves covered with a short pubescence. Leaves three to three and a half inches long, by an inch and a half wide, broad ovate, shortly acuminated, and amplexicaule. Panicle about four inches long, very similar to that of S. racemosa. The flowers are, however, as well as the petals, which are but little shorter than the stamens, and white. Style undivided, very short.
    Hab. In the valleys of the Rocky Mountains, about the sources of the Columbia river. Flowering about the middle of June.
     

    Watson (1871, p. 345) placed S. amplexicaulis as a variety of Smilacina racemosa

    Original Text
    Smilacina racemosa, Desf., Var. amplexicaulis. (S. amplexicaulis Nutt. Jour. Acad. Phil. 7. 58.) Leaves closely sessile and amplexicaul, shortly acuminate or only acute ; style at lease half as long as the ovary and equaling the stamens. — Distiguished at once from the usual form by its less acuminate sessile leaves and longer style and filaments. It is 2012 Hartweg and 353 Bridges from California, and 845 Fendler from New Mexico ; collected also by Wyeth and Lyall in Oregon and Washington Territory and by Bourgeau in the Rocky Mountains. 728 Coulter from California, Lyall's specimens from Lower Fraser River and Bourgeau's from the Winnipeg are the typical form, which extends southward to Southern California, New Mexico, and upper districts of the Gulf States. Clover Mountains, Nevada, and the Wahsatch ; 6-7,000 feet altitude ; May-september. (1,168)

    Literature Cited:
    - Harrington, H. D., 1954.  

    Harrington (1964, 2nd ed.) says this about Smilacina racemosa (L.) Desf.:

    Original Text
    Several varieties have been proposed for our western plants S. racemosa var. amplexicaulis (Nutt.) Wats. is the most distinct. --- Woods and thickets. Throughout most of temperate North America. Our records scattered in the western two-thirds of the state at 6500-10,000 feet.

    Literature Cited:
    - LaFrankie, James V., 1986b.  

    LaFrankie (1986) published a morphology and taxonomy of Maianthemum, placing M. amplexicaule as a subspecies of M. racemosum. LaFrankie also wrote the treatment for FNANM.

    Literature Cited:
    - LaFrankie, James V., 1986b.  

    LaFrankie, J. V., Jr. 1986. Transfer of the species of Smilacina Desf. to Maianthemum Wigg. (Liliaceae). Taxon. 35: 584-589. Unfortunately I don't have access to Taxon through JSTOR.

     

    Maianthemum racemosum var. amplexicaule (Nutt.) Dorn, Vasc. Pl. Wyoming 298 (1988): (1988).

    Literature Cited:
    - LaFrankie, James V., 2003.  

    LaFrankie (2003) writing in Flora of North America treats M. amplexicaule as a subspecies of M. racemosum.

    Literature Cited:
    - Weber, William A., and Ronald C. Wittmann, 2012.  

    Weber & Wittman (2012) treat M. amplexicaule (Nutt.) W. A. Weber as the species rank, noting “… FNA considers this to be a variety of M. racemosum; however, our species is diploid, while M. racemosum is tetraploid. Contrary to claims, these do not intergrade.”

    Literature Cited:
    - Ackerfield, Jennifer, 2015.  

    Ackerfield (2015) accepts LaFrankie's (2003) placement of M. amplexicaule as a subspecies of M. racemosum.

    Literature Cited:
    - Kim, Changkyun, and Joo-Hwan Kim, 2017.  

    Kim and Kim (2017) did not focus on M. racemosum, though their diagrams suggest the taxon split from the remainder of the genus s.s. early in the evolution of the genus.

     

    Other articles:
    • Golden Checklist Flora:  Allium cernuum;  

    Allium cernuum Roth “Nodding Onion”

     

     

    Other articles:
    • Golden Checklist Flora:  Allium × proliferum;  

    Allium × proliferum (Moench) Schrad. ex Willd. “Garden Onion”

     

    Literature Cited:
    - Willdenow, Carl Ludwig, 1809.  

    Willdenow (1809, p. 358) ...

    * 12. ALLIUM proliferum.
    A. caule nudo fistuloso tortuoso, umbella bulbifera prolifera, staminibus tricuspidatis.
    Allium proliferum. Schrad. cat. hort. goett.
    Habitat . . . . . . . . ♃ D.
    Spatha bivalvis, valvulis ovatis concavis. Umbella universalis bulbis tribus magnis ramisque longis tribus seu quatuor composita, quorum longissimus saepe bipedalis atque iterum tribus bulbis terminatus cum floribus tribus vel quatuor ; ex hac umbella saepe alia e bulbis tribus composita ramo semipedali suffulta elongatur. Rami laterales umbellae universalis tripollicares usque semipedales, apice bulbiferi cum floribus nonnullis. Corolla alba, nervo viridi ornata. Stamina corolla longiora, alterna obtuse trifida. Antherae virides.

     

    Other articles:
    • Golden Checklist Flora:  Allium textile;  

    Allium textile A. Nelson & J.F. Macbr. “Textile Onion”

     

    Literature Cited:
    - Nelson, Aven, and J. Francis Macbride, 1913.  

    Nelson & Macbride (1913, v. 56, p. 470) ...

    Original Text
    Allium textile, n. n. — A. reticulatum Fraser in Mem. Wern. Soc. 6:36. 1827; not A. reticulatum J. and C. Presl. Fl. Cech. 73. 1819.
    Our collections made in 1912 show that this species has a wider distribution than heretofore assigned to it. Specimens having been secured on the Snake River, at Shoshone Falls, it seem probable that it may extend quite into eastern Oregon.

     

    Other articles:
    • Golden Checklist Flora:  Asparagus officinalis;  

    Asparagus officinalis L. “Asparagus”

     

     

    Other articles:
    • Golden Checklist Flora:  Lilium philadelphicum;  

    Lilium philadelphicum L. “Wood Lily”

     

     

    Other articles:
    • Golden Checklist Flora:  Muscari botryoides;  

    Muscari botryoides (L.) Mill. “Common Grape Hyacinth”

     

     

    Other articles:
    • Golden Checklist Flora:  Prosartes trachycarpa;  

    Prosartes trachycarpa S. Watson “Roughfruit Fairybells”

     

     

    Other articles:
    • Golden Checklist Flora:  Streptopus amplexifolius;  

    Streptopus amplexifolius (L.) DC. “Clasping Twitedstalk”

     

    Literature Cited:
    - Lamarck, Jean-Baptiste, and Agustin de Candolle, 1815.  

    Lamarck and DeCandolle (1815, t. 3, p. 174)

    Original Text Interpreted Text
    CCXI. STREPTOPE. STREPTOPUS. CCXI. STREPTOPE. STREPTOPUS.
    Streptopus. Michaux. — Uvularia. Hall. — Uvulariae sp. Linn. Streptopus. Michaux. — Uvularia. Hall. — Uvulariae sp. Linn.
    Car. Le périgone est divisé profondément en six lanières munies à la base interne d’une cavité nectarifère ; les anthères sont plus longues que les filamens ; le fruit est une baie lisse à enveloppe mince. Car. The perigone is deeply divided into six strips provided at the internal base with a nectariferous cavity; the anthers are longer than the filaments; the fruit is a smooth berry with thin envelope.
    Obs. Ce genre , confondu par Linné avec l’uvulaire , mais don’t les anciens botarnstes connoissoient bien l’organisation , n’appartient pas même à la famille des liliacées , don’t l’uvulaire fait partie; il en diffère par son fruit qui est une baie, par ses stigmates très-courts , et par ses graines don’t la cicatricule est dépourvue d’arille ; toutes les espèces ont le pédicelle courbé ou tortillé dans le milieu. Observation. This genus, confused by Linnaeus with the uvular, but the organization of which the former botanists knew well, does not even belong to the Liliaceae family, of which the uvular is part of; it differs from it by its fruit which is a berry, by its very short stigmata, and by its seeds, the scar of which is devoid of aril; all species have curved pedicel or twisted in the middle.
    1856. Streptope embrassant. Streptopus amplexifolius; 1856. Streptope embrassant. Streptopus amplexifolius;
    Uvularia amplexifolia. Llnn. spec. 436. — Streptopus distortusi. Michaux. Fl. bor. am. i. p. 200.— Uvularia amplexicaulis. Delarb. FI. auv. 213.— Barr. t. 719 et 720. Uvularia amplexifolia. Llnn. spec. 436. — Streptopus distortusi. Michaux. Fl. bor. am. i. p. 200.— Uvularia amplexicaulis. Delarb. FI. auv. 213.— Barr. t. 719 et 720.
    Sa tige est haute de 5 décim. , rameuse , feuillée et cylindrique ; ses feuilles sont alternes , embrassantes , pointues , lisses et nerveuses ; ses fleurs sont petites , pendantes , solitaires et attachées à des pédoncules courbés dans leur milieu , et qui naissent à la base des feuilles; leur périgone est campanulé , et composé de six divisions lancéolées , distinguées chacune par une petite fossette à leur base intérieure : les étamines sont très-courtes; le fruit est une baie qui devient rougeâtre en mûrissant. ♃ Its stem is 5 decimeters high. , branched , leafy and cylindrical ; its leaves are alternate, embracing, pointed, smooth and sinewy; its flowers are small, hanging, solitary and attached to peduncles curved in the middle, and which arise at the base of the leaves; their perigone is bell-shaped, and composed of six lanceolate divisions, each distinguished by a small dimple at their inner base: the stamens are very short; the fruit is a berry that turns reddish as it ripens.
    On trouve cette plante dans les Alpes; les Pyrénées ; Je Jura; au Mont-d’Or ( Lemonn. ) , etc.; dans les montagnes du Forêt (Latour.). On la connoît vulgairement sous les noms de sceau de Salomon rameux , laurier alexandrin des Alpes , noms qui indiquent sa place dans l’ordre naturel. This plant is found in the Alps; the Pyrenees ; the Jura mountains; at Mont-d'Or (Lemonn.), etc.; in the mountains of the Forest (Latour.). It is commonly known by the names of branching Solomon's seal, Alexandrian laurel of the Alps, names that indicate its place in the natural order.

     

    Other articles:
    • Golden Checklist Flora:  Sisyrinchium montanum;  

    Sisyrinchium montanum Greene “Rocky Mountain Blue-Eyed Grass”

     

    Literature Cited:
    - Greene, Edward Lee, 1899c.  

    Greene (1899, v. 4, p. 33-34) ...

    S. montanum. Plant stout, erect, more than a foot high, herbage light-green, glabrous, not glaucescent : foliage rather copious but short, of less than half the length of the scapes, the broad leaves about 9-striate, the alternate lines commonly rather obscure; scapes ancipital, each of the broad, sharp-edged subentire wiags strongly 3-striate: spathes mostly solitary, their bracts very unequal, the outer of more than twice the length of the inner and 1½ to 2 inches long: perianths apparently dark-purple; capsules large (nearly ¼ inch in diameter), almost globose, very sparsely hairy.
    Meadows along the Mancos River, southern Colorado, Baker, Earle and Tracy (n. 113), 25 June, 1898 ; also by the same on Chicken Creek in the La Plata Mountains, at 9,000 feet, 7 July (n. 377). A large and doubtless showy species, exceeding even the Californian S. bellum in size.

     

    Other articles:
    • Golden Checklist Flora:  Calypso bulbosa;  

    Calypso bulbosa (L.) Oakes “Fairy Slipper”

     

     

    Other articles:
    • Golden Checklist Flora:  Coeloglossum viride;  

    Coeloglossum viride (L.) Hartm. “Longbract Frog Orchid”

    (Syn: Coeloglossum viride (L.) Hartm. ssp. bracteatum (Muhl.) Hultén,Dactylorhiza viridis (L.) R.M.Bateman, Pridgeon & M.W.Chase)

     

    Other articles:
    • Golden Checklist Flora:  Corallorhiza maculata;  

    Corallorhiza maculata (Raf.) Raf. “Summer Coralroot”

     

     

    Other articles:
    • Golden Checklist Flora:  Cypripedium parviflorum var. pubescens;  

    Cypripedium parviflorum Salisb. var. pubescens O. W. Knight “Greater Yellow Lady's Slipper”

    (Syn: Cypripedium calceolus L. var. parviflorum (Salisb.) Hultén)

     

    Literature Cited:
    - Rafinesque, Constantine Samuel, 1833.

    Other articles:
    • Golden Checklist Flora:  Goodyera oblongifolia;  

    Goodyera oblongifolia Raf. “Western Rattlesnake Plantain”

    Rafinesque (1833, v. 1, p. 76) ...

    10 Goodyera seu Tussaca oblongifolia, Raf. Fol. radic petiol oblongis ovatis acutis, 5 nervis non reticulatis, subtus glaucis caule gracile vaginato, spica laxiflora, fl. remotis hirsutis, bract. lanc. acut. ovar. eq. ovarium therto. — Oregon mts. subpedal, fl. white small.

     

    Other articles:
    • Golden Checklist Flora:  Spiranthes diluvualis;  

    Spiranthes diluvialis Sheviak. “Ute Lady's Tresses”

     

    Literature Cited:
    - Sheviak, Charles J., 1984.  

    Original Text
    TYPE: U.S.A. Colorado. Jefferson Co.: mesic to wet alluvial meadows along Clear Creek just W of junct. Rts. 6 & 58, Golden, 17 Jul 1982, C. J. Sheviak, J. K. Sheviak, W. Jennings, L. Long & S. Smookler 2257 (Holotype: NYS; isotype: NY).

     

    Other articles:
    • Golden Checklist Flora:  Spiranthes romanzoffiana;  

    Spiranthes romanzoffiana Cham. “Hooded Lady's Tresses”

     

     

    Other articles:
    • Golden Checklist Flora:  Leucocrinum montanum;  

    Leucocrinum montanum Nutt. ex A. Gray “Star Lily”

    As near as I can tell, the first common name applied to Leucocrinum montanum was 'White Crinum' in "Woolson & Co.'s descriptive catalogue of hardy perennial plants and price list : spring and fall of 1880." Miller (1884) in "A dictionary of English names of plants applied in England and among English-speaking people to cultivated and wild plants, trees, and shrubs" lists 'Californian Soap-Root' and 'Rocky Mountain Dwarf White Lily' as common names. 'Sand Lily' does not appear until 1904 in D. M. Andrews "Wholesale catalogue of rare seeds chiefly of plants and shrubs indigenous to the Rocky Mountains of Colorado." By 1926 the D. M. Andrews catalog lists the common name as 'Sandlily or Starlily."

    Literature Cited:
    - Gray, Asa, 1848.  

    A. Gray (1848) writing in his Revision of North American Melanthiaceae

    Original Text
    Leucocrinum montanum, Nutt. Ined.
    Radix e fibris plurimis crassitie pennae Corvi. Folia plurima, (ut videntur) crasiuscula, plana, 8 unc. Longa. Vix 2 lin. Lata, acutiuscula. Flores (6-8) foliis multò breviores; pedunculis omnibus radicalibus, unifloris, demùm ferè uncialibus. Perianthii tubus persistens, 2-3 unc. Longus, filiformis; limbus magnitudine florum Ornithogali umbellati. Stamina perianthium vix aequantia; antheris linearibus. Stylus staminibus paulò brevior; stigmatibus ferè obcordatis. Capsulae (immaturae) vix supra terram, membranaceae, 3-4 lin. Latae, (ut videntur loculucidè dehiscentes,) stylo et perianthio persistente coronatae. Semina ut in Colchico. — Floret Aprili.
    Hab. In planitiebus altis fluminis Platte, Nuttall!
    Obs. Genus a Colchico diversum, stylis coalitis, antheris adnatis, at capsulis membranaceis non inflatis, a Bulbocodia, unguibus perianthii totus concretis, etc. ; anutroque praesertim stigmatibus lamellatis, radice fascicilata, et defectu cormi. Si tamen Merendera, Bulbocodium, at Monocaryum, et monet doctiss. Brownio, * potius subgenera Colchici sunt, forsan hoc quoque illi referendum est.
    Specimina sicca a cel. Nuttallio lecta et mihi benevole communicata.

    Literature Cited:
    - Ornduff, Robert, and Marion S. Cave, 1975.  

    Sand-lily, Leucocrinum montanum Nutt., is a showy perennial that occurs widely in arid regions of the western United States. Recently, the second author reported chromosome counts of n = 11, 13, and 14 for this species (Cave, 1970). Plants from the Rocky Mountain region have n = 14 (see also Löve et al., 1971), those from several localities in California, western Nevada, and Oregon have n = 13, and one population from Nevada has n = 11. In addition, Cave noted that in some populations pollen is shed in tetrads and in others it is shed singly. In this paper we further discuss the variation in chromosome number and in the condition of pollen at the time of shedding (Ornduff and Cave, 1975).

       

    Studies of Prairie or Foothill Ecosystems

     

       

    Restoration and Effects of Attempts Thereof

     

    Literature Cited:
    - Wilson, Scott D., 1989.  

    Wilson, Scott D., 1989. The suppression of native prairie by alien species introduced for revegetation. Landscape and Urban Planning. 17(2), april 1989, pp. 113-119.

    Abstract

    Natural prairie has become uncommon in North America, making the revegetation of disturbed areas by native species a desirable goal. Alien species are often introduced for revegetation because of their abilities to stabilize and nitrify soil. The objective of this study was to test whether these attributes of introduced species would allow them to promote the recovery of native vegetation. Seven treatments (six commercially-available mixtures of introduced species and an unseeded control) were applied to a randomized field experiment in disturbed mixed-grass prairie in south-west Manitoba, Canada. Sampling eight years later revealed that introduced species suppressed native vegetation. Introduced species did not aid revegetation: plots seeded with introduced species did not produce significantly higher standing crop or below-ground biomass than did unseeded plots. Unseeded plots had the lowest frequency of bare ground. Allowing prairie to revegetate without sowing introduced species produced both the highest cover of bare ground and the greatest abundance of native species.

    Literature Cited:
    - Wilson, Scott D., and Joyce W. Belcher, 1989.  

    Wilson, Scott D., and Joyce W. Belcher, 1989. Plant and Bird Communities of Native Prairie and Intoduced Eurasian Vegetation in Manitoba, Canada. Conservation Biology. 3(1), pp. 39-44, March 1989.

    “… introduction of Eurasian plant species to North American prairie not only replaces the native plant community, but also produces significant changes in the species composition of a higher trophic level … ”

    Abstract: Large areas of North American prairie are dominated by Eurasian plant species introduced either for range improvement or accidentally as weeds. We examined the impact of introduced plants on both native vegetation and bird communities in a mosaic of North American mixed-grass prairie and Eurasian vegetation. We established ten transects five in areas of native prairie and five in areas dominated by introduced plant species. Each transect comprised five sampling stations separated by 100 m. Vegetation was sampled in four 0.5m2 quadrats at each station. The cooers of eight of the ten most common plant species varied significantly (p < 0.05) between native and introduced vegetation. One common native plans Andropogon scoparius, was absent in introduced vegetation Singing birds were identified to species at each station on three occasions during the breeding season All bird species found were native to prairie. The total number of birds did not vary between vegetation types Two out of eight bird species, upland sandpiper and Sprague's pipit were signifcantly more abundant in native prairie than in introduced vegetation No bird species were significantly more common in introduced vegetation. A correlation matrix calculated for all bird species and the ten most abundant plant species divided the bird community into two groups. The first group (western meadow lark, upland sandpiper, Sprague's pipit, Baird's sparrow and savannah sparrow) was positively correlated with native plant species and negatively with introduced plants, while the second (vesper sparrow, clay-colored sparrow, and grasshopper sparrow) was negatively correlated with native species and positively correlated with introduced Discriminant analysis separated transects from native and Eurasian vegetation on the basis of their respective bird communities. The results illustrate that the introduction of Eurasian plant species to North American prairie not only replaces the native plant community, but also produces significant changes in the species composition of a higher trophic level.

    Literature Cited:
    - Heidinga, Lawrence, and Scott D. Wilson, 2002.  

    Heidinga, Lawrence, and Scott D. Wilson, 2002. The Impact of an Invading Alien Grass (Agropyron cristatum) on Species Turnover in Native Prairie. Diversity and Distributions. 8(5), September 2002, pp 249-258.

    Abstract

    Alien invasions typically reduce species richness of habitats, but few studies have examined their effects on species turnover, the difference in species composition between localities. Agropyron cristatum (L.) Gaertn. (crested wheat grass) has been planted on 6-10 million ha of North American prairie, and is invading native prairie. We studied the invasion of A. cristatum into native prairie by measuring species composition along a gradient from maximum to minimum A. cristatum abundance. As A. cristatum increased, the abundance of most common native species decreased, but one appeared to be unaffected (Bouteloua gracilis (H.B.K.) Lag.), and another (Poa sandbergii Vasey) increased. The effect of A. cristatum on species turnover was investigated by examining species-area curves for areas from 0.5 m2 to 8.0 m2. Species diversity was reduced by 35% at high A. cristatum abundances at all areas. A. cristatum reduced the intercept of the species-area curve, but not the slope, suggesting that A. cristatum affected species turnover proportionally in all areas and habitats. This unusual result may indicate a homogeneous environment where species are distributed randomly. A. cristatum produced almost twice as many seeds as all native grasses combined. The number of seeds collected of native grasses and A. cristatum was highly correlated with the number of seed heads immediately nearby, but not with transect position. This suggests most seeds were dispersed over distances less than 5 m. In sum, the invasion of native prairie by A. cristatum might be related to high rates of seed production, and has the effect of decreasing species turnover by reducing the intercept of the species-area curve.

    Literature Cited:
    - Salesman, Jessica Bolwahn, and Meredith Thomsen, 2011.  

    Salesman, Jessica Bolwahn, and Meredith Thomsen, 2011. Smooth Brome (Bromus inermis) in Tallgrass Prairies: A Review of Control Methods and Future Research Directions. Ecological Restoration. 29(4), December 2011, pp. 374-381.

    Abstract. Smooth brome (Bromus inermis) is a cool-season rhizomatous grass of Eurasian origin that has escaped intentional plantings and spread widely in natural areas. A large body of work exists regarding smooth brome's biology and response to conservation management strategies designed to reduce its competitive effects on native prairie species, particularly for the tallgrass prairies of North America. Here we summarize that literature to improve restoration practice. In tallgrass prairie, smooth brome benefits from the early start of its growing season and its rhizomatous growth form, making it a strong competitor against native warm-season grasses. Late-spring burns timed to target smooth brome when root reserves are at their lowest have shown promise as a control strategy. Uncertainty remains, however, about the relative efficacy of fire, herbicide, mowing, or grazing to accomplish late-spring defoliation, the effect of repeated treatments, and the potential benefits of treatment combinations. The responses of resident or seeded natives to brome control treatments and/or the resulting decreases in brome cover also remain largely unexamined. Research focused on the questions we highlight would reduce costs associated with the control of smooth brome and increase confidence in the outcomes of restoration efforts.

    Literature Cited:
    - Grant, Todd A., Bridgette Flanders-Wanner, Terry L. Shaffer, Robert K. Murphy, and Gregg A. Knutsen, 2009.  

    Grant, Todd A., Bridgette Flanders-Wanner, Terry L. Shaffer, Robert K. Murphy, and Gregg A. Knutsen, 2009. An Emerging Crisis across Northern Prairie Refuges: Prevalence of Invasive Plants and a Plan for Adaptive Management. Ecological Restoration. 27(1), March 2009, pp. 58-65.

    Abstract

    In the northern Great Plains, native prairies managed by the U.S. Fish and Wildlife Service (Service) can be pivotal in conservation of North America's biological diversity. From 2002 to 2006, we surveyed 7,338 belt transects to assess the general composition of mixed-grass and tallgrass prairie vegetation across five "complexes" (i.e., administrative groupings) of national wildlife refuges managed by the Service in North Dakota and South Dakota. Native grasses and forbs were common (mean frequency of occurrence 47%-54%) on two complexes but uncommon (4%-13%) on two others. Conversely, an introduced species of grass, smooth brome (Bromus inermis), accounted for 45% to 49% of vegetation on two complexes and another species, Kentucky bluegrass (Poa pratensis) accounted for 27% to 36% of the vegetation on three of the complexes. Our data confirm prior suspicions of widespread invasion by introduced species of plants on Service-owned tracts of native prairie, changes that likely stem in part from a common management history of little or no disturbance (e.g., defoliation by grazing or fire). However, variability in the degree and type of invasion among prairie tracts suggests that knowledge of underlying causes (e.g., edaphic or climatic factors, management histories) could help managers more effectively restore prairies. We describe an adaptive management approach to acquire such knowledge while progressing with restoration. More specifically, we propose to use data from inventories of plant communities on Service-owned prairies to design and implement, as experiments, optimal restoration strategies. We will then monitor these experiments and use the results to refine future strategies. This comprehensive, process-oriented approach should yield reliable and robust recommendations for restoration and maintenance of native prairies in the northern Great Plains.

       

    Recreation in Foothill and Prairie Ecosystems

     

     

       

    Keys

     
     

    1830 Alabama - Georgia - South Carolina

     
     

    1834 - 1836 Oregon Trail, Hawai'i, Alta California

     
     

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    Vegetation Descriptions

     

     

       

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    Vouchers to Examine

     

    UTC

    Letters: Wednesday, November 5, 2014.  

    UTC10764: was determined Polemonium caeruleum whereas UTC19762, also bearing MEJ's #299 is determined Polemonium foliosissimum A. Gray. However, per Mary Barkworth, 11/25/2014, both are P. foliosissimum.

     

       

    Mystery Locations

     

    Locations: Gray Hill.  

    Gray Hill

    Harbouria trachypleura (A. Gray) J.M. Coult. & Rose. Whiskbroom Parsley. Mountain slope. Near Golden: Gray Hill. J. H. Ehlers 6848. 6/2/1938 ( RM184550 )

     

    West Cliff

    Quincula lobata (Torr.) Raf. (Syn: Physalis lobata Torr. ) Chinese Lantern. Golden, Road to West Cliff, Golden. Earl L. Johnston, with G. G. Hedgcock 813. 6/23/1917 ( RM101941 ).

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    Date and time this article was prepared: 4/27/2024 3:20:19 PM