Green Jay Cyanocorax yncas Scientific name definitions

Jordan C. Giese and Jared D. Hall
Version: 2.0 — Published January 19, 2024

Systematics

Systematics History

This species was previously placed in the monotypic genus Xanthoura (25, 26), a proposition that seems to have been first refuted by Zimmer (27), and thereafter its placement in Cyanocorax was solidified by Blake and Vaurie (28) and Hardy (29). South American populations are distinctive in color (generally darker green upperparts, or bluish in some, yellow underparts, nape whitish or blue, irides yellow or dark), vocalizations, social behavior (8, 9), and its montane distribution (mainly above 1,400 m, versus lowlands to 1,800 m in Middle American birds; 30, 13, 31, 32). It has thus been suggested that South American populations deserve separation as a different species (e.g., 33, 34). The molecular phylogeny of the genus Cyanocorax supplied by Bonaccorso et al. (7) provided further evidence that the Green Jay could be split into Middle American and South American groups, but their taxon sampling was far from complete. Gill et al. (35) currently treat the two groups at the species rank, though most other authorities continue to treat these groups as a single species (e.g., 36, 37, 10, 38).

Geographic Variation

Different authorities have accepted varying numbers of subspecies within the two groups: for example, between six and eight subspecies from North and Central America have been recognized in recent decades (39, 40, 41, 36, 37, 35), while in South America, the number of subspecies has been more stable at five (28, 40, 41, 36, 37, 35) or six (as here). Many of these subspecies, especially in the northern group, tend to intergrade and overlap (26, 13, 39), but northern and southern groups are reasonably well differentiated (differences are outlined under Systematics History) with the result that some authors treat them at the species level (35).

Subspecies


EBIRD GROUP (POLYTYPIC)

Green Jay (Green) Cyanocorax yncas [luxuosus Group]


SUBSPECIES

Cyanocorax yncas speciosus Scientific name definitions

Systematics History

Xanthoura luxuosa speciosa Nelson, 1900, Auk 17:265.—San Sebastián, Jalisco, Mexico. (42)

The holotype, an adult male collected on 13 March 1897 by Edward William Nelson (1855–1934) and Edward Alphonso Goldman (1873–1946), is held at the National Museum of Natural History, Smithsonian Institution, Washington, D.C. (USNM 156055; 43).

Distribution

Pacific slope of Mexico from northwest Nayarit to southern Jalisco.

Identification Summary

This subspecies is considered to be large and brightly colored, with a brown iris, forecrown yellowish, and underparts quite intense sulfur yellow (J. M. Marzluff in 41). Treated as a phylogenetic species by Peterson and Navarro Sigüenza (44).


SUBSPECIES

Cyanocorax yncas vividus Scientific name definitions

Systematics History

Xanthoura luxuosa vivida Ridgway, 1900, Auk 17:28.—Pluma, Oaxaca [Mexico]. (45)

The holotype, an adult male collected on 20 March 1895 by Edward William Nelson (1855–1934) and Edward Alphonso Goldman (1873–1946), is held at the National Museum of Natural History, Smithsonian Institution, Washington, D.C. (USNM 144810; 43).

Distribution

Pacific slope of southwest Mexico (Colima) to western Guatemala.

Identification Summary

Similar to luxuosus but considered to be slightly larger with brighter yellow rear underparts, a blue-tinged tail, and yellow irides (40, J. M. Marzluff in 41).


SUBSPECIES

Cyanocorax yncas luxuosus Scientific name definitions

Systematics History

Garrulus luxuosus Lesson, 1839, Revue Zoologique 2:100.—Mexico.

The holotype of Lesson’s name was said by him to be in the possession of a Dr. Abeillé who, in the original description of the Emerald-chinned Hummingbird (Abeillia abeillei), revealed it was a resident of Bordeaux at this time; the person honored was Dr. Grégoire Abeillé (1798‒1848; 46), and a catalog of his private collection numbering ~1,500 specimens (of which approximately 100 involved types described by Lesson) was published, presumably in anticipation of it being sold, two years later, in 1850 (47). The suggestion that the person concerned was the botanist and horticulturalist Paul-Emmanuel Abeille de Perrin (1797‒1868; 48) is quite erroneous. Its current whereabouts, if the specimen still exists, is unknown.

Synonyms:
C[yanocorax]. cyanocapillus Cabanis in von Tschudi, 1846, Untersuchungen über die Fauna Peruana, Ornithologie, p. 233.—Xalapa [= Jalapa Enríquez, Veracruz, Mexico; 19°34’N, 96°54’W]. Cabanis referred to “several” birds in the Museum für Naturkunde, Berlin (ZMB) from Xalapa; Steinheimer (49) noted that ZMB 1521, which is now labeled only “Mexico,” was nevertheless certainly one of this name’s syntypes, to which list he added ZMB 1522 (a male) and ZMB 1523 (sex/age unknown) both from Xalapa and believed to have been collected in January 1825 (50) by Graf Sebastian Albert Freiherr von Sack (1757–1829) and Ferdinand Deppe (1794–1861; 49).

Xanthoura luxuosa glaucescens Ridgway, 1900, Auk 17:28.—Fort Brown [= Brownsville, Cameron County], Texas (45). [Forth Brown in 28 is clearly a misprint.] The holotype, an adult male collected on 30 March 1876 by J. C. Merrill, is held at the National Museum of Natural History, Smithsonian Institution, Washington, D.C. (USNM 70593; 43). This subspecies was accepted by, among others, Madge and Burn (40), Gayou (24), Dickinson and Christidis (36), and del Hoyo and Collar (37), with its distribution generally believed to encompass southern Texas (USA) and northeast Mexico in western Nuevo León and Tamaulipas. Madge and Burn (40) considered it slightly paler and duller than luxuosus, with dark brown irides and blackish legs.

Cyanocorax yncas persimilis A. R. Phillips, 1966, Bulletin of the British Ornithologists’ Club 86:111.—2 km north of Ocotal Chico, near Cerro Santa Marta, southeastern Veracruz (51). Known from two syntypes, an immature (perhaps male) and an adult female collected by Florentino Francisco R. on 11 December 1962, originally in the Allan Robert Phillips (1914–1996) collection and now at the Delaware Museum of Nature and Science, Greenville (DMNH 21898 and DMNH 21899, respectively; 51, 52). Phillips (51) characterized this taxon by its more purplish-blue head, especially the sides and nasal tufts, and averaging marginally darker, less yellowish, more green, dorsally.

Distribution

Southern Texas (Rio Grande Valley) to eastern Mexico (northeast Puebla and Veracruz).

Identification Summary

Typical of Middle American subspecies in having a short blue frontal tuft, blue crown and nape, greenish-yellow underparts becoming yellower towards the vent; brown legs and dark irides (40, J. M. Marzluff in 41). This subspecies is reported to intergrade with vividus in the southeast Sierra de Tuxtlas and Catemaco region of southern Veracruz (52, J. M. Marzluff in 41, 37).


SUBSPECIES

Cyanocorax yncas centralis Scientific name definitions

Systematics History

Xanthoura luxuosa centralis van Rossem, 1934, Bulletin of the Museum of Comparative Zoology 77:397.—Secanquim, Alta Vera Paz, Guatemala. (53)

The holotype, an adult male collected by A. W. Anthony on 15 January 1926, is held at the University of California Los Angeles Donald R. Dickey Bird and Mammal Collection (UCLA 31838; 53).

Synonym:

Cyanocorax yncas confusus A. R. Phillips, 1966, Bulletin of the British Ornithologists’ Club 86:111.—4 and 6 km east of Pijijiapan, Chiapas (51). Known from two syntypes, an immature (perhaps female) and an immature female collected by Abraham Ramírez V. on 15 November 1964 and 6 October 1965, respectively, originally in the Allan Robert Phillips (1914–1996) collection and now at the Delaware Museum of Nature and Science, Greenville (DMNH 21933 and DMNH 18418, respectively; 51, 52). Phillips (51) described confusus as differing from vividus by its smaller size, yellow irides (except in juveniles?), and by being slightly darker above and, especially, on the head sides.

Distribution

Southeastern Mexico (extreme eastern Tabasco and adjacent parts of Chiapas east through extreme southern Quintana Roo), northern and eastern Guatemala, Belize, and northern Honduras.

Identification Summary

Has the underparts yellow or yellowish, tinged lime-green (occasionally approaching luxuosus; 40, J. M. Marzluff in 41).


SUBSPECIES

Cyanocorax yncas maya Scientific name definitions

Systematics History

Xanthoura luxuosa maya van Rossem, 1934, Bulletin of the Museum of Comparative Zoology 77:397.—Río Lagartos, Yucatan. (53)

The holotype, an adult female collected by Wilmot Wood Brown (1870?–1953) on 1 June 1893, is held at the Museum for Comparative Zoology, Cambridge, MA (MCZ 115266) (53, 54).

Synonym:

Xanthoura luxuosa cozumelae van Rossem, 1934, Bulletin of the Museum of Comparative Zoology 77:397.—Cozumel Island, off the coast of Quintana Roo, Mexico (53). The holotype is an unsexed adult collected in January 1886 (not 1866, as sometimes reported) by George Franklin Gaumer (1850‒1929), and held in the Natural History Museum, Tring (NHMUK 1886.9.9.862; 53, 55). Subspecies maintained by Madge and Burn (40) and J. M. Marzluff in dos Anjos (41). Madge and Burn (40) characterized this subspecies as small with a pale blue crown and bright yellow underparts.

Distribution

Southeastern Mexico (Yucatán Peninsula) and extreme southern Quintana Roo.

Identification Summary

Characterized by its pale but bright (sulfur-)yellow underparts and white forecrown (40, J. M. Marzluff in 41).


EBIRD GROUP (POLYTYPIC)

Green Jay (Inca) Cyanocorax yncas [yncas Group]

Available illustrations of subspecies in this group

SUBSPECIES

Cyanocorax yncas galeatus Scientific name definitions

Systematics History

Xanthoura yncas galeata Ridgway, 1900, Auk 17:27.—Western Colombia (45) [= El Edén, 8,300 ft, Central Andes (Tolima side), suggested by Chapman (56)].

The holotype, an adult (sex unknown) purchased in December 1878 from J. Wallace by Robert Ridgway, is held at the National Museum of Natural History, Smithsonian Institution, Washington, D.C. (USNM 81879; 43).

Distribution

Subtropical zone of Colombia west of East Andes, including the West Andes south to Valle; the Central Andes; and the west slope of the East Andes.

Identification Summary

Similar to the nominate subspecies but has yellow irides (brown in Colombian populations of yncas) and a larger frontal crest (40, J. M. Marzluff in 41).


SUBSPECIES

Cyanocorax yncas cyanodorsalis Scientific name definitions

Systematics History

Xanthoura yncas var. cyanodorsalis A. J. C. Dubois, 1874, Bulletins de l’Académie royale des sciences, des lettres et des beaux-arts de Belgique (2)38:492—“Nouvelle Grenade” [= Colombia].

In the original description, Alphonse Joseph Charles Dubois (1839–1921) mentioned that he had two specimens (syntypes) of cyanodorsalis; both are adults, were supplied by the natural history dealer Emile Parzudaki, and are held in the Royal Belgian Institute of Natural Sciences, Brussels (RBINS 3086 and RBINS 3087; O. Pauwels in litt. to G. M. Kirwan, August 2023).

Distribution

East slope of East Andes of Colombia (Norte de Santander to Cundinamarca) and northwest Venezuela (Zulia and southern Lara to Barinas).

Identification Summary

Considered to be smaller than galeatus, with a large deep blue frontal crest, narrow white band on forecrown, deep blue crown and hindneck, darker green upperparts (washed blue, especially in worn plumage), and legs darker; irides yellow (40, J. M. Marzluff in 41).


SUBSPECIES

Cyanocorax yncas andicolus Scientific name definitions

Systematics History

Xanthoura yncas andicola Hellmayr and von Seilern, 1912, Archiv für Naturgeschichte Abteilung A 78(5):72.—El Walle, Mérida, western Venezuela. (57)

The holotype, an adult male collected on 11 November 1897 by Salomón Briceño Gabaldón (1826‒1912), is held in the Zoologische Staatssammlung München (ZSM 1909.3422; 57). Hellmayr (25) himself subsequently repudiated andicolus, possibly because it had been originally described through misidentification of western Colombian galeatus as cyanodorsalis, but Zimmer (27) thought that it was, after all, probably valid.

Distribution

Mountains of northern Venezuela.

Identification Summary

Zimmer (27) was tempted to admit andicolus as valid, being close to guatimalensis, but the frontal crest averages slightly longer, the white post-frontal band is always well developed, and the bill is heavier, at least in most cases.


SUBSPECIES

Cyanocorax yncas guatimalensis Scientific name definitions

Systematics History

Xanthoura guatimalensis Bonaparte, 1850, Conspectus Generum Avium, Tome 1, p. 380.—Guatemala; error [= Caracas, Venezuela, fide van Rossem (53)], but Bonaparte’s mistake was earlier noticed by Schlegel (58) and also by Sclater (59).

This name is based on two syntypes: an adult labeled as being supplied by Reinhart Frans Cornelis van Lansberge (1804‒1873) that is held in the Naturalis Biodiversity Center, Leiden (RMNH.AVES.90618; 60); the other was said to be in the Royal Belgian Institute of Natural Sciences, Brussels (60), but cannot currently be located there (O. Pauwels in litt. to G. M. Kirwan, August 2023). Often dated from 1851 (see, e.g., 28), but we follow Zimmer (61), Dickinson and Christidis (36), and del Hoyo and Collar (37) in using 1850.

Synonym:

Xanthoura yncas var. caeruleocephala A. J. C. Dubois, 1874, Bulletins de l’Académie royale des sciences, des lettres et des beaux-arts de Belgique (2)38:493.—“Venezuela” [= Caracas]. This name was based on a single specimen in what is now the Royal Belgian Institute of Natural Sciences, Brussels (RBINS 3088; O. Pauwels in litt. to G. M. Kirwan, August 2023).

Distribution

Mountains of northern Venezuela (Falcón east to Sucre and Anzoátegui).

Identification Summary

Considered to be most similar to cyanodorsalis, but has a shorter crest, with less blue wash on the upperparts, and an even narrower white band on the forecrown; irides yellow and legs brownish (40, J. M. Marzluff in 41).


SUBSPECIES

Cyanocorax yncas yncas Scientific name definitions

Systematics History

Corvus yncas Boddaert, 1783, Table des planches enluminéez d'histoire naturelle de M. D’Aubenton, p. 38; based on Daubenton, Planches enluminées d’histoire naturelle, Plate 625.—Peru; type locality restricted by Zimmer (27) to Chilpes, Dept. Junín.

The holotype of this name is the bird illustrated in Daubenton’s plate, but no specimen that can be identified with the latter is known to be extant in the Muséum national d’Histoire naturelle, Paris.

Synonyms:

[Corvus] peruvianus J. F. Gmelin, 1788, Systema Naturae, Tomus 1, p. 373; based on Daubenton, Planches enluminées d’histoire naturelle, Plate 625.—Peru; type locality restricted by Zimmer (27) to Chilpes, Dept. Junín.

P[ica]. Chloronotos Wagler, 1827, Systema Avium, genus Pica [= signature 21, p. (9)].—Peru; type locality restricted by Zimmer (27) to Chilpes, Dept. Junín. Described from a specimen apparently in the property of the natural history dealers Ryals and Leadbeater, London. Van Rossem (53) apparently believed that this name referred to a composite, of which the main part could be applied to Venezuelan birds (herein guatimalensis), but he later accepted that this was a false premise (27). With hindsight, van Rossem seems to have engaged in nomenclatural gymnastics solely out of a desire not to apply a geographically inappropriate name (coined for what was believed to be a Central American bird), when such issues are commonplace in zoology.

Pica luteola Lesson, 1831, Traité d‘Ornithologie, Livraison 5, p. 331.—type locality said to be unknown. The holotype, a specimen reportedly from Peru (which can thus be considered terra typica) presented by Mn. Agasson (also reported as Ajassou), is held in the Muséum national d’Histoire naturelle, Paris (MNHN-ZO-2011-565; 62); Zimmer (27), on the authority of Berlioz, was doubtful that the specimen was the type of Lesson’s name, but it is nonetheless currently afforded holotype status. To stabilize nomenclature in favor of Carriker’s at the time recent erection of subspecies longirostris, Zimmer (27) restricted the type locality of luteola to Cajamarquilla, Dept. Pasco (formerly Junín, Peru).

C[yanocorax]. peruanus Cabanis in Tschudi, 1846, Untersuchungen über die Fauna Peruana, Aves, p. 232.—Peru. Name (which is an unnecessary of emendation of peruvianus, see above) was credited to Tschudi by Zimmer (27), but authorship should be clearly accorded to Cabanis, like other novel nomina in the same work.

Distribution

Southwestern Colombia (subtropical zone, in valleys of upper Cauca, Patía, and San Miguel, in Cauca and Nariño) south through eastern Ecuador and Peru to central Bolivia (La Paz, Cochabamba, and western Santa Cruz).

Identification Summary


SUBSPECIES

Cyanocorax yncas longirostris Scientific name definitions

Systematics History

Xanthoura yncas longirostris Carriker, 1933, Proceedings of the Academy of Natural Sciences of Philadelphia 85:30.—Soquián, 8,000 ft., Marañón Valley, Prov. Huamachuco, Dept. [La] Libertad, Peru. (63)

The holotype, an adult male collected by Melbourne Armstrong Carriker, Jr. (1879–1965) on 18 June 1932, is held at the Academy of Natural Sciences at Drexel University, Philadelphia (ANSP 108083; 63).

Zimmer (27) described this subspecies as “unsatisfactory but has just enough average distinction from the general series of yncas (from Ecuador to Bolivia) to give it a place”, and as a result it has been maintained ever since.

Distribution

Arid upper valley of Río Marañón, in northern Peru.

Identification Summary

Generally very similar to the nominate subspecies but is larger; irides yellow (40, J. M. Marzluff in 41).

Related Species

Phylogenetic relationships in the genus Cyanocorax were studied by Bonaccorso et al. (7), who recovered two clades within this genus. Cyanocorax yncas formed part of the larger of the two clades, which was also comprised of the following species: White-tailed Jay (Cyanocorax mystacalis), Cayenne Jay (Cyanocorax cayanus), Plush-crested Jay (Cyanocorax chrysops), White-naped Jay (Cyanocorax cyanopogon), Azure-naped Jay (Cyanocorax heilprini), Black-chested Jay (Cyanocorax affinis), Tufted Jay (Cyanocorax dickeyi), Bushy-crested Jay (Cyanocorax melanocyaneus), Yucatan Jay (Cyanocorax yucatanicus), San Blas Jay (Cyanocorax sanblasianus), and Purplish-backed Jay (Cyanocorax beecheii). However, within this clade, the precise relationships of Cyanocorax mystacalis and Cyanocorax yncas were consistently ambiguous across different analyses, with Cyanocorax yncas either found to be sister to the entire clade, or to the entire clade excluding Cyanocorax mystacalis (7). The wider-scale molecular insight provided by Jønsson et al. (64) resulted in a very similar arrangement (see also 65).

Hybridization

No Green Jay hybrids have been reported in wild populations (DCG), but a hybrid between this species and a Blue Jay (Cyanocitta cristata) has been recorded in captivity (66).

Hybrid Records and Media Contributed to eBird

  • Green x Blue Jay (hybrid) Cyanocorax yncas x Cyanocitta cristata

Nomenclature

The Green Jay is a familiar and distinctive bird throughout its range, and thus is known by a variety of names. Below is an incomplete list of names used for Cyanocorax yncas by language:

  • In Mayan languages, Green Jay is referred to as Kexkex or Kejkex (Ch'ol Maya, Chiapas, Mexico; 67), and Ses-ib (Yucatán, Mexico; 68).
  • In Shipibo (Peru), it is referred to as Iori (69).
  • In Spanish, it is regionally referred to as Queisque or Queixque Verde, Checla Verde, Pájaro Verde, Urraca de Montaña, Picachayote, Ruiseñor, Carriquí Verdiamarillo, Chara Querrequerre, Querrequerre, Chara Verde, Urraca Café, Urraca Querrequerre, Urraca Verde, and Urraquita Verde (70, 68).
  • In English, the Green Jay is also referred to as Inca Jay (70).

Fossil History

One Holocene (10,000 yr before present) record from the Yucatán Peninsula (71).

Recommended Citation

Giese, J. C. and J.D. Hall (2024). Green Jay (Cyanocorax yncas), version 2.0. In Birds of the World (S. M. Billerman, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.grnjay.02
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